xread 'Mutillidae analysis Brothers & Lelej 2017; NOTE numbering starts with 0'230 105 Hemipepsis 001100000010001000130--1000010000500001-00100000-1[02]0010--000010-110010330000100000000-00-0000-00010000000010001300--100002000501[01]00000200000000000001100011002-10102454000014002200310011[12]0100111000011000000-0000-2000000310100000010 Anthobosca 000[01]10002010000210110--1000220000000001-00000000-000000--000021-002000030000010000000-00-0010-00010200000000100100--000002000000000000000000000000000000012010-03000020000000001000000000000000[01]0000000000000-0000-1000081012100100200 Fedtschenkia 0011[12]0010010011000130--1200000001000000-00100000-000000--000111-000000420000000000000-00-0010-10010200001010001200--120001100000000000100000000000001000101001-000000[23]0000000001000020010200000[01]0001001000000-0010-1000001010101100200 Sapyginae 2011[01]001[02]010021[01][01]0120--1300001001000000-00[01]20000-100000--000000-0000000[13]000001000000[23]100-00[01]0-10100[02]02002010[01][01][12]200--130001100000002000[12]00000000100001000000001-000000[14]00000000010000000102000000[01]001001000000-00[01]0-20001-1010101100000 Acanthomutilla 201200101000212110221003210110000002011002122201011402500111111020107021100300221002200101320-00110102001121111220--2210120004211202211315[12]111101110122112111232140120111020001120121111120111-1110001220210110120-1060027221101300201 Allotilla 201200101120212220110--01003200000010111021002110110000--011211320104022000301221101210101320-00111115001321111100--01002200002101022121000000010000122102001222150101000020001100013121020-11-102002122121211012121000032200101300200 Amblotropidia 201200102100212010221003210110020001011402102101111002601111111020107022100300221002200101320-01110112001[12]23[01][01]121[01]00[01]21012020021120[02]011312[12]111101[12]101221111212[18]21431[02]0[01]11020[01]01120221111110111-1120001220210110120-1050027221101301201 Ancistrotilla 201200101231212110120--1310200[23]000010111021222010100006010013110201050211103002221320-0101[23]20-011121261015221[01]1100--1310220[34]0031120221231030000001101221[12]2101262140100010020001100103111010011-102001122122111012121006032200101300201 Apteromutilla 201200101120212120220--0200200000001011202112211010001100001111020109421100100222002[12]00101120-00110115001020122300--12101110001112122--------------01--1--1012[02]21631010100[12]003--------------------------022011012122000034110101300200 Areotilla 001210100020212200010--12000000000010110120000210022129100203113201050000003011200110-00-1020-001000120013231[01]1100--110001000021020001000501211022101110[23]1001232150000000020001000001121011-100102101122001010012101001030121101300210 Artiotilla 201200101100212010121003210210020001011002102201000002501111111020107421100300221002200101120-00110112001123001210--02101200002113022113151111101110122112101202143120111020101120121111120111-112101122021011012122050027221101300200 Atillum 201200141120212011232[01]210004[12]03[02]00010121021221010100020--011311620104322000300221012100101320-01110125001220001302021000420300112122[02]001[01]01200010100122131001252142101000020001100[01]42[12]41020-11-10200[12]1220210110120-200[07]030110101300200 Bischoffiella 201210111020212200100--01012100200202112020000211002016000113115201050010010012100010-0130320-0011011200142010110[02]0001112103001103[01]11112150110002210122111121232250010100000101[01]00000111110111-101002122012110312102083050201011400210 Bischoffitilla 20120010212021201022[01]00[01]200300010002011012132221000012601021311320105020[01]0030[01]2210120-0101220-00110113001220111200--[01]200310010311322212210120000[01]0001221[12]1001212150100010020001131121021120-11-11[23][01]011220210110120-20[02]00022[12]1101300200 Bothriomutilla 201200111232212110122001220210000001011112132201001200201011111420104021100300221[01]12100101320-01111126101322101100--1300220000211232202010211111110012211-101202140100010020001100133111010011-102001122122011012122000032200101300201 Brachymutilla 201200101120212110120--[013]2001100000010112021022210100002000[012]131112010242[12]0004012210[01]20-010132[01]100110115021020121[23]00--0200[12]100[01]0111[01]122--------------01--1--10121212310100003003--------------------------021[12]11012[01][12][12]0011-1010101300200 Cephalomutilla 2012001212312120102221012003100000010121021222011000001--0111113201040221001012320[02]2210101320-01110026101323101100--0200220000211312212310300001000012211210123214010100002000110000[12][12]21020-11-1120021220421110120-2015032201101300201 Ceratotilla 20120012210021201022[01]003200[23][012][01]0[01]0202011002122111011402[56]01111111[56]2010202110030[01]2210[03]22001013211001102130012201112010022103110102112221--------------01--1--10125212010101002002--------------------------021011012121000002212101300200 Chaetomutilla 2012001011002120002322032101200001020110021120110114025011112110201060221003002210320-0101320-01110[01]03001[12]210001020[04]12001213013113020102052110101100122111021212150121210020101120121111120111-1121011220210110120-10700222[12]2101300200 Chrestomutilla 201200111120212110110--13101100000020111020222211010001010112110201020210003012310320-0101220-00110113001323101100--01121100003113122113101300000000122121111222140101010020001100031111110011-102001122031111012111001032200101300200 Ctenotilla 2012001011002120001210012001100001010110021222010114025011111113201040211003012210320-0101320-0[01]110103001122100001002200120[01]012102222101152111101100122111111212150121210020101120121111120111-1121001220211110120-10[13]5022211101300200 Cystomutilla 201200101220212110120--13001100000010111021212010000011000111115201040210003002211120-0101320-00111115001321111200--0300120300211202211010140000110012211[12]101202140100000020001100001221010-11-102101122111111012112000044200101300201 Darditilla 2012001[34]12312121[12]022210320041002000101211212220101[01]01020[01]1012116201050211003002211[02]22[01]0101320-00111126001322101100--0200420000311302212110240000010012212-101242150100010020001000002221010-11-1020011221210113120-2006030200101300201 Dasylabris 20120010012021211012[01]001310100000002011102102211110000200021[23]11[02]20100022100102232032100101320-01110105001[12]22100100--13101[01][01][24]00211[23]0221211040000000001221[23][12]1012121001010100100011311511[12]1120011-1020021220321110120-1000032[12]00101300000 Dasylabroides 2012001[01]0120212010[12]20--121021002000[12]011102122111110000200011111020102[01]22100[23]002210[23][12]0-0101320-0111011500132[01]10[01][12]00--[01]2002[01][01][12][01]02112022101[12]01200001[01]001221[13][12]0[01]12121[02][01]1010100[12]00[02]1100021111010011-1020021220121110120-2000032[12]00101300200 Dasymutilla 2012001[014]123[12]2120[01]0[12][23][02]00130021000000[12]0111[01]2122201[01][01]0000[012]000[01]111102010[045][04]2[12]0[01]0[13]0[01]2[23]2[01][013]2[12][01]0101320-01111126101522[01]01[12]00--1200210000211[23]12212[03]10[34]0000[01]0[01]001221[13]21012[034]21[45]010[01]000020001100113[12]110[01]0[01]11-10200[12]122[01][13]22110120-200[06]033201101300201 Dentilla 201200101[01]00212110120--13001000000020110021222[12]1111002500111111020104021100401221002100101320-001101[01]1001[45]22[01]00[12]00--0200111210[23]1130221032021111011001221121012[16]2140100010030001120121111120111-11[12]00112202111101211100003622[01]101300100 Dilophotopsis 201200101122212110120--13001101000010111021002010110002000113110201040220003012322020-1101320-001121230014211[01]1100--02001110002101222120101[12]00001100122132011212143100010[01]200011000012[12]1020011-002[01]021222[02]10111120-2010042200101300201 Dimorphomutilla 201200132131212120230--33004[01][01]02000101110212[012]2010002110--01131152010502110030[01]221102200101[23]20-0011112[56]001321111100--02004[12][01]0[01]0210102212110[12]400000000122121101252150100010020001100001121010-11-001000122102[01]110120-2000032200101300201 Dolichomutilla 2012001121[02]0212000221[12]03210110[13]0030101101200220100000250[01]11111102010442[01]110300221002200101220-0311010[34]001122101210--02101203033113022113[01]22[01]21101010122112001212140100011020001020223111120011-10[12]0001220210110120-1020026220101300200 