Synapomorphies supporting Zethus and relationships between its lineages A proepisternum of Z. spinipes, ventral view; arrowheads indicate lamellae B second metasomal tergum of Z. yarrowi, dorsal view; arrow indicates stem C pronotum of Z. rodhaini, lateral view D pronotum of Z. planiclypeus, lateral view E pronotum of Z. robustus, lateral view; arrow indicates dorsal suture F ocelli of Z. mandibularis G pronotal carina of Z. smithii, postero-dorsal view; arrowhead indicates notch H latero-posterior region of the pronotum of Z. rodhaini; arrow indicates pre-tegular carina I lateral portion of mesoscutum of Z. spinipes; arrow indicates discoid puncture J mesepisternum of Z. jurinei; arrows indicate dorsal (upper arrow) and mid (lower arrow) sections of epicnemial carina K third metasomal tergum of Z. fraternus; arrow indicates gradulus L second metasomal tergum of Z. aztecus; arrow indicates lamella. M third metasomal sternum of Z. fraternus; arrow indicates gradulus N tegula of Z. aztecus O ocelli of Z. spinipes P third metasomal tergum of Z. spinipes; arrow indicates interruption of gradulus. Scale bars: 1.0 mm.