Annotated checklist of the megachilid bees of Corsica (Hymenoptera, Megachilidae)

Romain Le Divelec; Alexandre Cornuel-Willermoz; Matthieu Aubert; Adrien Perrard

doi:10.3897/jhr.97.114614

Abstract

Corsica stands as one of the largest Mediterranean Islands and has been the exploration ground for renowned entomologists like Charles Ferton. However, no synthesis on Corsican bees has been published so far. To fill this gap in knowledge, we propose an overview of the megachilid bee fauna of the island based on fieldwork, a thorough examination of material housed at the Muséum national d’Histoire naturelle (MNHN, Paris), data compilation from various collections and a comprehensive review of existing literature. We reviewed 5,886 specimens and we extracted 279 additional data from literature sources. These data confirm the presence in Corsica of 91 species of which two are endemic, including Hoplitis corsaria (Warncke, 1991) which is elevated to species rank stat. nov. One new synonymy is established: Megachile lucidifrons Ferton, syn. nov. of Megachile albocristata Smith, 1853. The presence in Corsica of 19 species is regarded as dubious or erroneous. Finally, the types of Megachilid bees housed at MNHN and described based on Corsican material are illustrated. Lectotypes are designated for Megachile sicula var. corsica Benoist, 1935, Osmia corsica Ferton, 1901, Osmia erythrogastra Ferton, 1905, Osmia lanosa Pérez, 1879, and Osmia lineola Pérez, 1895.

Keywords

Anthophila, Apoidea, mediterranean islands

Introduction

Bees (Apoidea, Anthophila) are essential pollinators, given their pivotal role in pollinating a wide variety of plants and their high level of specialization and dependence on specific flora for both larval and adult nutrition (Willmer 2011). This diverse group encompasses approximately 20,000 species globally, classified into seven families: Andrenidae, Apidae, Colletidae, Halictidae, Mellitidae, Megachilidae, and Stenotritidae. Notably, bee biodiversity hotspots are concentrated in regions with Mediterranean and xeric climates, including the Mediterranean basin (Orr et al. 2021). Despite their ecological significance, recent efforts in compiling checklists have highlighted a significant knowledge gap regarding bee diversity in Mediterranean islands (e.g. Varnava et al. 2019; Nobile et al. 2021). Particularly, the fauna of the Corsican Island remains largely unexplored.

Spanning 8,722 km², Corsica ranks as the fourth-largest Mediterranean island. Its topography is dominantly shaped by expansive mountain ranges covering much of the land, with the highest peak reaching 2,710 m. The prevailing climate is largely Mediterranean, characterized by hot and arid summers, erratic rainfall patterns, and the potential for intense rain events in autumn and spring. However, at higher elevations, the climate takes on an alpine character, marked by lower temperatures (DREAL 2017). The varied elevation gradients contribute to a diverse range of habitats, encompassing coastal dune grasslands, elevated pine and beech forests, and various maquis or shrub thickets. This mosaic of habitats, coupled with Corsica’s prolonged isolation from the mainland, fosters a rich array of fauna and flora, including numerous endemic species (Sabiani 2004).

The main investigations into Corsican wild bees occurred mainly from the late XIX^th century to the early XX^th century. During this period, Charles Ferton (1856–1921), a military professional stationed at the Bastion of Bonifacio from 1895 until his passing in 1921, published a series of notes and descriptions introducing new species (Ferton 1923). Starting in 1895, C. Ferton devoted considerable time observing and collecting wasps and bees in the Bonifacio area, although he also collected in other parts of the island. His extensive collection, now housed at the Muséum national d’Histoire naturelle (MNHN) in Paris, represents the largest collection of Corsican material. Additionally, C. Ferton published approximately thirty studies and notes on the diversity and behaviour of bees and wasps. Following Ferton’s era, few surveys were conducted in Corsica. The literature on Corsican Anthophila is therefore particularly scarce, difficult to access and disjointed, often dealing only with a few isolated taxa. Only a handful of works have undertaken a comprehensive review of certain genera (Rasmont and Adamski 1995: Bumblebees; Terzo et al. 2007: Ceratina Latreille, 1802; Liongo Li Enkulu 1988: Megachile Latreille, 1802).

In 2016, a research initiative led by C. Villemant, C. Fontaine, and A. Perrard (MNHN) aimed to investigate mimicry in Aculeata communities in Bonifacio. This project marked the beginning of the restoration and examination of Corsican material from Ferton’s collection. From 2016 onwards, extensive and collaborative sampling efforts resulted in an additional substantial influx of new data for the island, prompting the need for an updated checklist of Corsican bees, given the absence of synthesis works. For our first contribution to a comprehensive review of Corsican bees, we focused on the Megachilidae. The compilation of 65 publications suggests the presence of up to 90 taxa in Corsica. Many of these records, however, are questionable or outdated, relying on unclear nomenclature or species concepts. Recognizing the significant disparity between modern and historical taxonomies, a thorough revision of these records was imperative. Based on a bibliographic synthesis and an extensive review of collection material we propose an updated checklist of Corsican Megachilidae.

Material and methods

Study of the collections housed at MNHN

The historical data presented in this article predominantly stem from the examination of Charles Ferton’s collection. This material was initially stored in numerous boxes organized by locality and date within a distinct section of the MNHN collection (Fig. 1). It underwent labelling, taxonomic identification to the lowest level (species or subspecies), and digitization of the label data. The Corsican segment of Ferton’s collection encompasses over 14,000 Aculeate specimens, including 2,038 Megachilidae specimens, constituting approximately one-third of his entire Aculeate collection. The remaining two-thirds primarily originates from Algeria and Mainland France. Ferton’s collection also includes five handwritten books containing field trip details and observations (Figs 2, 3). Only the book for the years 1911–1920 is missing. An important part of his collection is directly linked to these manuscripts. Ferton’s practice of annotating page numbers on original locality labels facilitated the connection between detailed observations in the manuscripts and the corresponding specimens in his collection. The final two manuscripts primarily focused on Corsica have been particularly helpful for encoding the data associated to the specimens by offering insights into collecting methods and localities. In addition to Ferton’s collection, 337 Megachilidae specimens from Corsica were identified in various parts of the MNHN collections, including those of R. Benoist (material collected between 1920 and 1940), E. G. Dehaut (material collected by G. Bénard between 1909 and 1910), and J. Sichel (material collected in the 1860s without specifying localities). Several other collections, including those of J. Barbier, L. de Berland, J. Casewitz-Weulersse, J. de Gaulle, J. Hamon, S. Kelner-Pillault, A. L. M. Lepeletier de Saint-Fargeau, and H. Nouvel, also contain a limited number of specimens from Corsica.

Figures 1–3.

Collection and catalogue of C. Ferton 1 example of box housing unidentified specimens collected and mounted by C. Ferton 2 manuscript of C. Ferton with morphological notes 3 manuscript detailing observations about the nesting of some Megachilid bees (in red, additional information added by C. Ferton subsequent to the initial black draft).

Recent sampling

The study of historical collections was completed by recent samplings conducted within three different programs by (1) A standardized sampling initiative aimed at exploring bee and wasp communities was undertaken in 2017. Monthly sampling occurred from March to November 2017 in seven localities of Bonifacio explored by C. Ferton, using active netting and bi-weekly pan-trapping following the Westphal protocol (Westphal et al. 2008). The collected material was deposited at MNHN and the collections of the Observatoire Conservatoire des Invertébrés de Corse (OCIC), branch of the Office de l’Environnement de la Corse (OEC). (2) Between 2019 and 2021, bees were collected for two to four weeks per year across the island in the context of the expedition “La Planète Revisitée (Our Planet Revisited)–Corsica 2019–2022,” organized by MNHN. Collection methods included hand nets, pan traps, and Malaise traps. The collected specimens were distributed among one of the authors (RLD), MNHN, and OCIC. A barcoding program initiated during LPR expeditions resulted in the creation of dataset DS-MEGA1, available on BOLD (www.barcodinglife.org), featuring 89 barcoded specimens representing 51 species. (3) Since 2019, OCIC has undertaken a territorial actions plan for pollinators, including the Corsican Honeybee (Cornuel-Willermoz and Andrei-Ruiz 2021). Over the past three years, this program conducted multiple surveys in 144 localities, encompassing 67 towns, primarily using active netting, supplemented by pan traps and malaise traps. The collected specimens were distributed among one of the authors (ACW), and OCIC.

Additional data gathering

Additional data was also obtained from various sources, including literature review, other collections, and an online forum. The review of the literature was based on 66 published works and provided 279 original data of Corsican Megachilidae, ranging from the short mention ‘Corsica’ to the complete data. We also obtained data from J. Mann’s collection in the Naturhistorisches Museum, Vienna, data from various museums and private collections thanks to A. Müller, E. Dufrêne, G. Le Goff and P. Vignac. Finally, we used the data from “Le Monde des Insectes” website (https://insecte.org/), restricting our research to records with pictures, to confirm the specimen identification.

Conventions of the checklist

We present the resulting data as an annotated checklist in which species are sorted by tribe and by genus. Under each species name are listed the literature records. These mentions include the original name used in the publication, the bibliographic reference, as well as the published localities. Additional comments are presented between square brackets. A question mark means that it was not possible to assign a record with confidence to a species. We used an asterisk (*) to highlight taxa for which Corsican data are reported for the first time in this study.

The original data we compiled for each species comprise the number of examined specimens (sorted by sex, ‘NS’ is indicated when sex is unknown), the collecting period and the list of municipalities. Detailed records were uploaded online in ten datasets on Cardobs (https://cardobs.mnhn.fr/) (IDs: 884FEA17-4810-3B88-E053-5014A8C0FB66, 87FCFAFA-BC57-0572-E053-5014A8C04A0C, 90768076-0CF5-15F2-E053-5014A8C08F4A, A766A82F-BE8D-556D-E053-2614A8C0C12D, BF62B36D-1D84-2F86-E053-3014A8C0EFF8, E4C9E0B0-B16B-04D9-E053-3014A8C043D8, EB611548-937C-63F5-E053-0514A8C02064, EB6380B5-805D-5935-E053-0514A8C08744, EFF30F3C-ACD5-6B6E-E053-0514A8C0121F, EFA34354-3736-7BE7-E053-0514A8C00AD2). They can be consulted and downloaded on the Openobs data portal (https://openobs.mnhn.fr/) and MNHN data portal (https://science.mnhn.fr/). Special attention is given to type material (last section) for which complete label data are cited. Different labels are separated by a double slash. Additional information on labels is given between square brackets. Sex of the types is provided before the label information.

The specimens examined in this study are summarized in Table 1. To assess the species richness of the island, we compared these data to the checklists of the three largest mediterranean islands in Table 2: Sardinia, Sicily and Cyprus (Comba 2019; Varnava et al. 2020; Nobile et al. 2021; personal communication with Maurizio Cornalba). In order to estimate the completeness of our sampling effort of the Corsican Megachilid fauna, we used the Chao1 and abundance-based coverage estimators (ACE) (Chao 1984; Chao and Lee 1992; Colwell and Coddington 1994).

Table 1.

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Numbers of examined specimens by species in decreasing order.