Ephuta 20120010[12]1[02]02121[01]01[12][01]22432011000060[12]011302[01]000210[01]00010---21311320104021000002242032[02]00111[23]20-0011011400152[23]211[01]0[01]00332012000631122[01][01]1[12]3152122001[12][01][01]1221[123][12][01]012321401000[01]0020001021251[12][12]1120111-10200[12]1220522[01]00120-[12]007035221101300200 Ephutomma 201200102000212110120--1300100000001011002002211100002600111211020104021100300222032100101320-00110112011[24]22001200--02001112002113022103201111101000122122001212143100010020001120131111110111-1021021220221110120-1000037222101300200 Eurygnathilla 201200142020212010230--02003010100010110021222110010026010010113201050201003002210320-0101220-00110205001121001200--12003010001113022101101100001100122111101202140100010020001131221021120-11-112001122021011012122020022211101300200 Eurymutilla(Near) 2012001012312121101210012102101004010111021222110100006000111114201040221003002212320-0101320-01112126101422101100--1210220204211202202010400001000012211[12]101202140101010020001100141111010[01]11-102001122221111112122006032200101300201 Euspinolia 201200131120212020230--12004100000000111[01][12]122[01]0101[01]00[012][02]00011411720104323000300221032100101320-00110115001420001200--1210411000110122[02]10[013]0011000112[01]01221[12]100121214[02]10200002000110001[12]1[12][01]000011-102101122021011012121000035110101300201 Glossotilla 201200102100212110120--3200110010102011002102101111002500111111020107421100300221002200101320-0[01]110204001021011220--02001202012112020113201111101110122112111202143100011020001120123111120111-102[01]001220210110120-1050007222101300201 Gogoltilla 2012001022302121201210011004100000010111021212111120010--011311320108021100300221002210101320-00111126001222111200--00002110101112022103103000010100122131001232143101010020001100002221010-11-1031021220010110120-200503?201101300201 Hemutilla 201200101220212110120--130011000000101111210120100000[01]20001131162?105??[01]0003002221[01]20-0101320-00111115001321111200--03001[12][01][134]0[67]211202211110140000110012210203120215???[01]?[02]0020?01100001[12]21010-11-102101122111111012[01][12]20[01]0044200101300201 Hindustanilla 101210101000212120120--12000000000020114120200210105129100103111201054011003002200110-00-122110010011000102[01]121200--1200010000211-200--------------11--0--0012[01]214300001002003--------------------------000010012122004053121101300210 Hoplocrates 20120014[12]2202120212320[23]32004[01]0020401012102122[12]01000002[07]000[12]131162010432201030[12]2[12]10221[01]0101320-0[01]11012500122001120[01]0[12]2200411014312022[12]01115[12]1000000001221310012521[45]2101000[01]20001000141[12]41020-11-1020011220221110120-2000030110101300200 Hoplognathoca 201200122232212120232233200401200001012112121201000001100001011620104022100300221122210101320-00111126201421112200--02004113042112002121103400010000122102101242140101010020001000100111000011-1020001221210110120-1000030201101300201 Hoplomutilla 20120011123221211022210[01]200410000[06]010121121302010000002000[01]1211[56]201040211203022211222[01]0101[23]20-0[01]111126201[34]2210110[12]00[01]20042000[06]2111222021112400000200122121001242140100010[12]20001000033111020011-1010001221210110120-2010042200101300200 Karlissaidia 201200101100212010[12]2[01]003[23][01]011000000[12]011[04]02122[02]01111002601111[02]1102010402[12]10030[01]22[12]002200101320-01110102001[25]2[13]00121100020012[01][12]002112[12]22113[12]2[12]1111011[01]012211[12]1[02]12[01]214011001102000112022[13]111120111-1[01]100[01]1220[02]10110120-[01]05002722[01]101301201 Krombeinidia 201200101120212110121203[23]10110000002011002122211110002601111111020107421100300222002200101320-011101020014231112[12]0--121012020031130221132021111010[01]0122111101212140120111020101120121211120111-10[12][01]001220210110120-1000017221101300201 Kudakrumia 001120112000222210101001101?000004010114001100010120020---20310-201100100000110100010-00-1020-00100100001011211100--1100?1000410000000000000000000001100301012-1000000000000000100000001010000101001011100100-0020-1000032010101200200 Labidomilla 20120014210021202023[01]000200401000001011[04][01]21[02]20[12]1011302[06]01[01]11[123]1132010[45]021100300221032[01]00101[23]20-00110203001221111100--02004110[01]02112[02]221[01]110[01]200001[01]001221111012[01]21[45]01000[01]00200011311213[12]1010111-0[01]2001122021[01]110120-1020005211101300200 Liotilla 201200101100212220120--[01]301100000001011602110021010001610-213110201020210000002210020-010132[01]101110110021120222200--1201210000001-[12]0[02]--------------01--1--1012[01]212010000000003--------------------------021011012[01]120091-0110101300000 Lobotilla 201200101100212000220--3210120000001011402102201110012501111111020104422100300221002200101320-04110112001222001210--221012020021120221131211111011101221111012121431[12]0111020[01]01120221111120111-1110001220210110120-1000007221101301201 Lomachaeta 2012001012302122202220012112100000020111021202210000001000213113201050200003022211010-0101220-00111126001320210001001211210[12]00211201212310411000010012213-101232150100010020001101132121010-11-1020021221222114120-1006033200101300201 Lophomutilla 2012001112322121202322012104010100020101[01]2122201000010600[01]113115201050211000002221[03]2110101220-00111126001321111100--0200411000310322212[03]102400000000122132101242150100010[01]20001100001[12][12]1020011-002[01]021220[02]10110120-1000030200101300201 Lynchiatilla 201200101231212120121001210410000001011102122201011410100011[23]11520105021100300221102200101320-001121261013211[01]1100--110042000021120221231024000100001221[12]11012521501000100200011000101[12]1010011-1020011221210110120-1006032200101300201 Mickelomyrme 201200101100212110110--33001100000020110021022111110027--11121132010402[12]100300221002100101320-00110[01]0400152[23]1[01]0[01]00--[01]20012020031[01]30221[01]320[12]1[12][01]10100012211[12]101212140110110020101220121111120111-1[01]11001220[02]1011012121000037222101300100 MimecomutillaSS 201200102000212110120--0210110000102011002102211111002500111111020106421100300221032[02]00101320-0011020[34]001122100100--02101[12][01]2012113022102152110001100122112111212153121210020101120121111120111-1[01]21021220210110120-1000002212101300200 Mimecotilla 201200102100212110120--12[012]0110000102011002122211011002500111[01]11[03]20106[34]21100300221002200101320-0[01]110[12]0300112[12]100100--[01]20012000121130221000521100011001221[01]21112121[45]3120210020101120121111120111-1121021220210110120-1000002212101300200 Mutilla 201200112[01][02]021210012[01]00121010[01]00070101100212200101100250[01]11111132010402[12]10030122[01]0320-0101120-0111010[34]001[12]2[01]101000--021011100721[01]30221130521211[01]11[01]01221111012[01]21[45]0100011020001[01]20121111120011-1110001220211110120-1000036222101300200 MyrmillaSS 2012001421[02]0212010230--02003010100010110021222110010026010111113201050200003002210320-0101220-00110205001[13]21001200--120031100[02]2113[02]221211011000011001221111012121501000100200011311[24][13]021110-11-112001122021011012112037062221101300200 Myrmilloides 2012001421312121102320310004013000010111021212210000020--011111620105021100300221102210101120-001112160014201113020310004203002112222--------------01--1--10125215010001002001--------------------------1210110120-2008070110101300201 Myrmosa 0011201[13]0020012110110--130000000040[12]0111000200010010010--000310-201100120000111100003100-1020-00110000001120001200--130000100421002000100001000001101100201002-10000040[01]000000010000000000[01]000101001001100100-0120-01-0032010101200000 Myrmosula 001120130000212120110--22000102004010131000000010100010--020310-201100110000100000002100-1020-00110200001010111100--020001100410000000000000000000001100200002-1000001000000000100000000000000101001001100100-0120-0000032110101200200 Nanomutilla 001210111020212210100--1200000000001011[04]02020021012212910-20311320105000000300[01]200020-00-0020-0010011000132121[01]100--120001000011000000[12]1000110002[12]101110301012321500000000200011000[02]0[12][24]1021-10000200212200100-0120-1002060111101300210 