Corresponding author: Michael J. Sharkey (
Academic editor: G. Broad
Based on a cladistic analysis, a new genus of
Hymenoptera Institute Collection, University of Kentucky, Department of Entomology, Lexington, Kentucky, USA
Queen Sirikit Botanic Gardens, Chiang Mai, Thailand
Taiwan Agricultural Research Institute, Entomology Collection, Taichung, Taiwan
Generic concepts: This is the fourth of a series of papers (
The NJ analysis (
NJ tree showing 28S distances between species of
Maximum parsimony tree of some agathidine genera with a concentration near
Morphological terms used in this revision were matched to the Hymenoptera Anatomy Ontology (HAO;
All species are treated with a diagnosis and distributional data. They are illustrated with color photos using a JVC digital camera mounted on a Leica MZ16 microscope and Automontage® stacking software. Distributional data are listed for all species and a link to a Google map is included for all species. The descriptions are of holotypes and variation is given in parentheses.
Members of all three genera have distinct pegs on the anterior surface of the fore tibia (
Interantennal space with longitudinal keel that is sharply declivous posteriorly (
Unknown
Restricted to the eastern Palearctic (Taiwan) and the Oriental region. For a distribution map,
From the Greek
1a | Median tergite 1 mostly smooth or weakly sculptured |
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1b | Median tergite 1 mostly striate or rugosostriate | 2 |
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2a | Fore wing uniformly yellow, at most slightly darker apically | 3 |
2b | Fore wing clear basally, infuscate distally | 4 |
2c | Fore wing mostly evenly infuscate, very slightly clearer at extreme base |
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3a | Head mostly yellow, black color restricted to medial dorsal surface; larger specimens, ~ 9.6 mm |
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3b | Head mostly black, yellow color restricted to part of face and orbits of eyes; smaller specimens, ~ 6.2 mm |
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4a | Median tergite 1 (T1) weakly converging anteriorly; T2 quadrate and ~1.3× wider than long |
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4b | Median tergite 1 strongly converging anteriorly; T2 diverging posteriorly and ~1.5× wider than long |
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This species has the northern-most distribution and appears to be restricted to Taiwan where it is the only species present. Members come in two color morphs; one with the mesosoma bicolored and the other with the mesosoma entirely melanic.
Length: 6.5 (7.8) mm. Antenna with 38 (42) flagellomeres. Number of spines on fore, mid, and hind tibiae 6(5), 8(11), 15(16). First median tergite 1.6 times longer than wide; second median tergite 1.2 times wider than long; median tergite 1 rugosostriate, except smooth near posterior margin, sculpture more reticulated than congeners; median tergite 2 smooth (weakly rugosostriate near midlength). Color: (melanic color of mesopleuron more extensive in the sole paratype.)
Genbank Accession
Length: 8.4 mm. Antenna with 41 flagellomeres. Number of spines on fore, mid, and hind tibiae 6, 10, 13. First median tergite 1.5 times longer than wide; second median tergite 1.5 times wider than long; median tergite 1 entirely rugosostriate; median tergite 2 mostly weakly sculptured with striae, smooth posterolaterally.
Named after Kratae Sanok a collector for the TIGER project at Pa Hin Ngam National Park.
Genbank Accession
Length: 9.6(9.3) mm. Antenna with 44 flagellomeres. Number of spines on fore, mid, and hind tibiae 4(3), 6, 6(13). First median tergite 1.6 times longer than wide; second median tergite 1.4 times wider than long; median tergite 1 rugosostriate, except smooth near posterior margin; median tergite 2 mostly smooth with a weak indication of longitudinal sculpture at midlength. Color: (Melanic color of head restricted to ocellar triangle in the sole paratype.)
Named after Sailom Tongbunchai a collector for the TIGER project at Phuphan National Park.
Genbank Accession
Female:
Length: 8.4 (9.8) mm. Antenna with 43(45) flagellomeres. Number of spines on fore, mid, and hind tibiae 6, 8, 12(15). First median tergite 1.8 times longer than wide; second median tergite 1.2 times wider than long; median tergite 1 rugosostriate, except smooth near posterior margin; median tergite 2 mostly smooth with a weak indication of longitudinal sculpture at midlength (almost entirely smooth). Color: (melanic color of mesopleuron more extensive in one paratype; mid femur melanic in basal half).
Named after Winlon Kongnaka a collector for the TIGER project at Phuphan National Park.
Genbank Accession
Female:
We thank the staff at Queen Sirikit Botanic Garden in Chaing Mai for sorting the many hundreds of samples and for the Thai park staff for operating Malaise traps and other collection devices. A special thanks to Chaweewan Hutacharern for managing the Thai end of the TIGER project. Also many thanks to the curators around the world (London, Oxford, Warsaw, Leiden, Gainesville, Budapest, Beijing, Paris, Stockholm, Washington) for lending specimens and hosting MJS on visits. Special thanks to Kees van Achterberg for lending specimens and types from Vietnam. Funding was provided by NSF grants DEB-0542864 and EF-0337220.