Tribe	Species	N	Tribe	Species	N
Megachilini	Megachile argentata	535	Osmiini	Hoplitis praestans	33
Osmiini	Heriades crenulata	512	Anthidiini	Anthidium cingulatum	29
Osmiini	Osmia caerulescens	331	Megachilini	Coelioxys inermis	28
Osmiini	Hoplitis corsaria	265	Osmiini	Hoplitis fasciculata	24
Megachilini	Megachile sicula	257	Anthidiini	Icteranthidium laterale	23
Anthidiini	Anthidiellum strigatum	251	Anthidiini	Anthidium florentinum	21
Osmiini	Heriades rubicola	203	Osmiini	Chelostoma rapunculi	21
Osmiini	Hoplitis anthocopoides	186	Osmiini	Osmia emarginata	19
Osmiini	Hoplitis bisulca	167	Megachilini	Megachile ericetorum	18
Osmiini	Osmia niveata	167	Osmiini	Osmia erythrogastra	18
Osmiini	Osmia rufohirta	163	Anthidiini	Pseudoanthidium melanurum	16
Osmiini	Hoplitis bihamata	152	Megachilini	Megachile rotundata	14
Anthidiini	Anthidium taeniatum	142	Osmiini	Hoplitis manicata	14
Anthidiini	Rhodanthidium septemdentatum	117	Lithurgini	Lithurgus chrysurus	13
Osmiini	Osmia ligurica	116	Megachilini	Coelioxys brevis	13
Megachilini	Megachile melanopyga	112	Megachilini	Megachile albocristata	13
Osmiini	Osmia bicornis bicornis	107	Anthidiini	Stelis nasuta	12
Osmiini	Osmia ferruginea	100	Megachilini	Coelioxys conoideus	12
Megachilini	Coelioxys afer	98	Megachilini	Coelioxys mandibularis	12
Megachilini	Megachile centuncularis	89	Osmiini	Osmia nasoproducta	12
Osmiini	Hoplitis adunca	89	Osmiini	Hoplitis tridentata	11
Osmiini	Osmia versicolor	87	Anthidiini	Anthidium oblongatum	9
Osmiini	Hoplitis cristatula	82	Dioxyini	Dioxys cinctus	9
Osmiini	Osmia latreillei	82	Megachilini	Coelioxys acanthura	8
Megachilini	Megachile albisecta	78	Anthidiini	Stelis signata signata	7
Osmiini	Osmia submicans	67	Megachilini	Megachile deceptoria	7
Osmiini	Hoplitis aff. adunca	61	Anthidiini	Stelis murina	6
Megachilini	Megachile leachella	59	Megachilini	Coelioxys haemorrhoa	6
Osmiini	Hoplitis leucomelana	58	Osmiini	Chelostoma foveolatum	6
Osmiini	Chelostoma distinctum	57	Osmiini	Osmia cornuta cornuta	6
Megachilini	Megachile apicalis	54	Anthidiini	Stelis ornatula ornatula	4
Osmiini	Osmia signata signata	54	Dioxyini	Aglaoapis tridentata	4
Anthidiini	Anthidium manicatum	52	Anthidiini	Stelis punctulatissima	3
Osmiini	Heriades truncorum	50	Megachilini	Megachile burdigalensis	3
Osmiini	Hoplitis perezi	50	Megachilini	Megachile versicolor	3
Megachilini	Megachile pusilla	49	Anthidiini	Stelis minuta	2
Anthidiini	Stelis breviuscula	47	Lithurgini	Lithurgus cornutus	2
Anthidiini	Pseudoanthidium stigmaticorne	43	Megachilini	Coelioxys obtusus	2
Osmiini	Hoplitis acuticornis	43	Osmiini	Hoplitis ravouxi	2
Osmiini	Osmia aurulenta	38	Osmiini	Osmia melanogaster	2
Osmiini	Osmia scutellaris	38	Anthidiini	Icteranthidium grohmanni	1
Osmiini	Osmia tricornis	37	Megachilini	Coelioxys aurolimbatus	1
Anthidiini	Pseudoanthidium nanum	36	Osmiini	Osmia anceyi	1
Megachilini	Megachile lagopoda	34	Osmiini	Protosmia minutula	1

Table 2.

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Megachilidae diversity of the four largest Mediterranean islands.

Island	Corsica	Sardinia	Sicily	Cyprus
Size (km²)	8722	24090	25711	9251
Genera	17	15	17	15
Species	91	91	129	91
Endemic/subendemic species	2(3*)	2(3*)	3	3
Aglaoapis	1	1	0	0
Anthidiellum	1	1	1	2
Anthidium	5	3	8	5
Chelostoma	3	1	5	3
Coelioxys	10	8	14	12
Dioxys	1	2	3	2
Eoanthidium	0	0	0	1
Heriades	3	3	4	4
Hoplitis	15	15	19	9
Icteranthidium	2	0	2	2
Lithurgus	2	3	3	2
Megachile	15	18	20	17
Osmia	19	25	32	25
Protosmia	1	0	3	3
Pseudoanthidium	3	3	3	0
Rhodanthidium	2	2	3	1
Stelis	7	4	7	3
Stenoheriades	0	0	1	0
Trachusa	1	2	1	0

Results and discussion

Our synthesis is based on the examination of 5886 specimens (Table 1) and 279 data from the literature. It confirms the presence in Corsica of 91 species of Megachilidae, belonging to 17 different genera. Among them, twenty species and two genera are newly recorded in Corsica. Modern records are provided for 29 species which were only known from historical records. We could not get modern data for five species and further research is needed to clarify their presence in the island. Finally, the presence in Corsica of 19 species is regarded as dubious or erroneous.

It is noteworthy to highlight that Megachile sculpturalis Smith, 1853, an invasive Megachilid bee, has not been recorded in Corsica so far. This species arrived in France in 2008 (Vereecken and Barbier 2009) and is now widespread in a large part of Europe. It was recently found in Elba Island (Italy), only 50 km from Corsica (Ruzzier et al. 2020).

Based on our sample data, we found a Chao1 of 92.6 and an ACE of 92.955. Both estimators suggest that we detected most of the Megachilid species in the explored parts of the island. Additional efforts in the explored part may therefore result in few additional species for the checklist. However, the distribution of the data is uneven (Fig. 4) as Megachilidae data were available for only 131 of the 360 Corsican municipalities. Well-sampled sectors include Bonifacio, due to the Ferton data and the 2017 study, Vivario, and the coastline. These three areas concentrate a large proportion of the data. Under-sampled areas include the northwest (Cap Corse and Castagniccia region) and a large part of the southwest (Liamone and Taravo regions) probably due to the difficulty to access these rather remote regions. It is therefore possible than a few other species may be found in these areas. As such, the Territorial Actions Plan for wild pollinators defines these sectors as priority targets for more extensive inventories in the coming years.

Figure 4.

Distribution of the data in the different Corsican municipalities. Black dots are collecting stations with geographical coordinates. Municipalities for which we have imprecise collecting localities are highlighted in green (based on the quantity of data available).

In comparison to Megachilidae checklists from larger Mediterranean islands, Corsica exhibits notably high species richness (Table 2). Currently, only Sicily surpasses Corsica in Megachilidae diversity. This discrepancy can be attributed to the considerably larger size of Sicily and its proximity to the mainland, facilitating the establishment of new species from the continent. Conversely, despite being almost 2.76 times larger than Corsica, Sardinia displays a similar species richness. Since Corsica and Sardinia are very close islands, it is not surprising to find a similar diversity. However, their fauna is not entirely overlapping, possibly due to their different geological histories and the different habitats found in the two islands. Notably, mountainous habitats such as alpine grasslands are present in Corsica but not in Sardinia. Additionally, most bees sampling effort in Sardinia seem to have been restricted to the north-western part of the island, with only a few sites which have been recently sampled in the rest of the island (Floris et al. 2000; Quaranta et al. 2004; Nobile et al. 2021). It is therefore likely that this similarity in diversity between Sardinia and Corsica roots from a lack of knowledge on the Megachilidae fauna of the larger island.

Below, we list the species present in Corsica, along with the dubious and erroneous records and the Corsican type material housed at MNHN. This checklist confirmed the expected high diversity of Megachilidae in Corsica and call for similar efforts to address the local diversity of the other bee families.

Checklist of the Corsican Megachilidae

Anthidiini Ashmead, 1899

Anthidiellum Cockerell, 1904

1. Anthidiellum strigatum strigatum (Panzer, 1805)

Anthidium strigatum Panzer; Ferton (1901b: 92, 146): Bonifacio.

Anthidiellum strigatum (Panzer); Pagliano (1994: 392): Sotta.

Anthidiellum strigatum (Panzer); Kasparek (2022: 79): Corsica.

Material examined. 78♀, 165♂ & 8NS observed from April to October, between 1895 and 2021 in Albertacce, Aléria, Barbaggio, Asco, Bastelica, Bastia, Biguglia, Bonifacio, Calvi, Canale–di–Verde, Castellare–di–Casinca, Corbara, Corte, Coti–Chiavari, Evisa, Farinole, Figari, Furiani, Ghisonaccia, Ghisoni, Grosseto–Prugna, Lecci, Levie, Linguizzetta, Lucciana, Manso, Mausoléo, Ogliastro, Oletta, Olmi–Cappella, Palasca, Patrimonio, Penta–di–Casinca, Poggio–d’Oletta, Porto–Vecchio, Propriano, Quenza, Rospigliani, Saint–Florent, Santo–Pietro–di–Tenda, Sartène, Sorbollano, Tavera, Valle–di–Campoloro, Ventiseri, Vescovato, Vico, Vivario, Zicavo and Zonza.

Anthidium Fabricius, 1804

2. Anthidium cingulatum Latreille, 1809

Anthidium cingulatum Latreille; Nadig and Nadig (1934: 27): Monte d’Oro.

Anthidium cingulatum Latreille; Warncke (1981: 335, Map 29): Corsica.

Anthidium cingulatum Latreille; Kasparek (2022: 93): Corsica.

Material examined. 13♀ & 16♂ observed from May to October between 1900 and 2022 in Balogna, Bocognano, Bonifacio, Corte, Ghisoni, La Porta, Lumio, Mausoléo, Nocario, Palasca, Quenza, Sisco, Talasani, Vivario and Zonza.

3. Anthidium florentinum (Fabricius, 1775)*

Material examined. 15♀ & 6♂ observed from June to September between 2002 and 2022 in Ajaccio, Albertacce, Calacuccia, Castellare–di–Casinca, Lucciana, Propriano, Quenza, San-Giuliano, Talasani and Vico.

4. Anthidium manicatum manicatum (Linnaeus, 1758)

Anthidium manicatum Linnaeus; Ferton (1909a: 551–552): Bonifacio.

Anthidium manicatum (Linnaeus); Pagliano (1994: 396): Sotta.

Anthidium manicatum (Linnaeus); Kasparek (2022: 106): Corsica.

Anthidium manicatum (Linnaeus); Meunier et al. (2023): Corsica.

Material examined. 29♀ & 23♂ observed from May to October, between 1897 and 2021 in Aléria, Barbaggio, Bastelica, Bonifacio, Calacuccia, Casamaccioli, Castellare–di–Casinca, Corte, Ghisoni, Grosseto–Prugna, Lumio, Mausoléo, Meria, Palasca, Quenza, Santa–Maria–Poggio, Riventosa, Sisco, Tavera, Vivario and Zonza.

5. Anthidium oblongatum oblongatum (Illiger, 1806)*

Material examined. 1♀ & 8♂ observed from June to July, between 2019 and 2021 in Bocognano, Casamaccioli, Quenza, Solaro and Zonza.

6. Anthidium taeniatum Latreille, 1809*

Material examined. 45♀ & 97♂ observed from May to September, between 1895 and 2021 in Ajaccio, Bonifacio, Linguizzetta, Lucciana, Mausoléo, Patrimonio, Penta–di–Casinca, Prunelli–di–Fiumorbo, Sermano, Soveria, Talasani and Zonza.

Icteranthidium Michener, 1948

7. Icteranthidium grohmanni (Spinola, 1838)

Icteranthidium grohmanni (Spinola); Pagliano (1994: 395): Col de Celaccia.

Icteranthidium grohmanni (Spinola); Kasparek (2022: 163): Corsica.

Material examined. 1♂ observed in September 2020 in Tavera.