Nemka 201200102000212110120--1300210000001011002122201111002500111211020104022100400222031100101320-01110112011223000100--02001112002113022103201111101110122112111202140101[01]10030[01]01220121111120111-111000122021011012122000037222101300100 Odontomutilla 2012001021[02]0212100112[12][02]4320[23]10000002011112122001011[02]0040011111132010502[012]000[13]0[12]2210[012]1[01]0012112[01]0011101[01]400122110000[12][01][02]3320110000[23]1132[02][01]1[12]313212110111[01]1221111[02]1202150100030020001031133[12]21120-11-112102122021[01]120121[01]100002[24]201101300200 Odontomyrme 20120010113021221010100322021000000201110212220101120060001111142010402[12][01]003012202020-0101320-01112126001323211100--0210220000210302212310111000010012211110120214010[01]000020001100121111000011-102001122[12]11111012112006032201101300201 Odontophotopsis 201200101232212110120--13003100000020111021[02]2211011000200011[23]110201040220003002[23][12]2320-1101320-01112123211421001[23]00--[01]20011[01][01]04310[12]0221[01]010020000[01]0101221[12]2[01]112[02]214[03]101000020001200001[14][12]1010011-1021021222[01][12]011112[01]22000034201101300201 OdontotillaSS 201200132100212010231000210400000001011402122221000612601011211420104021100301220012100101320-01110210001120002300--02104110102112222--------------11--1--10127214010101002002--------------------------0211110120-2000005211101300200 Orientilla 2012001111202121101110012101[12]00000010111020[02]02110100001000213112201050200[01]03012320[23]2[01]000-112110[01]100115001[03]20200000--1210110010211200011[13]1[03]11[01]00001001221[23][12][01]012221501000100200010000[34]11111[12]0[01]11-10[12]00[12]12201210-012122000022200001300200 Paramyrmosa 001120130020012110110--01000100004010111000200010110010--020310-20110011000011110000[03][01]00-0020-00110000001020001200--020001000410002000000000000001001100301002-1000004000000000100000000000000101001001100100-0020-01-0032110101200000 Pertyella 201200141232212010232010000401[03]001020111121222010000120--011[23]1162010502112030[01]221112[12][01]0101120-00111126101321111100--00004110013112022121102400000000122121101242150100010020001100001111010011-1120011221220110120-1000030200101300201 Pherotilla 20121011102021220010[01]0001012100200[12]0[12]112121[12]002100020[01]6000113115201050010010012100010-0[01]30320-00110112001420[12][01]1[12]00--0[13][01]1210300[12]10[23]011112150111002[12]101221[012]11212322[03]0010100000101000000[12][12]1110111-10101[01]1220111[01]0312[01]020830502[01]100140[01]210 PhotomorphusSS 201200101121212110110--13003100000020111021212110110011000113116201054220003002211020-0101320-00111115101321111100--12002202083112022113100400010010122111011222150101000020001100110421010-11-102102122141111012122010032200101300201 Physetopoda 201200101000212[01][01]0120--1300100000002011002122201111002500111111020104021100301222002100101320-001101[01]20[01]1[45]23[01][01]1200--02001[12]02002113[02]22103[12]0111110110012211[12]1012[12]21401000100200011[13]0[12][23]1111120111-102102122021[01]110120-100003[67]220101300100 Platymyrmilla 2012001410202120102320002003010100010115021222110110026010110117201054201003002210320-0101220-00110115001120001200--12003010001112022101101100001100122112101282150100010020001131221021110-11-112001122021011012112020022211101300200 Pristomutilla 20120010110021201012[01]003310[23][01]0000[01]01011002122211011402500111[01]1102010[47]0221003002210[03]2200101321[01]001101[01]3001422100000--100022000021[01]30221131021100100001221[01]11012821[24]0100010020001120121[01][12]1120111-1[01][12]0001220210110120-10[02]0002212101300200 Promecilla 20120010110021211012120[01]310110000002011002122201[01]01001[05]0[01]111[123]11020104421100300222002[01]00101220-011101140[01]1422[01]01200--0210110400311302[12]1[01]31[05][12][01]111011001221121012121[45][03]1[01]0[01]10020[01]0122012[01]111110111-1111001220010110120-1000027222101300100 ProtophotopsisSS 2012001012302121100110012003100000020112021222010100001000113113201044210003002201310-0101220-00111126001421200000--03002100002111022121001110000100122121101282150100000000001130020321010-11-013000122131111312121000032200101300200 Pseudocephalotilla 201200101100212100122101310100000002011002122[02]01100002601111111020106422100300221012100101320-01110114001[45]2310010[01]000200110200[23]1130221[01]3[12][05]111110[01]1[01]01221[01]11012[12]212[03]10[01]010020001[12]20121111020111-1110001220210110121[12]1000032222101300200 Pseudomethoca 2012001[234]113[12]2121201[23][12]0[01][13][0123]004[01]0000[01]020111[01]212[02]20101100[12][016]00011[23]11[56]2010402[12]100300221102[02]10101320-00111126[01]01[03]2[01]11110[01]00[01][02]004[12]000031[01][12][01][02]2121[12]0[1234]200000[01]001221[12]1[01]012421[45]01000[01]0020001[01]0000[01][012][12]1010011-[01]0200[12]1221210110120-1010030200101300201 Pseudomutilla 201200142120212110230--0200301010001011[04]021222[12]100100260101111132010502[01]1003002210320-0101220-001102050011201[12]1200--12003[12]10001113[02]22--------------01--1--10123215010001002002--------------------------0210110120-2037062221101300200 Pseudomyrmosa 001120111[01]20212220110--12[01]00000004010111000000010100010--0203103201100110000100000000-00-1220-00100110001010111100--030001000410[01]0000000000000000000110020[01]0023100000100000000010000000[01]0[01]0000101001001100100-0120-2000032010101200200 Pseudophotopsis 211211100001[012]12110120--1200000[01]0000[01]010501000100000002300010310120013222[01][01]02002210310-01010211021[01]0010001[34]20111200--1200010[12]001100022004000100000000110021101212000131300010[23]01100001020000-0011110101110[01][01]01101212200[05]031010101210200 Radoszkowskitilla 2012001011202120001212042[01]0110000401011002122201111001501111111020107021100301221002200101320-01110112001423001210--121012000421122221131021111111101221111012[16]2150120011020001120221111120111-111[01]001220211110120-1050026221101300201 Reedomutilla 201200101230212100110--1[23]001100000010121021102010000010--0111[01]132010[45]0220003022421[13]2200101320-01111126001[35]23101100--0[23]0011000021230221231530000101001221[12]-0[02]12[03]214010[01]0[01]00200011001331[12]10[01]0011-102002[01]221322110120-2005033200101300201 Rhopalomutilla 2012101110[02]0212200100--02012100200[12]021120200001100000160002131152010500100100121[01]0010-0130320-00110112001[05]2010110[012]000221210300110[123][01][01][01]11215011[01]002210122121121232230010100000101[01]000001[12]1110111-[01]010[01]11220[13]210-3120-20830502[01]1011401210 Rimulotilla 201210111000212200101000211220000020[12]11202010021101201[06]000112115201050110010012100000-00-0320-00110112001420101200--0211220300210121011[02]1501100022101221211012322[03]0010100000101100000111100111-10100012201210-[23]120-2002000311001420210 Ronisia 20120011[02]100212010120--3200110000001011502122201011002600111111020104323100301221012100101320-01110013001222001100--12001200002113022113001121111100122112101212143101011020001020120111120011-1110001220211110120-1000036221101300200 Scaptodactyla 201200101232212120120--43003100000020111021020010110001000111113201040230003002212320-1101320-00112123111320001200--02002100002102022113000100001000122132001202153100000020001100003221010-11-1031021222121111120-2000044202101300200 Seyrigilla 201200111220212110120--12101100000010111020000110000001000213112201054200003012120321000-1121100110115001020100000--12001100102102002112151210000100122132001222150100000020001000031111120011-10100112201210-012122000032200001300200 SmicromyrmeSS 2012001020[02]0212110120--1300100000003011002122201111002500111111020104021100300221002100101320-00110102001423110100--02001[12]0200311302210320[12]1111011001221121012[01]2140110110020101120121111120111-10[12]10[012]122021[01]11012111000037220101300200 Smicromyrmilla 1012101210002121[12]01[12]0--1200000000001011[04]120200210[01]0[125]12[69][01]102031[01]320105401100300[12]200[13]10-00-1[023]21100100110001[12]2[12]111[12]00--120001000021[01]0200020040110[01]0[12]0111110[23]0[01]0121215[03]000010020001011131121011-100011002122001010012122004053121101300210 