First record from Costa Rica of the genus Caenophanes Foerster and description of a new species ( Hymenoptera , Braconidae , Doryctinae )

The genus Caenophanes Foerster is distinguished from the genus Heterospilus Haliday and one new species is described from Costa Rica which is the first species of Caenophanes described from the


Introduction
The small genus Caenophanes Foerster has, for a long time, been considered a synonym of Heterospilus Haliday but was recently resurrected as a valid genus by Belokobylskij (1993) who placed it in the tribe Doryctini.It is similar to genera in the tribe Heterospilini because of the absence or weakness of fore wing vein 2RS, but is distinguished by having the first subdiscal cell of the fore wing closed at its apex.Belokobylskij (2006) presented a key to the World genera of Doryctinae which have an absent or weak fore wing vein 2RS.Species of Caenophanes have been recorded from the Australasian, Oriental and Palaearctic Regions.During a recent study of the Heterospilini from Costa Rica (Marsh et al. 2013) one new species of Caenophanes was discovered.Specimens are also known from Mexico (Belokobylskij et al. 2011) and Wyoming (Scott Shaw, per. comm.)thus expanding the distribution of the genus to the entire Western Hemisphere.The purpose of this paper is to document the distribution of the genus Caenophanes for Central America and to present a description of the new species from Costa Rica.

Materials and methods
Specimens were found among the large collection of Heterospilini borrowed from the University of Wyoming.All specimens were examined using a Wild M5 binocular stereomicroscope and fluorescent illumination.Specimens for the scanning electron micrographs were gold/palladium coated using a Denton Desk II TSC turbo-pumped sputter coater and the micrographs were made on a Philips XL30 Environmental Scanning Electron Microscope.Minor levels of adjustments to the images were performed in Adobe Photoshop versions CS3 and CS4, and the plate was prepared in the same program.
Terminology for morphological characters largely follows that of Sharkey and Wharton (1997) and Marsh (2002).Terminology for surface sculpturing follows that of Harris (1979), Marsh (2002) and Marsh et al. 2013.
Label data for holotypes and paratypes is listed exactly as indicated on the labels including misspellings, punctuation, abbreviations and absent spaces.Lines on each label are separated by a bracketed semi-colon [;].
All type material is deposited in the Insect Museum at the University of Wyoming, Laramie, WY (ESUW) and the Department of Entomology, University of Illinois (UILL).Diagnosis.Small to moderate size, 1.5-3.5 mm; occipital carina not meeting hypostomal carina; fore tibia with single row of small spines; hind coxa with distinct antero-ventral basal tooth; basal sternal plate (acrosternite) of metasomal segment 1 short, less than ¼ length of tergum; fore wing second submarginal cell elongated, vein 2RS absent, vein r-m present, first subdiscal cell closed at apex, vein 3CU on same line as vein 1CU; hind wing vein SC+R present, male without stigma in hind wing.
Comments.Caenophanes is similar to Heterospilus by having the fore wing vein 2RS absent, but is distinct by having the fore wing first subdiscal cell closed at the apex.
A key to species was presented by Belokobylskij and Maeto (2009).
One new species from Costa Rica is described below which is the first record for the genus in the Western Hemisphere.