8. Icteranthidium laterale laterale (Latreille, 1809)

Anthidium laterale Latreille; Stöckl (2000: 276): Corsica.

Icteranthidium laterale (Latreille); Kasparek (2022: 165): Corsica.

Material examined. 13♀ & 10♂ observed from July to September, between 1896 and 2022 in Balogna Bonifacio, Ghisoni, Rospigliani, Santo–Pietro–di–Venaco, Tavera, Vico and Vivario.

Pseudoanthidium Friese, 1898

9. Pseudoanthidium melanurum (Klug, 1832)

Pseudoanthidium melanurum (Klug); Kasparek (2022: 188): Corsica.

Material examined. 4♀ & 12♂ observed from May to June, between 2017 and 2020 in Borgo, Castellare–di–Casinca, Penta–di–Casinca and Venzolasca.

10. Pseudoanthidium nanum (Mocsáry, 1880)

? Pseudoanthidium lituratum (Panzer); Pagliano (1994: 398): Bastia.

Pseudoanthidium nanum (Mocsáry); Litman et al. (2021): Ajaccio, Bonifacio, Lecci, Sartène, Zicavo, Albertacce, Corte, Ghisoni, Lozzi, Palasca, Vivario.

Pseudoanthidium nanum (Mocsáry); Kasparek (2022: 191): Corsica.

Material examined. 21♀ & 15♂ observed from May to October, between 1901 and 2022 in Albertacce, Balogna, Bonifacio, Calacuccia, Corte, Coti–Chiavari, Ghisoni, Grosseto–Prugna, Mausoléo, Montegrosso, Palasca, Renno, Sartène, Ventiseri, Vivario and Zicavo.

Remark. Pseudoanthidium nanum has long been a challenging species complex before the revision of Litman et al. (2021). Two species of this complex occur in Corsica. It is therefore not possible to assign the record of Pagliano (1994) to one of these two species with confidence. Our data, partially published in Litman et al. (2021), confirmed the presence of this taxon in Corsica.

11. Pseudoanthidium stigmaticorne (Dours, 1873)

Anthidium lituratum Latr.; Pérez (1879: 213) [Misidentification]: Corsica.

Anthidium lituratum Latr.; Ferton (1901b: 88): Corsica.

Anthidium lituratum Latr. (= peregrinum Costa); Ferton (1909a: 552–553, 575, 578) [Misidentification]: Pianottoli–Caldarello.

Pseudoanthidium scapulare (Latreille); Scheuchl and Willner (2016: 791) [Misinterpretation of the record of Ferton (1909a)]: Corsica.

Pseudoanthidium stigmaticorne (Dours); Litman et al. (2021): Ajaccio, Bonifacio, Pianottoli–Caldarello, Propriano.

Pseudoanthidium stigmaticorne (Mocsáry); Kasparek (2022: 198): Corsica.

Material examined. 19♀ & 24♂ observed from May to September, between 1897 and 2019 in Ajaccio, Bonifacio, Pianottoli-Caldarello and Propriano.

Rhodanthidium Insensee, 1927

12. Rhodanthidium septemdentatum (Latreille, 1809)

Anthidium septemdentatum Latreille; Ferton (1909a: 551): Bonifacio.

Anthidium septemdentatum Latreille; Nadig and Nadig (1934: 26): Cap Corse.

Anthidium septemdentatum Latreille; Kusdas (1974: 160): Calvi.

Rhodanthidium septemdentatum (Latreille); Kasparek (2022: 223): Corsica.

Material examined. 51♀, 64♂ & 2NS observed from April to July, between 1895 and 2021 in Ajaccio, Bonifacio, Calacuccia, Castellare–di–Casinca, Chisa, Corte, Farinole, Galéria, Grosseto–Prugna, L’Île Rousse, Mausoléo, Palasca, Patrimonio, Pianottoli-Caldarello, Porri, Porto–Vecchio, Santa–Maria–Poggio, Riventosa, Santo–Pietro–di–Tenda, Sotta, Ventiseri, Venzolasca, and Zonza.

13. Rhodanthidium sticticum (Fabricius, 1787)

Rhodanthidium sticticum (Fabricius); Kasparek and Lhomme (2019: 44): Porto–Vecchio.

Rhodanthidium sticticum (Fabricius); Kasparek (2022: 228): Corsica.

Remark. There is only one record from 1976 (Kasparek and Lhomme 2019). No modern record is known.

Stelis Panzer, 1806

14. Stelis breviuscula Nylander, 1841*

Material examined. 19♀ & 28♂ observed from May to October, between 1895 and 2021 in Ajaccio, Asco, Bonifacio, Casaglione, Farinole, Figari, Galeria, Grosseto–Prugna, Ogliastro, Oletta, Olmi–Cappella, Palasca, Patrimonio, Porto–Vecchio, Propriano, Riventosa, Saint–Florent, Santo–Pietro–di–Tenda, Sisco, Sorbollano, Urtaca and Ventiseri.

15. Stelis minuta Lepeletier, 1825*

Material examined. 1♀ & 1♂ collected in 1909 in Ghisonaccia and Porto–Vecchio.

Remark. No modern record.

16. Stelis murina Pérez, 1884

Figs 5–8

Stelis phaeoptera murina Pérez; Warncke (1992b: 356): Corsica.

Stelis phaeoptera (Kirby); Scheuchl and Willner (2016: 835): [quoting Warncke (1992b)].

Material examined. 2♀ & 4♂ observed from June to July, between 2019 and 2021 in Ghisoni, Mausoléo and Sisco.

Remark. Baker (1999) noted that the type material housed at MNHN does not include European specimens but only North African material in contradiction to the original description which also specify material from France and Spain. The unpublished catalogue of Pérez mentions specimens from Bizerte (Tunisia) and La Chiffa (Algeria). Warncke (1992b) designated one of the females from Bizerte as lectotype. Blüthgen (1930) already noted that the original description of Pérez did not match well with a syntype he examined. As a matter of fact, the modern species concept of S. murina (Warncke 1992b; Baker 1999; Aguib et al. 2014; Kasparek 2015) does not match with its lectotype and paralectotypes (coll. Pérez, MNHN). The specimens from Corsica are similar to the types of S. murina (including lectotype). Both females and males of S. murina are hardly distinguishable from S. phaeoptera (Kirby, 1802). They can be distinguished from it by their entirely pure white pilosity (Figs 5, 6) [dirty white to brown in S. phaeoptera, notably darker on the vertex and mesoscutum], except on the inferior side of the tarsi where it is dark brown (Fig. 6) [yellow gold in S. phaeoptera, see fig. 7 in Aguib et al. 2014] and by their dark brown to black tibial spurs (Fig. 6) [ochreous in S. phaeoptera, see fig. 7 in Aguib et al. 2014]. Finally, the barcode generated for the Corsican S. murina matches that of Moroccan specimens and is highly divergent from the European sequences of S. phaeoptera available in BOLD (Le Divelec and Wood unpublished). A detailed review of this species group is required. Considering that S. murina sensu modern authors occurs in Crete (Le Divelec unpublished) and that it matches the original description of S. murina cretica Mavromoustakis, 1963, it should most probably be referred to as S. cretica. Unfortunately, the types of S. murina cretica could not be located in Mavromoustakis collection and are seemingly lost (Santerre pers. comm.). Non-type specimens from Crete identified by Mavromoustakis as S. murina cretica are preserved in his collection and are morphologically similar to S. murina sensu modern authors. The female of S. cretica can be easily distinguished from that of S. murina by the widely notched apical margin of the clypeus (Fig. 9) [with straight indentation in S. murina (Fig. 7)] and by the conspicuous smooth medio-apical ridge on the last tergum (Fig. 10) [uniformly punctate in S. murina (Fig. 8)]. The male of S. cretica can be recognized by a notch bordered by a pair of teeth in the middle of the fourth sternum posterior margin (see fig. 4 in Aguib et al. 2014) [here with a conspicuous rake in S. murina, as in fig. 8 in Aguib et al. 2014].

Figures 5–10.

Comparative illustrations of Stelis murina Pérez and Stelis cretica Mavromoustakis (= Stelis murina sensu modern authors) 5–8 Stelis murina (from Corsica) 9, 10 Stelis cretica (from Crete) 5 pilosity of vertex and mesonotum 6 hind leg 7, 9 face 8, 10 tergum 6. Scale bars: 0.5 mm.

BOLD process ID. LPRCW173-19 (BIN, BOLD:AEC2025).

17. Stelis nasuta (Latreille, 1809)

Material examined. 6♀ & 6♂ observed from May to June, between 1902 and 2020 in Asco, Bonifacio and Mausoléo.

Remark. The whitish light drawings of Corsican individuals are remarkably reduced if not absent.

18. Stelis ornatula ornatula (Klug, 1807)*

Material examined. 2♀ & 2♂ observed from June to July, between 1909 and 2020 in Palasca, Patrimonio, Sorbollano and Vico.

19. Stelis punctulatissima punctulatissima (Kirby, 1802)*

Material examined. 2♀ & 1♂ collected in June 2020 in Lucciana.

20. Stelis signata signata (Latreille 1809)*

Material examined. 3♀ & 4♂ observed from May to September, between 1974 and 2021 in Galeria, Ghisonaccia, Ghisoni, Manso, Oletta, Rospigliani and Ventiseri.

Trachusa Panzer, 1804

21. Trachusa byssina (Panzer, 1798)

Trachusa byssina (Panzer); Kasparek (2022: 238): Corsica.

Remark. There is only one Corsican record (Kasparek 2022).

Dioxyini Cockerell, 1902

Aglaoapis Cameron, 1901*

21. Aglaoapis tridentata (Nylander, 1848)*

Material examined. 4♀ observed from June to September, between 2020 and 2021 in Ghisoni.

Remark. The morphology of the Corsican specimens is slightly different than that of the West palearctic specimens. The Corsican population might represent a distinct species. It was only observed in the Mountains where it seems to be very rare.

Dioxys Lepeletier & Serville, 1825

22. Dioxys cinctus (Jurine, 1807)

Dioxys cincta Jurine; Friese (1895: 109): Corsica.

Dioxys cinctus (Jurine); Bogusch (2023, Fig. 8): Corsica.

Material examined. 4♀ & 5♂ observed from May to June, between 1902 and 2021 in Bonifacio, Ghisonaccia, Oletta, Porto–Vecchio and Ventiseri.

Lithurgini Newman, 1834

Lithurgus Berthold, 1827

23. Lithurgus chrysurus Fonscolombe, 1834

Lithurge chrysura Fonscolombe; Canovai et al. (2000: 78): Corsica.

Material examined. 9♀ & 4♂ observed from June to August, between 2003 and 2022 in Balogna, Canale–di–Verde, Coti–Chiavari, Mausoléo, Noceta, Rutali and Vico.

24. Lithurgus cornutus fuscipennis Lepeletier, 1841

Lithurgus cornutus (Fabricius); Dufrêne et al. (2016: 18): Corsica.

Material examined. 2♀ observed from August to September, between 2020 and 2021 in Asco and Tavera.

Megachilini Latreille, 1802

Coelioxys Latreille, 1809

25. Coelioxys acanthura (Illiger, 1806)

Coelioxys acanthura (Illiger); Ferton (1901b: 92): Bonifacio.

Coelioxys acanthura (Illiger); Warncke (1992a: 57): Corsica.

Coelioxys acanthura (Illiger); Pagliano (1994: 375): Algajola.

Material examined. 4♀ & 4♂ observed from June to August, between 1898 and 2021 in Bonifacio, Calvi, Coti–Chiavari, Palasca, Prunelli–di–Fiumorbo and Rutali.

26. Coelioxys afer Lepeletier, 1841

Coelioxys afra Lepeletier; Pagliano (1994: 375): Bastia.