SphaeropthalmaSS 201200111121212110120--13001102000010111021102010000001000113110201040210003012222120-1101220-01111125101421001200--020011140021[01]10220001014000111001221[23]21012121431000000200011000012110[01]0011-1[01]20021222121111120-2000044202101300201 Spinulomutilla 201200102100212000121003210110020002011012112201010012501111[01]110201074211003012210320-0101320-0211021400112[12]001210--121012000421120211131[05]11111011[01]012211[12]101212140100011020001120221111120111-1110001220210110120-1050017222101300201 Stenomutilla 201200111120212[01]20[12][23][012]00[01]210110000001011102000211000000100021311220100020[01]0030123[12]0[13]20-00-112110[01]110115001020[12][01][01][123]0[01]00021011[01]0[01]021[01][12]0[02][02]1[12][123]1[05]110000010012213[12]0012221[05]01000[01]0020001[01]00041111[01][12]0[01]11-10[12]00[02]12201210-012122000032200001300200 Sulcotilla 2012001521202121101210033001100001010110021221111100027--111311020104022100400222032100101320-01110214001123100100--0200111201211302210320111110110012211101120214010101003000112012[01]111120111-111000122021011012112000037222101300100 Tallium 201200101120212110120--1200210100001011102122211011001800111[23]113201040210003002201[123]2[01]00101320-0011[12]115001321100100--0200220200211102201[13]101[12]000011001221[23]1011212140101000020001200001221010-11-102[01]02122[12]21[12]1101211[12]01[06]032200101300200 Timulla 201200102100212000220--321011000000[12]0110021[012][02][02]011[01]100260111111102010[47]022100300221032200101320-00110102001122001210--0200120[23]002112022103102111111100122111121212140120[01]11020[01]01120121111120111-1[01][12]0001220210110120-1050027221101301200 Tobantilla 2012001012302121201211003002200000020111021[12]12111120010--011111520108021100302232132210101320-00111026001223100100--02002200062113022113203010010100122111001232143100010020001100011221010-11-1020021221322110120-2005003200101300201 Tricholabiodes 2012001001[02]2212[12]101[12]0--13001100000010111021[02]2221100001[12]00021111020104021[01]0010[01]2220020-0101320-01110123211[34]21001200--1300111200310[12]022100[12]011000010001221[23]21012121501010[012]00200012000[023]11[12]1010011-102001022012111012[01]11000032200[01]01300200 Trispilotilla 20120010210021210012100321011000000[12]0110021221010000025001111110201044211003002210[01]2200101320-00110[12]02001222111210--22101200002112022113121111101[12]101221111[02]12[01]214[03]100011020001120121111120111-101000122021011012[01]11000027221101301001 TrogaspidiaSS 201200102100212000220--32101100000010110021[12]0001111002501111111020104021100300221002200101320-0[01]110102001222001210--02[01]012020[04]211202211312[12]1111011101221111212121401[12]0[01]1102000112012[13]111120111-10[12]0001220210110120-105002722[01]101301201 Tropidotilla 2012001[01]10[02]02120001[23]210321011000000[12]01100212220101100260011111102010402110030[12]2210320-0101320-01110104001[12]22[01]0[01][12]00--020012030021[01]302211305[12]12110110012211[12]1[12]12121401[012]0[01]11020[01]011[23]01211111[12]0011-10[12]000122021[01]110120-10[24]0036221101300200 Vianatilla 2012001322322120002222250004100200010111121222010010020--011311620105021120301221112210101220-001111260013221[01]1[12]00--[02][03]00421000311302212110340000100012213-1012521[45]010001000000100000[01][12][12]1010011-102001122101011012121000035200101300201 Viereckia 2012001[013]2000212110120--1200310010201011[05]021222[012]1011102601111111320104021100300221002210101320-0011010[35]001[13]2[02][12][01][01][01]00--120032001[02][12]1[01]2[02]221[01]1101100001[01]001221121012[13]21401000100200[03]1100121[13]11[01]20111-1[01]20011220[02]10110120-1000002212101300200 Wallacidia 201200102100212000221003310110000002011002122201111002601111211020107422100300221002200101320-01110102001223[01]01210--0210120300211212211312[12]11110111012211112121214012011102010112022[13]1111[12]0111-1010001220210110120-1050027221101301200 Xystromutilla 201200111[12][23][01]212110120--1300110[13]0000[23]0111021202210000010---2131102010[45]0210200012[23]1232[01]01101220-0111[12]1[12][35][01][01]1321001300--[01]200110[23]00311[12][02]2201[03]10140000[01]00012213110121214010000012000110000[01]2210[01]0-11-10[12][01]0112221[12][02]11112122005042201101300201 Yamanetilla 20120010212021210011222432031000000301111212200101100040011111132010502[01]0003022210010-0121120-0111010400122[01]11010212332011000031132001[12]3132121101100122111101212150100030020001031131421120-11-1131021220212120121[01]1000024201101300200 ; cc - 3 7 10 13 22.23 30.31 33 38.39 51 54 60.61 63 66 68.70 75 78.79 82 84 88 90 95 101 103.105 115.116 119 123 125 135 137 141 144 150 154.155 158 160.163 167 170 176 179.181 201 205.206 213.214 216.218 224.225 227; cc + 0.2 4.6 8.9 11.12 14.21 24.29 32 34.37 40.50 52.53 55.59 62 64.65 67 71.74 76.77 80.81 83 85.87 89 91.94 96.100 102 106.114 117.118 120.122 124 126.134 136 138.140 142.143 145.149 151.153 156.157 159 164.166 168.169 171.175 177.178 182.200 202.204 207.212 215 219.223 226 228.229; proc /; # $ ; cn {0 Both_Eye,_pubescence_and_pores 0=Both_present 1=Pubescence_absent,_pores_present 2=Both_absent; {1 Both_Pronotum,_ventral_pubescent_pit 0=Absent 1=Present; {2 Both_Metasternum_level 0=Not_depressed 1=Depressed; {3 Both_Metasternum,_form 0=Simple_and_flattened 1=With_V-shaped_carina,_arms_leading_to_metacoxae 2=With_posterior_median_process(es); {4 Both_Metacoxa_postero-dorsally 0=Simple 1=With_carinate_tubercle 2=With_lamellate_process; {5 Both_Sternum_I_lateral_rounded_pubescent_pit 0=Absent 1=Present; {6 Both_Tergum_II_and_sternum_I 0=Not_articulated 1=Articulated,_tergum_II_overlying_lateral_extremities_of_sternum_I; {7 Female_Head,_shape 0=Normal,_transverse,_rounded_posterolaterally 1=Normal,_long,_rounded_posterolaterally 2=Broad,_transverse,_rounded_posterolaterally 3=Broad,_long,_rounded_posterolaterally 4=Broad,_long,_rectangular_posterolaterally 5=Broad,_transverse,_protuberant/angular_posterolaterally; {8 Female_Occipital_carina 0=Distinct_and_reflexed,_complete 1=Distinct_and_reflexed,_dorsal_only 2=Absent_or_not_reflexed_and_scarcely_discernible; {9 Female_Eye_form 0=Weakly_convex,_following_head_contour 1=Moderately_convex,_distinct_from_head_contour 2=Strongly_convex,_disjunct_from_head_contour; {10 Female_Eye_shape 0=Oval,_inner_margin_more-or-less_convex,_long_axis_vertical 1=Oval,_inner_margin_obviously_sinuate_or_emarginate 2=Subcircular,_inner_margin_convex,_long_axis_vertical 3=Subcircular,_inner_margin_convex,_long_axis_horizontal; {11 Female_Eye_surface 0=Ommatidia_distinct 1=Ommatidia_faintly_distinguishable 2=Smooth,_ommatidia_not_distinguishable; {12 Female_Ocelli 0=Present,_functional 1=Present_but_rudimentary 2=Absent; {13 Female_Antennal_socket,_rim 0=Simple 1=Dorsally_expanded_as_“tubercle” 2=Frontal_ledge_overhanging_socket; {14 Female_Scape_radicle 0=Annular_differentiation,_not_angled 1=Annular_differentiation,_angled 2=Flangelike_expansion,_angled; {15 Female_Pedicel_length_I 0=Very_short,_<0.4X_length_of_flagellomere_I 1=Short,_>0.4<0.7X_length_flagellomere_I 2=About_as_long_as_flagellomere_I; {16 Female_Pedicel_shape 0=Shorter_than_wide 1=As_long_as_wide 2=Longer_than_wide; {17 Female_Flagellomere_number 0=10 1=11; {18 Female_Flagellomere_I,_length 0=<1X_length_of_flagellomere_II 1=1-1.5X_length_of_flagellomere_II 2=>1.8X_length_of_flagellomere_II; {19 Female_Flagellomere_I,_shape 0=Wider_than_long 1=About_as_long_as_wide 2=>1.3<2.0X_as_long_as_wide 3=>2.0X_as_long_as_wide; {20 Female_Head,_genal_carina 0=Absent 1=Present_but_weak 2=Strong; {21 Female_Head,_genal_carina_extent_(absent=-) 0=Ending_distant_from_hypostomal_carina 1=Ending_adjacent_to_hypostomal_carina 2=Continuous_with_hypostomal_carina; {22 Female_Head,_genal_armature_(absent=-) 0=Carina_simple 1=With_small_lamellate_tooth_anteroventrally 2=With_strong_conical_or_pyramidal_tooth_anteroventrally 3=With_teeth_posterodorsally_and_anteroventrally; {23 Female_Hypostomal_carina 0=Complete,_simple 1=Complete,_flangelike 2=Complete,_with_distinct_tooth_laterally 3=Complete,_with_tooth/tubercle/elevation_at_about_midlength 4=Strong_anterolaterally_but_obsolete_posteriorly 5=Strong_posteriorly_but_absent_anterolaterally; {24 Female_Pleurostomal_carina 0=Absent 1=Slight,_to_inner_mandibular_edge 2=Distinct,_with_hypostomal_carina_curved_to_outer_mand_artic 3=Distinct,_with_hypostomal_carina_straight_to_outer_mand_artic; {25 Female_Postgenal_carina/ridge 0=Absent 1=Distinct,_separate_from_hypostomal_carina 2=Distinct,_merging_with_hypostomal_carina; {26 Female_Postmandibular_carina_(ventral/posterior) 0=Absent 1=Present,_simple_blunt_ridge; {27 Female_Oral_and_mandibular_fossae 0=Continuous,_junction_about_half_mandible_width_or_more 1=Continuous,_junction_much_narrowed 2=Separated_by_anteriorly_unfused_depressed_cuticular_bridge 3=Separated_by_fused_much-depressed_cuticular_bridge 4=Separated_by_fused_superficial_cuticular_bridge; {28 Female_Mandible,_apical_teeth 0=Three 1=Two 2=One; {29 Female_Mandible_shape 0=Apically_not_expanded 1=Apically_expanded; {30 Female_Mandible,_ventral_basal_expansion 0=Absent 1=Present,_toothlike 2=Present,_flangelike,_apically_abrupt 3=Present,_flangelike,_apically_oblique; {31 Female_Mandible,_inner_basal_tooth 0=Absent 1=Present,_acute 2=Present,_mediobasal_obtuse_flange; {32 Female_Labio-maxillary_complex 0=Short 1=Elongated_prementum_and_stipes; {33 Female_Prementum 0=Evenly_convex_or_weakly_medio-longitudinally_carinate 1=With_posteromedian_domelike_tubercle/elevation 2=With_sharp_posterior_median_elevation 3=With_anteriorly-indented_posterior_domelike_elevation 4=Flattened,_depressed_to_weakly_concave 5=Longitudinally_convex_w_deep_narrow_anteromedian_groove 6=With_strong_long_narrow_median_carina 7=With_paired_medial_longitudinal_carinae; {34 Female_Maxillary_palpus 0=Six-segmented 1=Two-segmented 2=Unsegmented; {35 Female_Maxillary_palp_length 0=Shorter_than_foretibia 1=1-1.5X_length_foretibia 2=>1.5<2X_length_foretibia 3=>2X_length_foretibia; {36 Female_Labial_palpus 0=Four-segmented 1=Two-segmented 2=Unsegmented; {37 Female_Wings 0=Present 1=Absent; {38 Female_Flattened_decumbent_setae_on_mesosomal_dorsum 0=Absent 1=Laterally_flattened,_slender_arcuate 2=Laterally_flattened,_broad_lanceolate 3=Dorsoventrally_flattened,_slender_arcuate; {39 Female_Mesosoma_form_(winged=-) 0=More_or_less_parallel-sided 1=Mesopleuron_protub_well_ant_to_meta_spiracle,_propod_narrower_than_prothorax 2=Mesopleuron_protub_just_ant_to_meta_spiracle,_propod_narrower_than_prothorax 3=Ovate,_propodeum_about_as_broad_as_prothorax 4=Propodeum_much_broader_than_prothorax 5=Mesopleuron_dorsally_concave,_pronotum_broadest 6=Pronotum_broadest,_evenly_narrowed_posteriorly; {40 Female_Mesosoma,_dorsolateral_margin 0=Smooth,_sinuate_or_weakly_tuberculate 1=With_distinct_teeth; {41 Female_Pro-mesonotal_suture 0=Distinct_and_freely_articulating 1=Distinct_but_fused,_not_articulating 2=Obliterated_or_very_indistinct_and_fused,_not_articulating; {42 Female_Pronotum_lateral_length 0=About_as_long_as_dist_betw_pronotal_and_propodeal_spiracles 1=<0.8X_distance_between_pronotal_and_propodeal_spiracles; {43 Female_Pronotum,_humeral_angle 0=Rounded 1=Abrupt 2=Vertically_carinate_or_dentate 3=Prominent_tooth_or_spine; {44 Female_Pronotum,_dorsolateral_setose_area 0=Absent 1=Present,_dispersed_patch 2=Present,_clearly_delimited_tubercle/tuft; {45 Female_Pronotum,_anterodorsal_setose_area 0=Absent 1=Present,_dispersed_patch 2=Present,_clearly_delimited_tubercle/tuft; {46 Female_Pronotum,_posteroventral_margin 0=Distinct_and_complete 1=Indistinct_or_interrupted 2=Obliterated; {47 Female_Meso-metanotal_suture 0=Distinct 1=Absent_or_very_indistinct; {48 Female_Mesosoma,_scutellar_scale_(winged=-) 0=Absent 1=Present; {49 Female_Propodeum 0=>0.6X_as_long_as_wide 1=<0.6X_as_long_as_wide; {50 Female_Propodeum,_posterodorsal_margin_form 0=Smoothly_rounded 1=Abrupt_but_not_ridgelike 2=Carinate_or_ridgelike; {51 Female_Propodeum,_posterodorsal_margin_dentition 0=Smooth_or_tuberculate 1=With_one_weak_median_spine_or_vertical_tubercle 2=With_two_lateral_spines_or_teeth_only 3=With_three_spines 4=With_more_than_three_spines 5=With_two_median_teeth_and_two_lateral_spines_or_teeth 6=With_two_large_sublateral_cylindrical_spines; {52 Female_Propodeum_posterolateral_margin 0=Smooth_or_tuberculate 1=Dentate_or_spinose; {53 Female_Mesopleuron,_dorsal_region 0=Strongly_protuberant_ 1=Weakly_convex 2=Depressed; {54 Female_Mesopleural_ridge_(usually_setose) 0=Absent 1=Indistinct_and_joined_to_mesonotal_tubercle 2=Strong_and_joined_to_mesonotal_tubercle 3=Joined_to_pronotal_spiracle 4=Ventrally_only,_narrow_dorsal_ridge_to_mesonotal_tubercle 5=Ventrally_only,_narrow_dorsal_ridge_to_pronotal_spiracle 6=Present_only_ventrally 7=Dorsal_ridge_to_pronotal_spiracle,_ventrally_evanescent 8=Indistinct_and_joined_to_pronotal_spiracle 9=Fine_ridge_approaching_pronotal_spiracle; {55 Female_Mesopleural_ridge,_ventral_section_position_(absent=-) 0=Anterior_to_midpoint_of_mesocoxa 1=Dorsal_to_midpoint_of_mesocoxa; {56 Female_Mesopleural_ridge,_ventral_section_form_(absent=-) 0=Blunt 1=Sharply_carinate; {57 Female_Meso-metapleural_suture,_direction_(indistinguishable=-) 0=Weakly_curved_(separate_from_mesopleural_ridge) 1=Strongly_angled_(joining_mesopleural_ridge); {58 Female_Meso-metapleural_suture_development 0=Distinct 1=Distinct_ventrally_only 2=Obliterated_on_surface; {59 Female_Meso-metapleural_"bridge" 0=Absent 1=Present; {60 Female_Metapleural-propodeal_suture_development 0=Entirely_distinct 1=Obliterated_dorsally,_distinct_ventral_to_endophragmal_pit 2=Obliterated_dorsally,_vague_ventral_to_endophragmal_pit 3=Entirely_obliterated_on_surface 4=Distinct_dorsally,_obliterated_ventral_to_endophragmal_pit; {61 Female_Mesosternum_anterior_to_mesocoxae 0=Smoothly_rounded 1=Paired_transverse/oblique_carinae_(may_be_toothed_mesally) 2=Paired_lamellate_projections_mesally; {62 Female_Mesocoxae,_contiguity 0=Contiguous_mesally 1=Slightly_separated_mesally; {63 Female_Metasternum,_posterior_median_process_(absent=-) 0=Shorter_than_coxal_height,_tridentate 1=Shorter_than_coxal_height,_shallowly_bidentate 2=Shorter_than_coxal_height,_deeply_bidentate 3=Shorter_than_coxal_height,_unidentate 4=Longer_than_coxal_height,_tridentate 5=Longer_than_coxal_height,_unidentate_acute 6=Longer_than_coxal_height,_unidentate_obtuse 7=Shorter_than_coxal_height,_transverse_crenulate_ridge; {64 Female_Metacoxal_cavities 0=Open 1=Partially_closed 2=Closed; {65 Female_Tarsomeres,_apicoventral_median_ovoid_pulvillus 0=Absent 1=On_1st-4th_tarsomeres; {66 Female_Tarsal_claws 0=Mid-ventrally_toothed 1=Simple 2=Apically_deeply_bifid; {67 Female_Arolia 0=Present 1=Absent; {68 Female_Foretibia,_inner_(anterior)_secretory_structure 0=None 1=Broad_coarsely_setose_delimited_patch 2=Linear_to_oval_finely_perforated_depression 3=Vertically_elongate_groove/pore 4=Obliquely_elongate_groove/pore 5=Obliquely_oval_to_circular_pore 6=Obliquely_elongate_groove/pore_and_linear_finely_perf_depres 7=Two_apical_separated_obliquely_oval_pores 8=Linear_to_oval_finely_perforated_convexity 9=Basal_elongate/oval_and_separated_apical_round_pores; {69 Female_Foretibia,_outer_(posterior)_secretory_structure 0=None 1=Linear_to_oval_finely_perforated_depression 2=Vertically_elongate_groove/pore 3=Obliquely_elongate_groove/pore 4=Obliquely_oval_to_circular_pore; {70 Female_Forecalcar_blade 0=Linear_narrow,_margin_entire 1=Linear_narrow,_margin_finely_pectinate 2=Expanded,_longish_>0.4X_length_calcar 3=Expanded,_almost_square,_<0.4X_length_calcar 4=Concave,_narrow,_apically_expanded; {71 Female_Meso-_and_metatibial_articulated_spines,_mean_number 0=0-4 1=5-9 2=10-14 3=>14; {72 Female_Metacoxa,_mesally 0=Smoothly_rounded 1=Carinate; {73 Female_Metatibia,_apex_dorsally 0=Evenly_rounded 1=With_elevated_tubercle_bearing_spine(s) 2=With_cylindrical_process_bearing_spine; {74 Female_Metatibia,_posterior_(inner)_surface 0=Convex,_setose,_punctate 