Material examined. 53♀ & 45♂ observed from April to November, between 1895 and 2021 in Bonifacio, Evisa, Figari, Galeria, Grosseto–Prugna, Oletta, Palasca, Porto–Vecchio, Quenza, Saint–Florent, Santo–Pietro–di–Tenda, Sermano, Soveria, Talasani, Vero, Vescovato, Vivario, Zicavo and Zonza.

27. Coelioxys aurolimbatus aurolimbatus Förster, 1853*

Material examined. 1♂ collected in July 2019 in Mausoléo.

Remark. This species is known from only one mountainous station.

28. Coelioxys brevis Eversmann, 1852

Coelioxys brevis Eversmann; Warncke (1992a: 53): Serra–di–Ferro, Linguizzetta.

Material examined. 8♀ & 5♂ observed from May to July, between 1972 and 2021 in Aléria, Ghisonaccia, Linguizzetta, Palasca, Solaro, Tallone and Ventiseri.

29. Coelioxys conoideus (Illiger, 1806)*

Material examined. 3♀ & 9♂ observed from July to September, between 1897 and 2019 in Cozzano, Mausoléo and Vivario.

Remark. This species has only been collected in mountains.

30. Coelioxys echinatus Förster, 1853

Coelioxys rufocaudatus Smith; Nadig and Nadig (1934: 62): Cap Corse.

Coelioxys echinata Förster; Warncke (1992a: 62): Corsica.

Remark. We could not confirm the occurrence of C. echinatus in Corsica. This species has been recorded twice in the past. It is unlikely to be confused with any other species, but we cannot exclude a confusion with C. brevis. We believe these records to be accurate for now.

31. Coelioxys haemorrhoa haemorrhoa Förster, 1853

Coelioxys haemorrhoa Förster; Le Divelec and Dufrêne (2020: 6): Bonifacio.

Material examined. 3♀ & 3♂ observed from the end of May to September, between 1895 and 2017 in Bonifacio and Venzolasca.

32. Coelioxys inermis (Kirby, 1802)

Coelioxys acuminata Nylander; Ferton (1909a: 551): Bonifacio.

Coelioxys inermis (Kirby); Warncke (1992a: 65): Corsica.

Material examined. 10♀ & 18♂ observed from May to November, between 1896 and 2022 in Ajaccio, Asco, Balogna, Bonifacio, Corte, Evisa, Poggio–di–Venaco, Rospigliani, Vivario and Zonza.

33. Coelioxys mandibularis Nylander, 1848*

Material examined. 7♀ & 5♂ observed from May to October, between 2009 and 2021 in Bonifacio, Ghisonaccia, Levie, Mausoléo, Moncale, Olmi–Cappella, Palasca and Quenza.

34. Coelioxys obtusus Pérez, 1884*

Material examined. 1♀ & 1♂ observed in July 2019 in Palasca.

Megachile Latreille, 1802

35. Megachile albisecta (Klug, 1817)

Megachile sericans Fonscolombe; Ferton (1901b: 90–92, 145): Bonifacio.

Megachile albisecta Klug in Germar; Benoist (1940: 47): Corsica.

Creightonella albisecta (Klug); Liongo Li Enkulu (1988, Map 19): Corsica.

Creightonella albisecta (Klug); Pagliano (1994: 370): Sotta.

Material examined. 34♀ & 44♂ observed from June to September, between 1896 and 2022 in Balogna, Bonifacio, Coti–Chiavari, Ghisoni, Manso, Omessa, Palasca, Patrimonio, Piana, Propriano, Rospigliani, Rutali, Serra–di–Ferro and Vico.

36. Megachile albocristata Smith, 1853

Figs 39–42

Megachile (Chalicodoma) lucidifrons sp. nov.; Ferton (1905: 57–58): Bonifacio.

Megachile (Chalicodoma) lucidifrons Ferton; Ferton (1909b: 407): Bonifacio.

Chalicodoma albocristata Smith; Liongo Li Enkulu (1988, Map 20): Cap Corse.

Chalicodoma lucidifrons Ferton; Liongo Li Enkulu (1988, Map 44) [quoting Ferton (1905)].

Material examined. 9♀ & 4♂ observed from June to July, between 2019 and 2022 in Asco, Mausoléo, Santo–Pietro–di–Venaco, Sermano and Sisco.

Ecology. The ecology of Megachile albocristata is poorly known. The specimens were all observed in mountainous regions, specifically on sun-exposed rocky slopes covered with scrub vegetation (Fig. 11). These habitats featured prominent fractured rock formations and scree slopes where numerous individuals were observed in flight. During our investigations, we chanced upon a concealed nest beneath a boulder, situated within a cavity (Fig. 12). The layer (Fig. 13) covering the cells of the nest consist of a composite material comprising vegetable paste and ground gravels that the female takes great care to calibrate. The nesting behavior of M. albocristata appears to closely resemble that of M. lefebvrei, as detailed by Ferton (1909a: 544–547). It has been observed by Ferton on Teucrium maritimum (Lamiaceae). We observed this species visiting flowers of Teucrium in Mausoléo.

Figures 11–13.

Nesting site of Megachile albocristata 11 adret covered with sparse Oak wood (Quercus ilex) in the Tartagine valley (Mausoléo) 12 boulder sheltering a nest of Megachile albocristata 13 nest of Megachile albocristata.

Remark. In Corsica, M. albocristata was previously known from only one uncertain historical record (Liongo Li Enkulu 1988). Our observations thus confirm the presence of this taxon in Corsica.

37. Megachile apicalis Spinola, 1808

Megachile apicalis Spinola; Ferton (1909a: 550): Bonifacio.

Megachile apicalis Spinola; Nadig and Nadig (1934: 26): Cap Corse.

Megachile apicalis Spinola; Liongo Li Enkulu (1988, Map 23): Corsica.

Material examined. 39♀ & 15♂ observed from from June to September, between 1895 and 2022 in Balogna, Bonifacio, Canale–di–Verde, Corte, Lucciana, Mausoléo, Oletta, Palasca, Patrimonio, Propriano, Rutali, Sartène and Vico.

48. Megachile argentata schmiedeknechti Costa, 1884

Figs 47–49

Megachile xanthopyga sp. nov.; Pérez (1895: 25).

Megachile xanthopyga Pérez; Ferton (1897: 48): Bonifacio.

Megachile xanthopyga Pérez; Ferton (1909a: 550): Bonifacio.

Megachile schmiedeknechti Costa; Nadig and Nadig (1934: 26): Cap Corse.

Megachile schmiedeknechti Costa; Benoist (1940: 65): Bonifacio, Vivario.

Megachile schmiedeknechti Costa; Rebmann (1968: 30): Corsica.

Megachile schmiedeknechti Costa; Kusdas (1974: 160): Calvi.

Megachile pilidens Alfken; Liongo Li Enkulu (1988, Map 54): Corsica.

Megachile schmiedeknechti Costa; Liongo Li Enkulu (1988, Map 57): Corsica.

Megachile pilidens Alfken; Pagliano (1994: 373): Venzolasca.

Megachile schmiedeknechti Costa; Pagliano (1994: 374): Golfe de Liscia.

Megachile argentata schmiedeknechti Costa; Praz and Bénon (2023: 170): Corsica.

Material examined. 299♀, 234♂ & 2NS observed from April to October, between 1895 and 2022 in Ajaccio, Albertacce, Aléria, Asco, Balogna, Bastelica, Biguglia, Bonifacio, Canale–di–Verde, Castellare–di–Casinca, Centuri, Conca, Coti–Chiavari, Evisa, Farinole, Figari, Ghisonaccia, Ghisoni, Grosseto–Prugna, La Porta, Lecci, Levie, Linguizzetta, Lucciana, Mausoléo, Nocario, Noceta, Oletta, Palasca, Patrimonio, Penta–di–Casinca, Piana, Poggio–d’Oletta, Porto–Vecchio, Propriano, Quenza, Riventosa, Saint–Florent, Santo–Pietro–di–Tenda, Sartène, Sermano, Serra–di–Ferro, Sisco, Solaro, Sorbollano, Soveria, Talasani, Tavera, Ventiseri, Venzolasca, Vescovato, Vico, Vivario, Zicavo and Zonza.

Remark. This taxon has been recently relegated to the status of a subspecies of Megachile argentata (Fabricius, 1793) (Praz and Bénon 2023). This subspecies occurs in West Mediterranean islands such as Corsica, Sardinia, and Malta (Praz and Bénon 2023). Notably, both M. a. argentata (formerly identified as M. pilidens, now recognized as a synonym of M. argentata) and M. a. schmiedeknechti have been documented in Corsica (Liongo Li Enkulu 1988; Pagliano 1994). However, the distinction of these two taxa primarily relies on the colour of their pilosity which might pose challenge in discerning between old, discoloured specimens, particularly in males. Upon scrutinizing numerous specimens, we believe that the Corsican records of M. pilidens (= M. a. argentata) refer to M. a. schmiedeknechti.

38. Megachile burdigalensis Benoist, 1940*

Material examined. 1♀ & 2♂ observed from June to September, in 2020 in Bastelica, Oletta and Patrimonio.

39. Megachile centuncularis centuncularis (Linnaeus, 1758)

Megachile centuncularis Linnaeus; Ferton (1909a: 551): Bonifacio.

Megachile centuncularis (Linnaeus); Liongo Li Enkulu (1988, Map 27): Corsica.

Material examined. 58♀ & 31♂ observed from April to November, between 1895 and 2022 in Ajaccio, Balogna, Bonifacio, Corscia, Evisa, Ghisoni, Grosseto–Prugna, La Porta, Mausoléo, Oletta, Orto, Ota, Palasca, Porto–Vecchio, Riventosa, Rospigliani, San–Martino–di–Lota, Santo–Pietro–di–Venaco, Serra–di–Fiumorbo, Serra–di–Scopamène, Sisco, Sorbollano, Vivario and Zonza.

40. Megachile deceptoria Pérez, 1890

Megachile deceptoria Pérez; Liongo Li Enkulu (1988, Map 30): Cap Corse.

Material examined. 4♀ & 3♂ observed from May to July, between 2020 and 2021 in Ghisonaccia, Saint–Florent, Santo–Pietro–di–Tenda and Serra–di–Fiumorbo.

Remark. The species was only known from one historical data (Liongo li Enkulu 1988). Our observations thus confirm the presence of this taxon in Corsica.

41. Megachile ericetorum Lepeletier, 1841

Chalicodoma ericetorum (Lepeletier); Liongo Li Enkulu (1988, Map 32): Corsica.

Chalicodoma ericetorum melaleuca Zanden ssp. nov.; Zanden (1989: 72): Algajola.

Chalicodoma ericetorum (Lepeletier); Canovai et al. (2000: 78): Corsica.

Megachile ericetorum melaleuca Zanden; Le Goff (2004: 10): Vivario.

Material examined. 8♀ & 10♂ observed from May to September, between 1897 and 2021 in Ajaccio, Albertacce, Barrettali, Corte, Mausoléo, Riventosa, Sermano and Vivario.

Remark. The Corsican populations were assigned to the subspecies M. ericetorum melaleuca (Zanden, 1989) which is characterized by a pure white pilosity (dirty white to yellowish in the nominative subspecies). This subspecies has been synonymised with M. ericetorum by Praz and Dorchin (2018).

42. Megachile lagopoda lagopoda (Linnaeus, 1760)

Megachile lagopoda (Linnaeus); Liongo Li Enkulu (1988, Map 38): Corsica.

Megachile lagopoda (Linnaeus); Pagliano (1994: 372): Col de Celaccia.

Material examined. 13♀ & 21♂ observed from July to September, between 1896 and 2021 in Corscia, Ghisoni, Mausoléo, Piana, Quenza, Renno, Tavera, Vivario and Zicavo.

43. Megachile leachella Curtis, 1828

Figs 35–37

Megachile argentata var. fossoria Ferton, var. nov.; Ferton (1909a: 550): Bonifacio, Propriano.

Megachile dorsalis Pérez; Alfken (1923: 8): Corsica.

Megachile argentata Fabricius; Nadig and Nadig (1934: 26): Golfe de Sagone.

Megachile argentata var. fossoria Ferton; Benoist (1940: 66): Bonifacio, Vivario.

Megachile fossoria Ferton; Liongo Li Enkulu (1988, Map 35) [quoting Ferton (1909a)].

Megachile leachella Curtis; Liongo Li Enkulu (1988, Map 40): Corsica.

Megachile leachella Curtis; Pagliano (1994: 372): Bastia.

Megachile leachella Curtis; Meunier et al. (2023): Ajaccio.

Megachile leachella Curtis; Praz and Bénon (2023, supplementary material): Aléria, Borgo, Ghisonaccia.

Material examined. 30♀ & 29♂ observed from May to September, between 1897 and 2021 in Bonifacio, Calenzana, Cervione, Corbara, Farinole, Ghisonaccia, Grosseto–Prugna, Linguizzetta, Palasca, Penta–di–Casinca, Propriano, Santo–Pietro–di–Tenda, Solaro, Talasani, Tallone, Ventiseri, Vico and Zonza.

Remark. Schwarz and Gusenleitner (2011) delineated several forms of M. leachella. In the context of France, two distinct forms can be recognized: the variety dorsalis Pérez, 1880, prevalent in Southern France, and the variety fossoria Ferton, 1905, found in Corsica. Baker (in Liongo Li Enkulu 1988) proposed considering M. fossoria as a valid species. Nevertheless, the extensive variability exhibited by M. leachella across its entire distribution range complicates the clear demarcation of taxa. Praz and Bénon (2023) thus maintained the synonymy of this taxon with M. leachella.

44. Megachile melanopyga melanopyga Costa, 1863

Megachile melanopyga Costa; Benoist (1931, 63): Corsica.

Megachile melanopyga Costa; Liongo Li Enkulu (1988, Map 47): Corsica.

Megachile melanopyga Costa; Ebmer (1997: 52): Tavignano.

Material examined. 68♀ & 44♂ observed from April to November, between 1895 and 2022 in Ajaccio, Aléria, Balogna, Bastelica, Bocognano, Bonifacio, Calenzana, Castellare–di–Casinca, Coti–Chiavari, Croce, Ghisonaccia, Ghisoni, Grosseto–Prugna, Linguizzetta, Manso, Mausoléo, Moncale, Nocario, Oletta, Olmi–Cappella, Palasca, Patrimonio, Piana, Porto–Vecchio, Quenza, Santo–Pietro–di–Tenda, Sartène, Sorbollano, Sotta, Talasani, Tavera, Ventiseri, Vivario and Zonza.

46. Megachile pusilla Pérez, 1884

Megachile pusilla Pérez; Ferton (1909a: 543): Bonifacio.

Megachile variscopa Pérez; Benoist (1940: 68): Corsica.

Megachile atratula Rebmann; Liongo Li Enkulu (1988, Map 24): Corsica.

Megachile albohirta (Brullé); Ornosa et al. (2007: 121) [Misinterpretation]: Corsica.

Material examined. 29♀ & 20♂ observed from May to October, between 1895 and 2021 in Biguglia, Bonifacio, L’Île Rousse, Lucciana, Palasca, Poggio–d’Oletta, Propriano, Saint–Florent, Santo–Pietro–di–Tenda, Talasani and Ventiseri.

47. Megachile rotundata (Fabricius, 1793)

Megachile rotundata (Fabricius); Pagliano (1994: 374): Col de Celaccia.

Material examined. 6♀ & 8♂ observed from May to July, between 2002 and 2021 in Bonifacio, Coti–Chiavari, Grosseto–Prugna, Olmi–Cappella, Palasca and Ventiseri.

49. Megachile sicula corsica Benoist, 1935

Figs 43–45

Megachile sicula var. perezi Lichtenstein; Friese (1899: 39, 176): Corsica.

Megachile sicula Rossi; Ferton (1909a: 550): Bonifacio.

Megachile perezi Lichtenstein; Ferton (1909a: 550): Bonifacio.

Megachile perezi Lichtenstein; Friese (1911: 212): Corsica.

Megachile sicula f. corsica Benoist var. nov.; Benoist (1935: 103).

Chalicodoma sicula var. corsica Benoist; Benoist (1940: 45): Bonifacio.

Chalicodoma sicula Rossi; Kusdas (1974: 160): Calvi.

Chalicodoma corsica Benoist; Liongo Li Enkulu (1988, Map 29): Corsica.

Chalicodoma sicula Rossi; Liongo Li Enkulu (1988: 87, Map 62): Corsica.

Chalicodoma sicula perezi Lichtenstein; Liongo Li Enkulu (1988, Map 64): Corsica.

Chalicodoma sicula Rossi; Bürgis (1995: 27): Calvi.

Material examined. 187♀, 69♂ & 1NS observed from March to August, between 1855 and 2021 in Ajaccio, Albertacce, Aléria, Asco, Bonifacio, Corte, Evisa, Farinole, Galéria, Mausoléo, Porto–Vecchio, Saint–Florent, Santo–Pietro–di–Tenda, Serra–di–Ferro, Solaro, Sotta, Ventiseri, Vivario and Zonza.

Remark. Following Tklaců’s suggestions, Liongo Li Enkulu (1988) formally recognized the Corso-Sardinian variety as a distinct species. Subsequently, Rasmont et al. (1995) and Ornosa et al. (2007) adhered to this classification, designating C. corsica as a valid species. This taxonomic distinction lacks substantive morphological or molecular justifications, with only variations in the coloration of pilosity and integument being established (Benoist 1935, 1940). It is acknowledged that considerable variability exists in coloration and pilosity among several species within the Chalicodoma subgenus. In light of these considerations, we currently regard C. corsica as a subspecies of M. sicula. It is noteworthy that the CO1 sequence from a Corsican specimen stands isolated in comparison to the sequences available on BOLD for M. sicula, displaying a notable divergence from other clusters. A more comprehensive investigation is imperative to ascertain the precise taxonomic status of this Corsican taxon concerning other subspecies of M. sicula.

50. Megachile versicolor Smith, 1844

Megachile versicolor Smith; Liongo Li Enkulu (1988, Map 65): Corsica.

Megachile versicolor Smith; Pagliano (1994: 374): Bastia.

Material examined. 3♂ observed from May to June, between 2003 and 2017 in Oletta and Vivario.

Osmiini Newman, 1834

Chelostoma Latreille, 1809*

51. Chelostoma distinctum Stöckhert, 1929*

Material examined. 25♀ & 32♂ observed from March to June, between 1906 and 2021 in Ajaccio, Bastelicaccia, Cauro, Evisa, Ghisonaccia, Grosseto–Prugna, Lozzi, Murzo, Pianottoli-Caldarello, Poggio–di–Venaco, Santo–Pietro–di–Tenda, Sorbollano, Sermano, Sotta, Venzolasca, Zigliara and Zonza.

Remark. The morphology of the Corsican specimens is slightly different than that of the other European specimens. The Corsican population might represent a distinct species.

52. Chelostoma foveolatum Schletterer, 1889*

Material examined. 6♀ observed in June 2003 in Grosseto–Prugna.

53. Chelostoma rapunculi Lepeletier, 1841*

Material examined. 9♀ & 12♂ observed from May to July, between 1897 and 2021 in Afa, Asco, Cateri, Corbara, Evisa, Grosseto–Prugna, Mausoléo, Olmi–Cappella, Quenza, Santa–Maria–Poggio, Sermano, Sorbollano, Vivario, Zigliara and Zonza.

Heriades Spinola, 1808

54. Heriades crenulata Nylander, 1856

Heriades crenulata Nylander; Benoist (1931: 132): Corsica.

Eriades crenulatus Nylander; Kusdas (1974: 160): Calvi.

Heriades crenulata Nylander; Pagliano (1994: 379): Bastia.

Heriades crenulata Nylauder; Marchal and Chardonnet (2001: 203): Serra–di–Fiumorbo.

Material examined. 247♀ & 265♂ observed from May to the beginning of November, between 1855 and 2021 in Ajaccio, Asco, Aullène, Bonifacio, Calenzana, Calvi, Cargèse, Corbara, Coti–Chiavari, Ersa, Farinole, Figari, Galéria, Ghisonaccia, Ghisoni, Grosseto–Prugna, Lecci, Linguizzetta, Mausoléo, Meria, Moncale, Oletta, Ota, Palasca, Patrimonio, Penta–di–Casinca, Piana, Poggio–di–Venaco, Poggio–d’Oletta, Porto–Vecchio, Propriano, Quenza, Saint–Florent, San–Martino–di–Lota, Santo–Pietro–di–Tenda, Serra–di–Ferro, Serra–di–Scopamène, Serriera, Sisco, Sorbollano, Sotta, Tavera, Ventiseri, Vivario and Zonza.

55. Heriades rubicola Pérez, 1890

Heriades rubicola Pérez; Benoist (1931: 132): Corsica.

Heriades rubicola Pérez; Pagliano (1994: 379): Venzolasca.

Material examined. 115♀ & 88♂ observed from May to October, between 1895 and 2021 in Aléria, Bonifacio, Calvi, Casaglione, Corbara, Ghisonaccia, Linguizzetta, Lumio, Moncale, Morosaglia, Ogliastro, Oletta, Palasca, Patrimonio, Penta–di–Casinca, Pietracorbara, Poggio–di–Venaco, Porto–Vecchio, Propriano, Saint–Florent, Santa–Maria–Poggio, Santo–Pietro–di–Tenda, Serra–di–Ferro, Talasani, Sisco, Venaco and Ventiseri.

56. Heriades truncorum (Linnaeus, 1758)

Heriades truncorum Linnaeus; Ferton (1901b: 93, 143): Propriano, Vivario.

Examined material. 30♀ & 20♂ observed from June to August, between 1896 and 2019 in Asco, Bonifacio, Evisa, Ghisoni, Mausoléo, Riventosa, Propriano, Serra–di–Ferro, Serra–di–Scopamène, Sorbollano, Vivario and Zicavo.

Hoplitis Klug, 1807

57. Hoplitis acuticornis (Dufour & Perris, 1840)

Osmia acuticornis Dufour & Perris; Benoist (1931: 41): Corsica.

Material examined. 31♀ & 12♂ observed from April to July, between 1895 and 2021 in Asco, Bonifacio, Borgo, Corte, Evisa, Mausoléo, Santo–Pietro–di–Venaco, Ventiseri, Vivario, Zicavo and Zonza.

Remark. The morphology of the Corsican specimens is slightly different than that of the other European specimens. The Corsican population might represent a distinct species.

58. Hoplitis aff. adunca (Panzer, 1798)*

Material examined. 23♀ & 38♂ observed from May to June, between 1895 and 1907 in Bonifacio, Porto–Vecchio and Propriano.

Remark. This species will be described soon (Le Divelec in prep.). It seems to be restricted to Southern Corsica and Sardinia. No modern record is known for Corsica.

59. Hoplitis adunca (Panzer, 1798)

Osmia adunca Panzer; Radoszkowski (1887: 288): Corsica.

Osmia adunca Panzer; Benoist (1931: 36): Corsica.

Osmia morawitzi Pérez; Benoist (1931: 36) [Misidentification]: Corsica.

? Osmia benoisti Alfken; Warncke (1992: 115) [probably quoting Benoist (1931)]: Corsica.