1=Flattened_and_broadened,_smooth_delimited_area; {75 Female_Metatibia,_posteroapical_secretory_structure 0=Absent 1=Present,_delimited_patch_of_dense_setae 2=Present,_linear_setose_felt-line-like 3=Present,_a_small_pore 4=Present,_a_deep_narrow_longitudinal_groove; {76 Female_Metatibia,_apical_spurs 0=Both_unmodified 1=Inner_modified_as_cleaner; {77 Female_Tergum_I_profile 0=Broadly_convex 1=Anterior_&_dorsal_faces_merging 2=Anterior_&_dorsal_faces_distinct,_bounded; {78 Female_Tergum_I_base 0=Simple 1=With_paired_vertical_ridges 2=With_paired_expanded_"auricles"; {79 Female_Tergum_I_shape 0=>=0.5X_length_TII,_gradually_broadened_posteriorly,_sessile 1=>=0.4X_length_TII,_parallel-sided_posteriorly,_discont 2=<0.5X_length_TII,_gradually_broadened_post,_sessile_on_2nd 3=<0.5X_length_TII,_gradually_broadened_post,_constrict_apically 4=<0.25X_length_TII,_entirely_parallel-sided,_discont; {80 Female_Tergum_I_apical_width 0=>0.75X_width_tergum_II 1=>0.5_<0.75X_width_tergum_II 2=<0.5X_width_tergum_II; {81 Female_Tergum_I/propodeum_pubescence 0=Simple 1=Some_brachyplumose 2=Some_fully_plumose; {82 Female_Tergum_I_apex,_pale_pubescent_markings 0=None 1=Median_pale_spot 2=Paired_pale_spots 3=Pale_band; {83 Female_Tergum_II_length 0=<0.75X_length_tergum_III-VI 1=0.75-1.25X_length_tergum_III-VI 2=>1.25X_length_tergum_III-VI; {84 Female_Tergum_II_discal_markings_number 0=None 1=Odd_number_(unpaired) 2=Even_number_(paired) 3=Broad_band; {85 Female_Tergum_II_discal_markings_composition_(absent=-) 0=Plumosity_only 1=Integumental; {86 Female_Tergum_II_apical_fringe_setae 0=Entirely_simple 1=Some_densely_plumose; {87 Female_Tergum_II_felt_line,_presence 0=Absent 1=Present; {88 Female_Tergum_II_felt_line,_form_(absent=-) 0=Linear 1=Broad_patch 2=Invaginated,_elongate_or_pitlike 3=Small_indefinite_anterior_patch; {89 Female_Tergum_III_stridulitrum 0=Absent 1=Present; {90 Female_Tergum_VI_form 0=Evenly_sculptured 1=Evenly_sculptured_except_apical_area_much_finer/smoother 2=With_smooth/ish_unbounded_longitudinal_median_area 3=With_distinct_bounded_pygidial_plate; {91 Female_Sternum_I_differentiation 0=Overlapping_sternum_II 1=Briefly_declivous_and_abutting_sternum_II 2=Depressed_posteriorly,_constricted_and_abutting_sternum_II; {92 Female_Sternum_II_felt_line,_presence 0=Absent 1=Present; {93 Female_Sternum_II_felt_line,_form_(absent=-) 0=Dispersed_traces_only 1=Distinctly_compact_and_linear; {94 Female_Sternum_VI_sting_aperture 0=Lateral_areas_differentiated,_sting_aperture_slitlike 1=Lateral_areas_dorsomesally_produced,_sting_aperture_circular; {95 Female_Sternum_VI_armature 0=Without_processes 1=With_pair_apical_processes_(apex_notched) 2=With_pair_acute_lateral_teeth_basally 3=With_two_pairs_lateroventral_tubercles 4=With_two_pairs_apical_processes/teeth_(apex_4-lobed); {96 Female_Gonapophysis_IX_sting_curvature 0=Weakly_convexly_arcuate_dorsally 1=Strongly_convexly_arcuate_dorsally,_apex_directed_downwards; {97 Male_Head,_width_across_mandibular_bases 0=<0.6X_head_width 1=>0.6X_head_width; {98 Male_Head_pubescence 0=Entirely_simple 1=Some_brachyplumose 2=Some_fully_plumose; {99 Male_Occipital_carina 0=Distinct_and_reflexed,_complete 1=Distinct_and_reflexed,_dorsal_only 2=Absent_or_not_reflexed_and_scarcely_discernible; {100 Male_Eye,_form 0=Weakly_convex,_following_head_contour 1=Moderately_convex,_distinct_from_head_contour 2=Strongly_convex,_disjunct_from_head_contour; {101 Male_Eye,_shape 0=Broadly_oval,_inner_margin_convex_to_weakly_sinuate 1=Broadly_oval_inner_margin_acutely_but_shallowly_emarginate 2=Broadly_oval,_inner_margin_acutely_and_deeply_emarginate 3=Subcircular,_inner_margin_sinuate_to_weakly_emarginate 4=Subcircular,_inner_margin_acutely_and_deeply_emarginate 5=Subcircular,_inner_margin_roughly_convex,_long_axis_vertical 6=Subcircular,_inner_margin_roughly_convex,_long_axis_horizontal; {102 Male_Eye_surface 0=Ommatidia_distinct 1=Ommatidia_faintly_distinguishable 2=Smooth,_ommatidia_not_distinguishable; {103 Male_Ocelli 0=Present,_normal 1=Present,_much_enlarged 2=Absent; {104 Male_Antennal_socket,_rim 0=Simple 1=Dorsally_expanded_as_"tubercle" 2=Frontal_ledge_overhanging_socket; {105 Male_Antennal_scrobe,_dorsal_tubercle/carina 0=No_tubercle,_no_carina 1=Weak_transverse_carina_only 2=Strong_transverse_carina_only 3=Secretory_tubercle_only 4=Secretory_tubercle_and_weak_carina 5=Secretory_tubercle_and_strong_carina; {106 Male_Scape,_radicle 0=Annular_differentiation,_not_angled 1=Annular_differentiation,_angled 2=Flangelike_expansion,_angled; {107 Male_Scape,_ventral_ridges 0=None 1=One_only 2=Two,_one_less_developed 3=Two,_equally_developed; {108 Male_Pedicel,_length 0=Very_short,_<0.4X_length_of_flagellomere_I 1=Short,_>0.4<0.8X_length_flagellomere_I 2=About_as_long_as_flagellomere_I; {109 Male_Pedicel,_shape 0=Distinctly_shorter_than_wide 1=About_as_long_as_wide 2=Distinctly_longer_than_wide; {110 Male_Flagellomere_I,_length 0=<0.6X_length_of_flagellomere_II 1=Subequal_to_flagellomere_II 2=>1.3X_length_of_flagellomere_II; {111 Male_Flagellomere_I,_shape 0=Wider_than_long 1=About_as_long_as_wide 2=>1.3<2.0X_as_long_as_wide 3=>2X_as_long_as_wide; {112 Male_Flagellomere_I,_ventral 0=Cylindrical 1=Weakly_flattened_ventrally 2=Strongly_flattened_and_broadened; {113 Male_Head,_genal_carina 0=Absent 1=Present_but_weak 2=Strong; {114 Male_Head,_genal_carina_extent_(absent=-) 0=Ending_distant_from_hypostomal_carina 1=Ending_adjacent_to_or_continuous_with_hypostomal_carina; {115 Male_Head,_genal_armature_(absent=-) 0=Carina_simple,_unarmed 1=With_small_lamellate_tooth_anteroventrally 2=With_strong_conical_tooth_anteroventrally 3=With_teeth_posterodorsally_and_anteroventrally 4=With_strong_short_lamellate_tooth_posterodorsally; {116 Male_Hypostomal_carina 0=Simple 1=Flangelike 2=With_tooth/tubercle/elevation_at_about_midlength 3=Strong_anterolaterally_but_obsolete_posteriorly; {117 Male_Pleurostomal_carina 0=Absent 1=Slight,_to_inner_mandibular_edge 2=Distinct,_with_hypostomal_carina_curved_to_outer_mand_artic 3=Distinct,_with_hypostomal_carina_straight_to_outer_mand_artic; {118 Male_Postgenal_carina/ridge 0=Absent 1=Distinct,_separate_from_hypostomal_carina 2=Distinct,_merging_with_hypostomal_carina; {119 Male_Postmandibular_carina 0=Absent 1=Present,_simple_blunt_ridge 2=Present,_broad_smooth_tubercle; {120 Male_Oral_and_mandibular_fossae 0=Continuous,_junction_about_half_mandible_width_or_more 1=Continuous,_junction_much_narrowed 2=Separated_by_anteriorly_unfused_concealed_cuticular_bridge 3=Separated_by_fused_much-depressed_cuticular_bridge 4=Separated_by_fused_superficial_cuticular_bridge; {121 Male_Mandible,_apical_teeth 0=Four 1=Three 2=Two; {122 Male_Mandible,_shape 0=Apically_not_expanded 1=Apically_expanded; {123 Male_Mandible,_ventral_basal_tooth 0=Absent 1=Present,_small 2=Present,_enlarged 3=Present,_lamellate_oblique 4=Present,_lamellate_abrupt; {124 Male_Mandible,_inner_basal_tooth 0=Absent 1=Present; {125 Male_Prementum 0=Evenly_convex_or_weakly_medio-longitudinally_carinate 1=With_posteromedian_domelike_tubercle/elevation 2=With_sharp_posterior_median_elevation 3=With_anteriorly-indented_posterior_domelike_elevation 4=Flattened,_depressed_to_weakly_concave 5=Longitudinally_convex_w_deep_narrow_anteromedian_groove 6=With_strong_long_narrow_median_carina 7=With_paired_longitudinal_carinae 8=Flattened_with_posterior_median_transverse_lamellate_projection; {126 Male_Maxillary_palp_length 0=Shorter_than_foretibia 1=>1<1.5X_length_foretibia 2=>1.5<2X_length_foretibia 3=>2X_length_foretibia; {127 Male_Labial_palp_mid_segments 0=More_or_less_cylindrical 1=Flattened_and_expanded,_asymmetrical; {128 Male_Laterally_flattened_decumbent_setae_on_mesosomal_dorsum 0=Absent 1=Present,_slender_arcuate 2=Present,_broad_lanceolate; {129 Male_Pro-mesonotal_suture_(indistinguishable=-) 0=Weakly_concave 1=Strongly_but_evenly_concave 2=Broadly_V-shaped_(laterally_straight,_mesal_third_curved) 3=Abruptly_V-shaped_(laterally_straight,_mesally_angled); {130 Male_Pronotum,_humeral_angle_above 0=Smoothly_rounded 1=Blunt 2=Vertically_carinate_or_dentate 3=Prominent_tooth_or_spine; {131 Male_Pronotum,_dorsolateral_setose_area 0=Absent 1=Present,_dispersed_patch 2=Present,_clearly_delimited_tubercle/tuft; {132 Male_Pronotum,_anterodorsal_setose_area 0=Absent 1=Present,_dispersed_patch 2=Present,_clearly_delimited_tubercle/tuft; {133 Male_Mesoscutum,_length_(apterous/brachypterous=-) 0=Short_anterior_to_tegulae 1=Extended_far_anterior_to_tegulae; {134 Male_Mesoscutum,_notaulus_(apterous/brachypterous=-) 0=Present_and_complete 1=Present_but_incomplete 2=Absent; {135 Male_Mesoscutum,_parapsidal_furrow_(apt/brachypterous=-) 0=Obvious,_complete 1=Obvious_posteriorly_only,_absent_or_scar_anteriorly 2=Obvious_groove_anteriorly_only,_absent_posteriorly 3=Much_reduced,_at_most_a_superficial_scar_anteriorly 4=Interrupted,_faint_groove_posteriorly,_superficial_scar_anteriorly; {136 Male_Mesoscutum,_posterolateral_margin_(apt/brachypterous=-) 0=Rounded 1=Lobed_but_flattish 2=Dentate_and_protruding_upwards; {137 Male_Mesoscutellum_(apterous/brachypterous=-) 0=Simple,_even_with_metanotum 1=Pulvinate_with_smooth_median_ridge 2=Posteromesally_produced_(conical)_with_smooth_median_ridge 3=Laterally_produced_as_curved_posteriorly_dentate_flange 4=Posteriorly_produced_and_overhanging_metanotum 5=Swollen,_discontinuous_with_metanotum; {138 Male_Axilla_posterolateral_dorsal_margin_(apt/brachypterous=-) 0=Rounded,_posterolateral_surface_convex_or_flat 1=Narrowly_rounded,_posterolateral_surface_concave 2=Carinate,_posterolateral_surface_concave 3=Flangelike,_apex_broadly_obtuse 4=Flangelike,_apex_strongly_dentate; {139 Male_Axilla_anterolateral_dorsal_extremity_(apt/brachypterous=-) 0=Evenly_rounded_(vertical_ridge_may_be_present_ventrally) 1=With_slight_vertical_ridge_broadening_ventrally 2=With_strong_evenly_developed_vertical_ridge_or_flange 3=With_blunt_tubercle_or_tooth_dorsally 4=With_abrupt_broad_vertical_flange_dorsally; {140 Male_Tegula,_length_(apterous/brachypterous=-) 0=Short,_round,_distant_from_transscutal_articulation 1=Slightly_elongate,_reaching_transscutal_artic_or_slightly_beyond 2=Elongate,_extending_well_beyond_transscutal_articulation; {141 Male_Tegula,_form_(apterous/brachypterous=-) 0=Evenly_convex_to_flattish_posteriorly 1=Posteriorly_recurved 2=Longitudinally_angulate_basally; {142 Male_Tegula,_free_posterior_inner_margin_(apt/brachypterous=-) 0=More_or_less_straight_or_convex 1=Distinctly_concave; {143 Male_Propodeum,_length_(apterous/brachypterous=-) 0=About_as_long_as_high 1=Much_shorter_than_high; {144 Male_Propodeum,_disc_(apterous/brachypterous=-) 0=Evenly_sculptured 1=Larger_basal_fields_and_smaller_distal_fields 2=Three_large_fields_bordered_by_strong_longitudinal_carinae; {145 Male_Propodeum_disc/declivity_(apterous/brachypterous=-) 0=Broadly_rounded 1=Distinct_but_merging 2=Abruptly_differentiated; {146 Male_Propodeum,_dorsolateral_margin_(apterous/brachypterous=-) 0=Rounded 1=Carinate_or_distinctly_angled; {147 Male_Propodeum,_posterolateral_margin 0=Smooth_or_tuberculate 1=Dentate_or_spinose; {148 Male_Prepectus 0=Articulating_with_mesepisternum 1=Fused_to_mesepisternum; {149 Male_Meso-metapleural_suture,_fusion_(apt/brachypterous=-) 0=Articulating 1=Immovable_although_not_fused 2=Partially_or_entirely_fused; {150 Male_Meso-metapleural_suture,_shape_(apt/brachypterous=-) 0=Almost_straight 1=Posteriorly_convex 2=Sinuate; {151 Male_Meso-metapleural_"bridge" 0=Absent 1=Present,_fused; {152 Male_Metapleural-propodeal_suture_(apt/brachypterous=-) 0=Entirely_distinct 1=Obliterated_dorsal_to_endophragmal_pit,_distinct_ventrally 2=Obliterated_dorsal_to_endophragmal_pit,_vague_ventrally 3=Entirely_obliterated_on_surface; {153 Male_Oblique_metapleural_suture_(apt/brachypterous/obliterated=-) 0=Running_anteroventrally_from_pit 1=Running_horizontally_from_pit 2=Running_anterodorsally_from_pit; {154 Male_Mesosternum,_anteromesal_to_mesocoxae 0=Smoothly_rounded 1=Paired_transverse/oblique_carinae_(may_be_mesally_toothlike) 2=Paired_lamellate_projections_mesally; {155 Male_Mesosternum,_midway_to_anterior_margin 0=Evenly_convex 1=With_distinct_paired_teeth_or_tubercles 2=With_distinct_paired_(separated)_transverse_carinae/ridges 3=With_paired_longitudinal_high_lamellae_acuminate_apically; {156 Male_Mesocoxae,_contiguity 0=Contiguous_mesally 1=Slightly_separated_mesally; {157 Male_Mesocoxa,_insertion 0=Large_basicoxite,_cavities_large_and_approximated 1=Large_basicoxite,_cavities_large_and_widely_separated 2=Reduced_basicoxite,_cavities_small_and_widely_separated; {158 Male_Metasternum,_posterior_median_process_(absent=-) 0=Shorter_than_coxal_height,_tridentate 1=Shorter_than_coxal_height,_shallowly_bidentate 2=Shorter_than_coxal_height,_deeply_bidentate 3=Shorter_than_coxal_height,_unidentate 4=Longer_than_coxal_height,_unidentate_acute 5=Longer_than_coxal_height,_unidentate_obtuse 6=Shorter_than_coxal_height,_tridentate_w_median_tooth_incised 7=Longer_than_coxal_height,_tridentate_with_median_tooth_acu 8=Shorter_than_coxal_height,_transverse_crenulate_ridge; {159 Male_Metacoxal_cavities 0=Open 1=Partially_closed 2=Closed; {160 Male_Tarsal_claws 0=Mid-ventrally_toothed 1=Simple 2=Ventrally_basally_lamellate,_distinct_apex_acute 3=Apically_deeply_bifid; {161 Male_Foretibia,_inner_(anterior)_secretory_structure 0=None 1=Broad_coarsely_setose_delimited_patch 2=Linear_to_oval_finely_perforated_depression 3=Vertically_elongate_groove/pore 4=Obliquely_elongate_groove/pore 5=Obliquely_oval_to_circular_pore 6=Basal_elongate/oval_and_apical_round_separated_pores; {162 Male_Foretibia,_outer_(posterior)_secretory_structure 0=None 1=Linear_to_oval_finely_perforated_depression 2=Obliquely_elongate_groove/pore 3=Obliquely_oval_to_circular_pore; {163 Male_Forecalcar_blade 0=Linear,_narrow,_margin_entire 1=Expanded,_longish_>0.5X_length_calcar 2=Expanded,_almost_square,_<0.4X_length_calcar; {164 Male_Foretarsomeres,_apicoventral_median_ovoid_pulvillus 0=Absent 1=On_4th_tarsomere_only 2=On_3rd_&_4th_tarsomeres 3=On_2nd-4th_tarsomeres 4=On_1st-4th_tarsomeres; {165 Male_Meso-_and_metatibial_articulated_spines,_mean_number 0=0-4 1=5-9 2=10-14 3=15-19 4=20-24 5=>24; {166 Male_Mesotarsomeres,_apicoventral_median_ovoid_pulvillus 0=Absent 1=On_4th_tarsomere_only 2=On_3rd_&_4th_tarsomeres 3=On_2nd-4th_tarsomeres 4=On_1st-4th_tarsomeres; {167 Male_Metacoxa,_mesally 0=Simple 1=Carinate 2=Dentate 3=With_setaceous_pit; {168 Male_Metatibia,_posterior_longitudinal_smooth_glabrous_ridge/carina 0=Absent 1=Present; {169 Male_Metatibia,_apex_dorsally 0=Evenly_rounded 1=With_elevated_tubercle_bearing_spine(s) 2=With_distinct_cylindrical_process_bearing_spine; {170 Male_Metatibia,_posteroapical_secretory_structure 0=Absent 1=Present,_delimited_patch_of_dense_setae 2=Present,_a_small_pore 3=Present,_a_deep_narrow_longitudinal_groove; {171 Male_Metatibia,_apical_spurs 0=Both_unmodified 1=Inner_modified_as_cleaner; {172 Male_metatarsomeres,_apicoventral_median_ovoid_pulvillus 0=Absent 1=On_4th_tarsomere_only 2=On_3rd_&_4th_tarsomeres 3=On_2nd-4th_tarsomeres 4=On_1st-4th_tarsomeres; {173 Male_Wings_and_tegula 0=Fully_developed 1=Brachypterous,_wing_exceeding_propodeum_apex 2=Brachypterous,_wing_shorter_than_propodeum_base 3=Apterous; {174 Male_Forewing,_extent_of_venation_(apterous/brachypterous=-) 0=Reaching_distal_margin 1=Ending_before_distal_margin; {175 Male_Forewing,_Sc+R_(apterous/brachypterous=-) 0=<0.5X_length_of_1st_abscissa_of_RS 1=Subequal_to_1st_abscissa_of_RS 2=>1.5X_length_of_1st_abscissa_of_RS; {176 Male_Forewing,_pterostigma_sclerotization_(apt/brachypterous=-) 0=Entirely_well_sclerotized 1=Sclerotization_reduced_anteriorly 2=Unsclerotized 3=Faintly_sclerotized; {177 Male_Forewing,_pterostigma_delimitation_(apt/brachypterous=-) 0=Complete_delimited_by_distinct_veins_or_complete_sclerotized 1=Vein_SC_lost/much_reduced,_pterostigma_not_delimit_basally; {178 Male_Forewing,_pterostigma_base_(apterous/brachypterous=-) 