Material examined. 44♀ & 45♂ observed from April to August, between 1895 and 2021 in Aléria, Bonifacio, Cervione, Corbara, Corte, Galeria, Ghisonaccia, Grosseto–Prugna, Lucciana, Oletta, Piana, Poggio–di–Venaco, Saint–Florent, Santo–Pietro–di–Tenda, Tallone, Ventiseri, Venzolasca, Vico, Vivario and Zonza.

60. Hoplitis anthocopoides (Schenck, 1853)

Hoplitis anthocopoides Schenck; Scheuchl and Willner (2016: 435) [quoting Müller (2022a)].

Hoplitis anthocopoides Schenck; Müller (2022a): Corsica.

Material examined. 104 ♀ & 82♂ observed from May to August, between 1895 and 2021 in Ajaccio, Aléria, Aregno, Bonifacio, Borgo, Calvi, Casaglione, Centuri, Cervione, Corbara, Corscia, Corte, Ersa, Ghisonaccia, Grosseto–Prugna, L’Île Rousse, Lucciana, Lumio, Oletta, Palasca, Piana, Poggio–di–Venaco, Propriano, Santo–Pietro–di–Tenda, Serra–di–Ferro, Sotta, Ventiseri, Vico, Vivario and Zonza.

61. Hoplitis bihamata (Costa, 1885)

Figs 51–53

Osmia corsica Ferton sp. nov.; Ferton (1901a: 61–63): Bonifacio, Monte Renoso.

Osmia corsica Ferton; Benoist (1931: 36): Corsica.

Osmia corsica Ferton; Nadig and Nadig (1934: 26): Monte d’Oro.

Osmia corsica Ferton; Stanek (1969: 28): Corsica.

Osmia marchali Pérez; Warncke (1992c: 109) [misidentification]: Col de Verde.

Material examined. 83♀ & 69♂ from April to September, between 1898 and 2021 in Asco, Barrettali, Bocognano, Bonifacio, Calvi, Casaglione, Corte, Evisa, Ghisoni, Mausoléo, Olcani, Patrimonio, Porto–Vecchio, Quenza, Santo–Pietro–di–Tenda, Serra–di–Scopamène, Sisco, Vivario and Zonza.

Remark. Corso-sardinian endemic.

62. Hoplitis bisulca (Gerstäcker, 1869)

Figs 59–61

Osmia lanosa Pérez; Ferton (1897: 42): Bonifacio.

Osmia lanosa Pérez; Ferton (1901b: 88–89): Bonifacio.

Osmia lanosa Pérez; Ferton (1905: 58–59): Bonifacio.

Osmia lanosa Pérez; Ferton (1909a: 538): Bonifacio.

Osmia bisulca Gerstäcker; Benoist (1931: 33): Corsica.

Material examined. 97♀ & 70♂ observed from May to August, between 1895 and 2021 in Bonifacio, Canale–di–Verde, Patrimonio and Zonza.

63. Hoplitis corsaria (Warncke, 1991), stat. nov.

Osmia crenulata Morawitz; Morawitz (1871: 208): Corsica.

Osmia crenulata Morawitz; Schmiedeknecht (1885–1886: 161): Corsica.

Osmia crenulata Morawitz; Ducke (1900: 171): Corsica.

Osmia crenulata Morawitz; Ferton (1901a: 63): Bonifacio.

Osmia crenulata Morawitz; Benoist (1931: 35): Corsica.

Osmia annulata corsaria Warncke ssp. nov.; Warncke (1991a: 734): Ajaccio.

Osmia annulata corsaria Warncke; Le Goff (2010: 7): Grosseto–Prugna.

Type material examined. Holotype: Corse 15-6-1981 [15.VI.1981] Ajaccio leg. Perraudin // Holotypus Osmia annulata corsaria War // Holotypus // ♂ // Coll. K. Warncke O. Ö. Landesmuseum Linz/Austria-egg.93. [Examined on pictures: https://www.europeana.eu/fr].

Material examined. 151♀ & 114♂ observed from May to July, between 1895 and 2021 in Ajaccio, Aléria, Aullène, Bonifacio, Canale–di–Verde, Coggia, Corbara, Ghisonaccia, Grosseto–Prugna, Lumio, Oletta, Palasca, Porto–Vecchio, Propriano, Sartène, Serra–di–Ferro, Sollacaro, Ventiseri, Venzolasca, Vico, Vivario and Zonza.

Remark. Old European records of Osmia crenulata (Morawitz, 1871) refer to H. annulata (Latreille, 1811) and not specifically to the East Mediterranean H. annulata crenulata. Corsican records only refer to H. annulata corsaria. Hoplitis annulata is a species complex. The morphology of the Corsican endemic H. a. corsaria, especially that of the male, is remarkably distinct from that of the two other subspecies (Table 3). Plus, the CO1 sequences of the Corsican specimens are significantly diverging (around 3.7%) from the Iberian sequences of H. annulata published on BOLD. For these reasons, we consider H. corsaria to be a distinct species. The nominative subspecies and H. a. crenulata might also represent distinct species as they are morphologically different and have distinct nesting behaviour (Le Goff, 2010). However, the morphological differences are tenuous and without the examination of an extensive material or molecular evidence we cannot exclude intraspecific variability. Hoplitis corsaria is only known from Corsica to date.

Table 3.

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Main morphological differences between H. corsaria and H. annulata.

Features	H. corsaria	H. a. annulata	H. a. crenulata
Gena punctation	♀: Gena with shallow punctation, more sparsely spaced by uneven interspaces that can reach the diameter of one puncture.	♀: Gena with coarse and dense subcontiguous punctation, with narrow linear interspaces.	♀: Gena with coarse and dense subcontiguous punctation, with narrow linear interspaces.
Gena punctation	♂: Gena with remarkably minute and sparse punctuation, its lower half with large and almost impunctate area near outer orbit (Fig. 17).	♂: Gena with coarse subcontiguous punctuation, its lower half uniformly punctate, at most with a small impunctate area (Fig. 14).	♂: Gena with coarse subcontiguous punctuation, its lower half densely punctate, at most with a small impunctate area (Fig. 20).
Vertex length	Around two times the diameter of posterior ocellus	Around two times the diameter of posterior ocellus	More than two times the diameter of posterior ocellus
Mesepisternum punctation	Punctation sparse, punctures separated by conspicuous flat interspaces (dense around scrobal area) (Fig. 19).	Punctation subcontiguous, mostly with linear to carina-shaped interspaces (Fig. 16).	Punctation subcontiguous, mostly with carina-shaped interspaces (Fig. 22).
Metasoma	Punctation of terga 2–4 fine and remarkably sparse (most interspaces > 1–2 puncture diameters) with conspicuous micropunctation in-between, tegument smooth and shiny (Fig. 18).	Punctation of terga 2–4 denser (interspace ≤ 1 puncture diameter) and somewhat coarser so the segment appears matte, micropunctation hardly distinct, tegument of terga smooth, at most slightly shagreened basally (Fig. 15).	Punctation of terga 2–4 denser (interspace ≤ 1 puncture diameter) and coarser so the segment appears matte, micropunctation hardly distinct, tegument of terga shagreened, at least basally (Fig. 21).

Figures 14–22.

Comparative illustrations of H. corsaria (Warncke) and H. annulata (Latreille) 14–16 H. annulata annulata (Latreille) (from Spain) 17–19 H. corsaria (Warncke) (from Corsica) 20–22 H. annulata crenulata (Morawitz) (from Greece) 14, 17, 20 gena of male 15, 18, 21 gaster of female (15 terga 2-4 18 terga 1-4 21 terga 1-3) 16, 19, 22 mesepisternum. Scale bars: 1 mm.

BOLD process ID. LPRCW186-19, LPRCW187-19, LPRCI1815-21, LPRCI1824-21, LPRCI1968-21 (BIN, BOLD:AEC2169).

64. Hoplitis cristatula (van der Zanden, 1990)

Osmia cristata Fonscolombe; Ferton (1897: 42): Bonifacio.

Osmia cristata Fonscolombe; Ferton (1901b: 88): Bonifacio.

Osmia cristata Fonscolombe; Ferton (1909a: 538): Bonifacio.

Osmia cristata Fonscolombe; Benoist (1931: 34): Corsica.

Material examined. 48♀ & 34♂ observed from May to July, between 1895 and 2021 in Ajaccio, Aléria, Bonifacio, Calenzana, Casamaccioli, Coggia, Farinole, Grosseto–Prugna, L’Île Rousse, Ogliastro, Oletta, Santa–Maria–Poggio, Santo–Pietro–di–Tenda, Serra–di–Ferro, Vivario and Zonza.

65. Hoplitis fasciculata (Alfken, 1934)

Figs 23–28

Hoplitis fasciculata (Alfken); Müller (2022a): Corsica.

Material examined. 13♀ & 11♂ observed from June to July, between 1899 and 2021 in Bonifacio, Propriano, Prunelli–di–Fiumorbo and Rutali.

Remark. The generic Corsican record of H. fasciculata mentioned on the website of Müller (2022a) comes from our records that are here published for the first time. Corsica represents the western limit of the distribution of this species. The female is easily distinguishable from other Anthocopa Lepeletier & Serville, 1825 by the white scopa (Fig. 23), the crenulate clypeus free margin (saw-edged) (Fig. 25) and the rugose areolate sculpture of the scutum (Fig. 27). The male can be recognized by the conspicuously bilobed tergum 7 (Fig. 28), the bidentate mandibule (Fig. 26). Its last sternum has a rounded apical margin and is apically depressed with a smooth and shiny area at the end of which is a small tubercle bearing a tuft of backwardly directed setae (Fig. 28).

Figures 23–28.

Hoplitis fasciculata (Alfken) 23, 25, 27 female 24, 26, 28 male 23, 24 habitus in lateral view 25, 26 head in front view 27 habitus in dorsal view 28 gaster in ventral view. Scale bars: 1 mm

66. Hoplitis leucomelana (Kirby, 1802)

Osmia leucomelana Kirby; Warncke (1991b: 716): Ajaccio.

Hoplitis leucomelana (Kirby); Pagliano (1994: 383): Col de Celaccia.

Hoplitis leucomelana Kirby; Le Goff (2004: 11): Vivario.

Material examined. 33♀ & 25♂ observed from June to September, between 1896 and 2020 in Albertacce, Aullène, Biguglia, Bonifacio, Calacuccia, Casamaccioli, Coti–Chiavari, Evisa, Grosseto–Prugna, Oletta, Patrimonio, Piana, Sermano, Serra–di–Ferro, Serra–di–Fiumorbo, Sisco, Sorbollano, Tavera, Venaco, Vico, Vivario, Zigliara and Zonza.

67. Hoplitis manicata (Morice, 1901)

Hoplitis manicata Morice; Le Goff (2004: 19): Vivario (2B).

Material examined. 9♀ & 5♂ observed in June 2002 and 2003 in Vivario.

Remark. In Corsica, this species is only known from one station in the mountains of Vivario (Le Goff 2004). Corsica represents the western limit of the distribution of this species.

68. Hoplitis perezi (Ferton, 1895)

Osmia perezi Ferton; Ferton (1897: 42–43): Bonifacio.

Osmia perezi Ferton; Ferton (1901b: 89–90): Bonifacio.

Osmia perezi Ferton; Ferton (1909a: 538): Bonifacio.

Osmia perezi Ferton; Benoist (1931: 35): Corsica.

Anthocopa perezi (Ferton); Tkalců (1969: 330): Bonifacio.

Material examined. 31♀ & 19♂ observed from May to July, between 1895 and 2021 in Bonifacio, Casamaccioli, Corte, Grosseto–Prugna and Sermano.

69. Hoplitis praestans (Morawitz, 1893)

Figs 63–66

Osmia praestans Morawitz; Ducke (1900: 110): Corsica.

Osmia lineola Pérez; Ferton (1901a: 63): Corsica.

Osmia praestans Morawitz; Friese (1911: 80): Corsica.