0=With_interruption/constriction_in_C_and_Sc+R 1=With_interruption/constriction_in_Sc+R_only 2=Without_interruptions/constrictions; {179 Male_Forewing,_pterostigma_shape_(apterous/brachypterous=-) 0=Elongate,_broader_than_base 1=Elongate,_as_narrow_as_base 2=Short,_broader_than_base 3=Short,_as_narrow_as_base 4=Very_short,_narrowed_from_base 5=Minuscule/absent; {180 Male_Forewing,_radial_(marginal)_cell_apex_(apt/brachypterous=-) 0=Acute 1=Blunt 2=Rounded 3=Obtuse_with_posterior_spur; {181 Male_Forewing,_vein_RS2_(apterous/brachypterous=-) 0=Absent 1=Present_and_complete,_basally_tubular/solid_nebulous 2=Present_and_complete,_(pigmented)_spectral 3=Present_but_basally_absent,_pigmented_nebulous/spectral 4=Present_as_short_stub_only; {182 Male_Forewing,_closed_submarginal_cells_(apt/brachypterous=-) 0=Three,_all_veins_tubular 1=Three,_vein_3r-m_nebulous 2=Two,_all_veins_tubular 3=Two,_vein_2r-m_nebulous 4=One; {183 Male_Forewing,_cell_1R1_(first_submarginal)_(apt/brachypterous=-) 0=Rudiment_of_crossvein_1r-rs_present 1=Rudiment_of_crossvein_1r-rs_absent; {184 Male_Forewing,_vein_RS_third_abscissa_bulla_(apt/brachypt=-) 0=Present 1=Absent; {185 Male_Forewing,_vein_RS_third_abscissa_course_(apt/brachypt=-) 0=With_distinct_angle 1=With_weak_angle 2=Straight_or_very_weakly_and_evenly_curved; {186 Male_Forewing,_cell_1S_(second_submarginal)_(apt/brachypt=-) 0=Sessile_anteriorly 1=Petiolate_anteriorly; {187 Male_Forewing,_crossvein_3r-m_(absent/apterous/brachypterous=-) 0=With_bulla 1=Without_bulla; {188 Male_Forewing,_jugal_lobe_(apterous/brachypterous=-) 0=Present 1=Absent; {189 Male_Hindwing,_basal_hamuli_occurrence_(apt/brachypterous=-) 0=Present 1=Absent; {190 Male_Hindwing,_basal_hamuli_position_(apt/brachypterous/none=-) 0=Dispersed 1=Basal_cluster; {191 Male_Hindwing,_apical_hamuli_(apterous/brachypterous=-) 0=<11 1=>10; {192 Male_Hindwing,_vein_RS_junction_with_vein_SC_(apt/brachypt=-) 0=At_acute_angle 1=At_right_angle; {193 Male_Hindwing,_crossvein_r-m_(apterous/brachypterous=-) 0=Distal 1=Proximal,_complete 2=Proximal,_incomplete 3=Absent; {194 Male_Hindwing,_vein_M_free_apical_section_(apt/brachypterous=-) 0=Present 1=Absent; {195 Male_Hindwing,_vein_Cu_free_apex_(apterous/brachypterous=-) 0=Present 1=Absent; {196 Male_Hindwing,_crossvein_cu-a_(apterous/brachypterous=-) 0=Present,_tubular/solid 1=Present,_nebulous 2=Absent; {197 Male_Hindwing,_vein_A,_free_apical_sectn_(apt/brachypterous=-) 0=Present 1=Absent; {198 Male_Hindwing,_anal_lobe_(apterous/brachypterous=-) 0=Moderate_incision 1=Shallow_definite_notch 2=Not_indicated_on_margin_(at_most_very_shallowly_sinuate); {199 Male_Hindwing,_jugal_lobe_(apterous/brachypterous=-) 0=Present,_large_with_incision_about_half_length 1=Present,_small_with_incision_nearly_to_base 2=Absent; {200 Male_Tergum_I/propodeum_pubescence 0=Entirely_simple 1=Some_brachyplumose 2=Some_fully_plumose; {201 Male_Tergum_I,_shape 0=Gradually_broadened_posteriorly,_>0.5X_length_2nd,_sessile 1=Gradually_broadened_postly,_>0.5X_length_2nd,_constricted 2=Gradually_broadened_postly,_<0.5X_length_2nd,_sessile 3=Gradually_broadened_postly,_<0.5X_length_2nd,_constricted 4=Parallel-sided_postly,_>=0.4_×_length_tergum_II,_discontinuous 5=Entirely_parallel-sided,_<0.5X_length_2nd,_discontinuous; {202 Male_Tergum_I,_apical_width 0=>0.75X_width_tergum_II 1=>0.5<0.75X_width_tergum_II 2=<0.5X_width_tergum_II; {203 Male_Tergum_I,_profile 0=Broadly_convex 1=Anterior_&_dorsal_faces_merging 2=Anterior_&_dorsal_faces_distinct; {204 Male_Tergum_II,_felt_line,_presence 0=Absent 1=Present; {205 Male_Tergum_II,_felt_line,_form_(absent=-) 0=Dispersed_traces 1=Linear 2=Invaginated; {206 Male_Tergum_II,_apical_fringe 0=Setae_simple_to_slightly_flattened_slender_arcuate 1=Some_setae_densely_plumose 2=Setae_apically_split 3=Setae_strong_and_curved 4=Setae_few,_long,_strong,_convergent; {207 Male_Tergum_III,_stridulitrum 0=Absent 1=Present; {208 Male_Sternum_I,_differentiation 0=Overlapping_sternum_II 1=Briefly_declivous_and_abutting_sternum_II 2=Depressed_posteriorly,_constricted_and_abutting_sternum_II; {209 Male_Sternum_II,_lateral_felt_line,_presence 0=Absent 1=Present; {210 Male_Sternum_II,_lateral_felt_line,_form_(absent=-) 0=Lateral_traces_only 1=Distinct_but_minute 2=Well_developed; {211 Male_Sternum_VII 0=Entirely_exposed,_about_as_long_as_sternum_VI 1=Partly_exposed,_much_shorter_than_sternum_VI 2=Concealed; {212 Male_Hypopygium_(sternum_VIII),_visibility 0=Almost_entirely_exposed,_lat_marg_entire_or_shallow_notched 1=Almost_entirely_concealed,_deeply_tri-_or_pentalobate; {213 Male_Hypopygium_(st_VIII),_sculpt_exposed_surface_(hidden=-) 0=Convex_to_flat,_+-evenly_sculptured,_punctate_to_smooth 1=Concave,_punctured_to_smooth_with_lateral_longit_carina 2=Evenly_convex,_median_tubercle_on_basal_half 3=Convex_mediolongitudinally,_abrupt_lateral_depression 4=Convex,_median_smooth_ridge 5=Convex_to_flat,_sublateral_paired_longitudinal_oblique_ridges 6=Convex,_median_Y-shaped_ridge 7=Longit_smooth_median_depression,_dense_punctures_laterally 8=Median_excavation,_lateral_peg-like_projection; {214 Male_Hypopygium_(sternum_VIII),_apex 0=Simple,_rounded_or_obtuse 1=Shallow_broad_median_emargination 2=Simple_deep_narrow_median_emargination 3=Broad_lobed_median_emargination 4=Deep_narrow_median_emargination_with_internal_sclerites 5=Median_tooth_or_peg 6=Two_small_approximated_teeth_or_slight_notch 7=Broadly_bilobed 8=Two_small_lateral_teeth 9=Two_separated_moderate_teeth; {215 Male_Cercus 0=Present 1=Absent; {216 Male,_cercus_form_(absent=-) 0=Elongate,_cylindrical_or_weakly_evenly_broadened_apically 1=Elongate,_strongly_clavate_(with_narrow_basal_stalk) 2=Elongate,_narrow,_flattened 3=Short,_base_narrow,_distinctly_flattened 4=Short,_base_widened,_apex_narrowed,_distinctly_flattened 5=Short,_flattened_basally,_clavate_apically 6=Short,_evenly_clavate 7=Vestigial 8=Broad-based_diskiform,_flattened; {217 Male_Gonobase,_form 0=Compl,_dors_&_ventr_lengths_similar,_long_as_paramer_base 1=Complete,_dors_&_ventr_lengths_similar,_very_short_annular 2=Compl,_dors_shorter_than_ventr,_ventr_long_as_paramer_base 3=Compl,_dors_&_ventr_lengths_similar,_longer_th_paramer_base 4=Compl,_dors_shorter_than_ventr,_ventr_longer_th_paramer_bas 5=Compl,_dors_shorter_than_ventr,_much_shorte_th_paramer_bas 6=Dors_incompl,_dors_short_thn_ventr,_ventr_long_as_paramer_bas 7=Dorsally_absent,_very_short; {218 Male_Gonostylus,_form,_lateral_view 0=Short,_lamellate_with_rounded_apex 1=Short,_tapered_with_narrow_or_acute_apex 2=Elongate,_tapered_with_acute_apex 3=Elongate,_lamellate_with_rounded_apex; {219 Male_Gonostylus,_curvature,_lateral_view 0=Apically_upcurved 1=Apically_straight 2=Apically_downcurved; {220 Male_Gonostylus,_dorsal_transv_suture_distant_from_gonobase 0=Present,_extending_at_least_halfway_to_lateral_margin 1=Absent_or_short,_longit_suture_distant_from_gonobase 2=Absent,_longit_suture_reaching_gonobase; {221 Male_Gonostylus,_parapenial_lobe 0=Absent 1=Present; {222 Male_Gonostylus,_dorsal_oblique_heavy_setae 0=None 1=Present,_arising_under_dorsal_flange; {223 Male_Gonapophysis_IX_(penis_valve),_fusion 0=Fused_dorsally_for_most_of_length 1=Free_for_most_of_length; {224 Male_Gonapophysis_IX_(penis_valve),_form 0=Apex_elongate,_rounded,_no_ventral_tooth 1=Apex_rounded,_ventral_tooth_about_midway 2=Apex_dorsally_produced,_ventral_tooth_about_midway 3=Apex_dorsally_simple,_ventral_tooth_on_apical_half 4=Apex_rounded,_produced,_ventral_prominence_about_midway; {225 Male_Gonapophysis_IX_(penis_valve),_articul_spines/long_setae 0=Absent 1=Present,_strong_short_spines 2=Present,_thick_long_setae; {226 Male_Gonapophysis_IX_(penis_valve),_right 0=Same_shape_as_left_gonapophysis_IX 1=Longer_and_more_elaborate_than_left_gonapophysis_IX; {227 Male_Volsella,_basal_lobe 0=Present,_distinct_prominent_inner_projection 1=Present,_rounded_ventral_lamellate_expansion 2=Absent; {228 Male_Volsella,_digitus 0=Present 1=Absent_or_scarcely_discernible; {229 Male_Volsella,_paracuspis 0=Absent 1=Present; ;