Osmia praestans Morawitz; Benoist (1931: 42): Corsica.

Osmia praestans Morawitz; Kusdas (1974: 160): Calvi.

Osmia praestans Morawitz; Warncke (1991b: 734): Calvi.

Material examined. 13♀ & 20♂ observed from May to July, between 1855 and 2021 in Bonifacio, Calvi, Ghisonaccia, Ghisoni, Linguizzetta, Mausoléo, Pianottoli-Caldarello, Ventiseri, Vivario and Zonza.

70. Hoplitis ravouxi (Pérez, 1902)

Hoplitis ravouxi (Pérez); Müller (2022a): Corsica.

Hoplitis loti Morawitz; Le Goff (2004: 11) [Misidentification]: Vivario.

Material examined. 1♀ observed in June 2003 in Vivario & 1♂ in May 2011 in Balogna.

71. Hoplitis tridentata (Dufour & Perris, 1840)

Osmia mocsaryi Friese; Ducke (1900: 154) [Misinterpretation]: Corsica.

Osmia mocsaryi Friese; Friese (1911: 92) [quoting Ducke (1900)]: Corsica.

Osmia mocsaryi Friese; Benoist (1931: 38) [quoting Ducke (1900)]: Corsica.

Material examined. 5♀ & 6♂ observed from May to June, between 1897 and 2021 in Borgo, Lucciana, Oletta, Sermano, Vescovato and Vivario.

Remark. This species has been recorded from Corsica under the name of O. mocsaryi, a species with which H. tridentata was regularly confused in the past (see below).

Osmia Panzer, 1806

72. Osmia anceyi Pérez, 1879

Osmia anceyi Pérez; Müller (2018: 312): Verghia.

Material examined. 1♀ collected in June 2001 in Coti-Chiavari.

73. Osmia aurulenta (Panzer, 1799)

Osmia aurulenta (Panzer); Müller (2022a): Corsica.

Material examined. 33♀ & 5♂ observed from April to July, between 1897 and 2021 in Barbaggio, Bocognano, Ghisoni, Grosseto–Prugna, Lozzi, Lucciana, Palasca, Riventosa, Sartène, Sermano, Vico and Vivario.

74. Osmia bicornis bicornis (Linnaeus, 1758)

Figs 29, 30

Osmia bicornis Latr.; Ferton (1911: 368): Bonifacio.

Osmia rufa rufa (Linnaeus); Peters (1977: 291): Ajaccio, Bastia, Calvi, Corte, Nonza, Monte d’Oro, Porto–Vecchio, Saint–Florent, Santo–Pietro–di–Venaco, Sari-Solenzara, Vivario.

Figures 29–34.

Remarkable variety of Corsican Osmia 29, 30 Osmia bicornis bicornis (Linnaeus) 31, 32 Osmia latreillei iberoafricana Peters 33, 34 Osmia versicolor corrusca Erichson 29, 31, 32 lateral view 30, 32, 34 dorsal view. Scale bars: 1 mm.

Osmia bicornis globosa (Scopoli); Ungricht et al. (2008: 177): Corsica.

Material examined. 60♀ & 47♂ observed from March to June, between 1897 and 2021 in Bocognano, Bonifacio, Calacuccia, Casaglione, Corte, Ghisonaccia, Riventosa, Porto–Vecchio, Quenza, Serra–di–Ferro, Serra–di–Scopamène, Sollacaro, Sotta, Ventiseri, Vico, Vivario and Zonza.

75. Osmia caerulescens caerulescens Linnaeus, 1758

Osmia aenea (Linnaeus); Ferton (1897: 40): Bonifacio.

Osmia cyanea (Fabricius); Ferton (1905: 93): Bonifacio.

Osmia aenea (Linnaeus); Ferton (1911: 368): Bonifacio.

Material examined. 222♀ & 109♂ observed from March to September, between 1855 and 2021 in Ajaccio, Albitreccia, Aléria, Asco, Aullène, Balogna, Biguglia, Bocognano, Bonifacio, Calenzana, Casaglione, Casamaccioli, Casanova, Centuri, Coggia, Corbara, Coti–Chiavari, Cozzano, Evisa, Farinole, Furiani, Galéria, Ghisonaccia, Ghisoni, Grosseto–Prugna, Linguizzetta, Lozzi, Lucciana, Mausoléo, Oletta, Olmi–Cappella, Palasca, Palneca, Patrimonio, Penta–di–Casinca, Pianottoli-Caldarello, Poggio–di–Venaco, Porto–Vecchio, Propriano, Prunelli–di–Fiumorbo, Quenza, Riventosa, Rogliano, Rutali, Saint–Florent, Santa–Maria–Poggio, Santo–Pietro–di–Tenda, Serra–di–Ferro, Serra–di–Fiumorbo, Sisco, Sorbollano, Sorbo–Ocagnano, Sotta, Talasani, Ventiseri, Vescovato, Vico, Vivario, Zicavo and Zonza.

76. Osmia cornuta cornuta (Latreille, 1805)

Osmia cornuta (Latreille); Peters (1977: 337): Corsica.

Material examined. 3♀ & 3♂ observed from March to June, between 1976 and 2021 in Borgo, Grosseto–Prugna, Riventosa and Sotta.

77. Osmia emarginata emarginata Lepeletier, 1841

Osmia emarginata Lepeletier; Ferton (1901b: 85, 89): Evisa, Vivario.

Osmia emarginata emarginata Lepeletier; Tkalců (1971: 224): Monte d’Oro, Vivario.

Osmia emarginata emarginata Lepeletier; Peters (1977: 310): Quenza, Rutali, Vivario.

Material examined. 12♀ & 7♂ observed from May to July, between 1897 and 2021 in Corte, Evisa, Lozzi, Serra–di–Ferro, Vivario, Zicavo and Zonza.

78. Osmia erythrogastra Ferton, 1905

Figs 55–57

Osmia erythrogastra Ferton sp. nov.; Ferton (1905: 57): Bonifacio.

Osmia erythrogastra Ferton; Ferton (1909b: 407): Bonifacio.

Osmia erythrogastra Ferton; Benoist (1931: 33): Bonifacio.

Osmia erythrogastra Ferton; Müller (2020: 230): Bastia.

Material examined. 10♀ & 8♂ observed from May to August, between 1904 and 1964 in Bastia and Bonifacio.

Remark. No modern records.

79. Osmia ferruginea Latreille, 1811

Osmia igneopurpurea Costa; Ferton (1897: 39): Bonifacio.

Osmia ferruginea Lep.; Ferton (1899: 72): Bonifacio.

Osmia ferruginea Latreille; Ducke (1900: 212): Corsica.

Osmia ferruginea Lep.; Ferton (1901a: 65): Corsica.

Osmia ferruginea Lep.; Ferton (1901b: 143): Bonifacio.

Osmia ferruginea Lep.; Ferton (1905: 89–93): Bonifacio.

Osmia ferruginea Lep.; Ferton (1909a: 575, 577): Bonifacio.

Osmia ferruginea var. igneopurpurea Costa; Benoist (1931: 32): Corsica.

Osmia ferruginea igneopurpurea Costa stat. nov.; Warncke (1992d: 109): Corsica.

Material examined. 65♀ & 35♂ observed from March to June, between 1855 and 2020 in Barbaggio and Bonifacio.

Remark. The subspecies igneopurpurea Costa, 1882, has been documented in Corsica, Sardinia, Sicily, and Malta (Warncke 1992d). This subspecies is characterized by a dark scopa and dark hairs on the hind tibia. However, Corsican specimens exhibit significant variability, ranging from typical characteristics of the nominative subspecies to those of the subspecies igneopurpurea. The presence of numerous intermediate specimens further complicates the differentiation between subspecies. Given this variability, we suggest considering both dark and light hairy Corsican specimens of O. ferruginea as part of a single taxon. The subspecies igneopurpurea may just be a synonym of Osmia ferruginea.

80. Osmia latreillei (Spinola, 1806)

Figs 31, 32

Osmia latreillei (Spinola); Ducke (1900: 236): Corsica.

Osmia latreillei Lep.; Ferton (1905: 59, 93): Bonifacio.

Osmia latreillei (Spinola); Benoist (1931: 30): Corsica.

Osmia latreillei iberoafricana Peters; Tkalců (1975b: 184): Corsica.

Osmia latreillei latreillei (Spinola); Ungricht et al. (2008: 153): Corsica.

Material examined. 35♀ & 47♂ observed from March to June, between 1895 and 2021 in Ajaccio, Bonifacio, Corte, Ghisonaccia and Riventosa.

Remark. Tkalců (1975b) assigned the Corso-Sardinian populations to the subspecies iberoafricana. However, the situation remains unclear as he noted that these populations exhibit intermediate characteristics, with some specimens closely resembling the nominate subspecies found in mainland France and Italy. Müller (2022a) argues that a subspecies rank is unjustified and proposes its synonymy with O. latreillei.

81. Osmia ligurica Morawitz, 1868

Osmia ligurica Morawitz; Kusdas (1974: 160): Calvi.

Material examined. 76♀ & 40♂ observed from May to August, between 1895 and 2021 in Albertacce, Aléria, Asco, Barbaggio, Bonifacio, Calacuccia, Castellare–di–Mercurio, Corte, Ghisonaccia, Grosseto–Prugna, Lumio, Mausoléo, Moncale, Oletta, Palasca, Patrimonio, Pianottoli-Caldarello, Porto–Vecchio, Propriano, Santo–Pietro–di–Tenda, Sermano, Serra–di–Ferro, Sisco, Sotta, Venaco, Ventiseri, Vivario and Zonza.

82. Osmia melanogaster melanogaster Spinola, 1808

Osmia notata (Fabricius); Benoist (1931: 28) [Misinsterpretation, see Tkalcu 1975a]: Corsica.

Material examined. 2♂ collected in Ajaccio during the XIXth century (J. Pérez coll.).

Remark. No modern records.

83. Osmia nasoproducta Ferton, 1909

Figs 68–70

Osmia nasoproducta Ferton sp. nov.; (Ferton 1909b: 406–407): Bonifacio.

Material examined. 8♀ & 4♂ observed from March to June, between 1902 and 2017 in Bonifacio.

Remark. It is a rare and poorly known species. Its male was recently described (Le Goff 2016). The original description (Ferton 1909b) represented the only Corsican record until now.

84. Osmia niveata (Fabricius, 1804)

Osmia fulviventris var. albiscopa Alfken; Benoist (1931: 29): Corsica.

Osmia fulviventris Panzer; Kusdas (1974: 160): Calvi.

Osmia fulviventris niveata (Fabricius): Tkalců (1975a: 307): Bastia, Porto–Vecchio, Ville–di–Pietrabugno.

Material examined. 114♀ & 53♂ observed from March to July, between 1896 and 2021 in Altagène, Bonifacio, Calenzana, Calvi, Centuri, Coggia, Corbara, Ersa, Grosseto–Prugna, Lozzi, Manso, Mausoléo, Olcani, Oletta, Osani, Palasca, Patrimonio, Pianottoli-Caldarello, Poggio–di–Venaco, Porto–Vecchio, Prunelli–di–Fiumorbo, Quenza, Riventosa, Santo–Pietro–di–Tenda, Serra–di–Ferro, Sorbo–Ocagnano, Sotta, Venzolasca, Vivario and Zonza.

Remark. Corsican females, characterized by a pure white scopa, were previously considered a distinct subspecies named O. niveata albiscopa. However, the males are not distinguishable, and the barcode of Corsican specimens cannot be distinguished from other European counterparts. This suggests that the light hairy insular population is conspecific with mainland populations.

85. Osmia rufohirta Latreille, 1811

Osmia rufohirta Latreille; Ferton (1897: 37–39): Bonifacio.

Osmia rufohirta Latreille; Ferton (1899: 70–72): Bonifacio.

Osmia fossoria Pérez; Ducke (1900: 127) [Misinterpretation]: Corsica.

Osmia rufohirta Latreille; Ferton (1901b: 89, 145): Bonifacio.

Osmia rufohirta Latreille; Ferton (1905: 62, 83–95): Bonifacio.

Osmia rufohirta Latreille; Ferton (1909a: 579–580): Bonifacio.

Osmia rufohirta Latreille; Ferton (1911: 381–382): Bonifacio.

Osmia rufohirta Latreille; Benoist (1931: 40): Corsica.

Material examined. 101♀ & 62♂ observed from March to the beginning of July, between 1893 and 2021 in Bonifacio, Corte, Farinole, Patrimonio, Sermano and Vivario.

86. Osmia scutellaris Morawitz, 1868

Osmia scutellaris Morawitz; Le Goff (2004: 11): Vivario.

Osmia scutellaris Morawitz; Müller (2018: 320): Zonza.

Material examined. 19♀ & 19♂ observed from May to the beginning of July, between 1973 and 2021 in Bonifacio, Calacuccia, Casaglione, Grosseto–Prugna, Oletta, Palasca, Riventosa, Porto–Vecchio, Quenza, Santo–Pietro–di–Tenda, Sermano, Sotta, Vivario and Zonza.

87. Osmia signata signata Erichson, 1835

Osmia vidua Gerstäcker; Ferton (1897: 40–42): Bonifacio.

Osmia vidua Gerstäcker; Ferton (1901a: 64): Bonifacio.

Osmia vidua Gerstäcker; Benoist (1931: 27): Corsica.

Material examined. 34♀ & 20♂ observed from May to August, between 1895 and 2021 in Ajaccio, Bonifacio, Calenzana, Calvi, Coti–Chiavari, Grosseto–Prugna, Mausoléo, Olcani, Patrimonio, Porto–Vecchio, Propriano, Prunelli–di–Fiumorbo, Sartène, Serra–di–Ferro, Sollacaro and Sotta.

88. Osmia submicans submicans Morawitz, 1870

Osmia submicans submicans Morawitz; Tkalců (1977: 94): Bonifacio.

Osmia submicans Morawitz; Zanden (1991: 65): Ajaccio, Saint–Florent.

Material examined. 51♀ & 16♂ observed from March to July, between 1898 and 2021 in Bonifacio, Galéria, Ghisonaccia, Ghisoni, Mausoléo, Muracciole, Oletta, Olmi–Cappella, Palasca, Riventosa, Porto–Vecchio, Sotta, Ventiseri, Vivario and Zonza.

89. Osmia tricornis Latreille, 1811

Osmia tricornis Latreille; Ferton (1901b: 97): Bonifacio.

Osmia tricornis Latreille; Ferton (1911: 368): Bonifacio.

Osmia tricornis Latreille; Benoist (1931: 23): Corsica.

Osmia tricornis Latreille; Kusdas (1974: 160): Calvi.

Osmia tricornis Latreille; Peters (1977: 337): Corsica.

Material examined. 12♀ & 25♂ observed from March to April, between 1896 and 2021 in Bonifacio, Calenzana, Corte and Santo–Pietro–di–Tenda.

90. Osmia versicolor corrusca Erichson, 1835

Figs 33, 34

Osmia versicolor Latreille; Ferton (1905: 85): Bonifacio.

Osmia versicolor Latreille; Kusdas (1974: 160): Calvi.

Osmia versicolor corrusca Erichson stat.nov.; Zanden (1984: 183): Bonifacio.

Material examined. 72♀ & 15♂ observed from February to the beginning of July, between 1895 and 2020 in Barbaggio, Bonifacio, Ghisonaccia and Patrimonio.

Remark. Osmia v. corrusca is allegedly restricted to Corsica, the Balearic Islands, the Iberian Peninsula and to North Africa (Warncke 1992d).

Protosmia Ducke, 1900*

91. Protosmia minutula (Pérez, 1896)*

Material examined. 1♂ observed in July 2019 in Mausoléo.

Dubious records

The Corsican records of Megachile marginata Smith, 1853 (as Megachile picicornis Morawitz), M. pyrenaea Pérez, 1890), M. pyrenaica Lepeletier, 1841 published by Liongo Li Enkulu (1988) could not be confirmed and are likely to be mistakes. The Corsican record of H. claviventris (Thomson, 1872) published by Warncke (1988) most probably refers to H. leucomelana, a very common species in Corsica which is morphologically very similar. Finally, the presence of O. notata in Corsica is documented by Nadig and Nadig (1934) and Warncke (1988) who probably quoted the previous authors. These records most probably refer to the closely related O. signata, a common species in Corsica.

Erroneous records

Hoplitis benoisti (Alfken, 1935)

Osmia morawitzi Pérez; Benoist (1931: 36): Corsica.

Osmia benoisti Alfken; Warncke (1992: 115): Corsica.

Remark. The specimens reported by Benoist (1931) are preserved in his collection and belong to H. adunca. The Corsican record of Warncke (1992) is either a quotation of Benoist (1931) or refer to the new species, H. aff. adunca.

Hoplitis loti (Morawitz, 1867)

Osmia loti Morawitz; Ferton (1901a: 64): Bonifacio.

Hoplitis loti Morawitz; Le Goff (2004: 11) [= H. ravouxi (Pérez), Le Goff pers. comm.].

Remark. No such specimen has been located in Ferton’s collection. It is a mountainous species, unlikely to occur in Bonifacio.

Hoplitis marchali (Pérez, 1902)

Osmia marchali Pérez; Warncke (1992c: 109): Col de Verde.

Remark. The specimen reported by Warncke (1992c) is preserved in his collection and belong to H. bihamata. Hoplitis marchali is present in the south of the Iberian Peninsula, North Africa, and Sicily (Warncke 1992c; Baldock et al. 2018).

Hoplitis mocsaryi (Friese, 1895)

Osmia mocsaryi Friese; Ducke (1900: 154): Corsica.

Remark. Despite significant samplings in Corsica, no specimen of. H. mocsaryi were observed. The specimen(s) originally recorded from Corsica could not be located. The original record of Ducke (1900) was quoted by many authors (e.g. Friese 1911; Benoist 1931; Moczar 1958; Pagliano 1994; Rasmont et al. 1995, 2017; Ornosa et al. 2007; Scheuchl and Willner 2016). However, Friese, Ducke and Benoist misinterpreted the identity of the female of H. mocsaryi which they confused with H. tridentata. The female of H. mocsaryi was actually described for the first time by Noskiewicz (1934). According to Noskiewicz (1934), Ducke also misidentified many males. Therefore, it seems more likely that H. mocsaryi was confused with H. tridentata.

Megachile albohirta (Brullé, 1839)

Megachile albohirta (Brullé); Ornosa et al. (2007: 121) [Misinterpretation]: Corsica.

Remark. The Corsican record of Megachile albohirta in Ornosa et al. (2007) is a mistake that probably comes from a misinterpretation of the Corsican record of M. variscopa Pérez, 1895 by Benoist (1940). Following Tkalců (1993), Ornosa et al. (2007) considered M. variscopa as a junior synonym of M. albohirta. However, they are two distinct species (Gonzalez et al. 2010; Praz 2017). Megachile albohirta is restricted to North Africa and Canary Islands (Brullé 1839; Nadig and Nadig 1933; Tkalců 1993) while M. pusilla (senior synonym of M. variscopa, see Praz and Bénon 2023) occurs in Western Europe (Soltani et al. 2017).

Megachile lefebvrei Lepeletier, 1841

Megachile lefebvrei Lepeletier; Ferton (1920: 336): Bonifacio.

Chalicodoma lefebvrei lefebvrei Lepeletier; Tkalců (1975b: 187): Corsica.

Remark. No Corsican specimen was collected or examined during this study. M. lefebvrei, was vaguely reported from Corsica by Tkalců (1975b) who cites the dark form M. lefebvrei lefebvrei. Since no material was presented by the author, this could either be an error in the text or based on the paper of Ferton (1920). Ferton (1920) mentions an observation of M. lefebvrei in Bonifacio on July 31, 1915. However, only one Hymenoptera collected in 1915 is in his collection, suggesting that Ferton likely ceased his regular field trips in Corsica from 1915 onwards. Additionally, the date of observation seems unlikely. Upon comparing Ferton (1920) with the corresponding specimens in his collection, it becomes evident that he frequently switched the years 1915 and 1914. This discrepancy is not surprising, considering Ferton’s passing during the preparation of the publication, leading to an incomplete revision of the draft. Finally, the entire publication focuses on Ferton’s collecting activities in Algeria and occasionally in Provence. It appears more plausible to us that the specimen was observed in Nemours, where Ferton was definitely present in July 1914 and where the nominative form of M. lefebvrei is known to occur. In his box from the year 1914, a female of M. lefebvrei from Nemours is present with a label referring to his lost manuscript, where he had evidently recorded the observations reported in his publication.

Megachile sicula sicula (Rossi, 1792) and Megachile s. perezi Lichtenstein, 1880

Megachile sicula var. perezi Lichtenstein; Friese (1899: 39, 176): Corsica.

Megachile sicula Rossi; Ferton (1909a: 550): Bonifacio.

Megachile perezi Lichtenstein; Ferton (1909a: 550): Bonifacio.

Megachile perezi Lichtenstein; Friese (1911: 212): Corsica.

Chalicodoma sicula Rossi; Liongo Li Enkulu (1988, p.87, Map 62): Corsica.

Chalicodoma sicula perezi Lichtenstein; Liongo Li Enkulu (1988, Map 64): Corsica.

Remark. Both M. sicula and M. perezi are mentioned in Corsica by Ferton and Friese. The record of M. perezi is a mistake as no specimen of M. s. perezi has been found in Ferton’s collection. Liongo Li Enkulu (1988) also notes Corsican records for C. corsica, along with instances of C. sicula sicula (Rossi, 1792) and C. s. perezi (Lichtenstein, 1880). These two taxa are very close to C. s. corsica and are prone to confusion, particularly among males with highly variable pilosity. We believe that all these records exclusively pertain to C. s. corsica, which is the sole taxon we have observed on the island.

Osmia caerulescens cyanea (Fabricius, 1793)

Osmia cyanea (Fabricius); Ferton (1905: 93): Bonifacio.

Remark. Ferton labelled his specimens of O. caerulescens with the name cyanea. The mention of Osmia caerulescens cyanea (Fabricius, 1793) is erroneous. This subspecies is strictly North African (Tkalců 1970).

Osmia cyanoxantha Pérez, 1879

Osmia cyanoxantha Pérez; Benoist (1931: 31): Corsica.

Remark. No specimen of O. cyanoxantha has been located in MNHN collections. We believe that the record of Benoist resulted from a misidentification.

Osmia kohlii Ducke, 1899

Osmia kohlii Ducke; Ferton (1905: 59) [Misinterpretation, see Peters (1977)]: Bonifacio.

Remark. No such specimen is in Ferton’s collection. The Corsican record has moreover been invalidated by Peters (1977) who consider this species to be restricted to southern Italy and Sicily. It also occurs in Malta according to Müller (2022a).

Osmia lhotelleriei Pérez, 1887

Osmia fossoria Pérez; Ducke (1900: 127) [Misinterpretation]: Corsica.

Remark. Osmia fossoria Pérez, junior synonym of O. lhotelleriei, is restricted to North Africa and Levant (Müller 2022). The record of Ducke (1900) refers to O. rufohirta.

Types of Megachilid bees described from Corsica and housed at MNHN

Megachile argentata var. fossoria Ferton, 1909

Figs 35–38

Megachile argentata var. fossoria Ferton, 1909a: p.550. Corsica: Bonifacio, Propriano (Lectotype designated by Schwarz and Gusenleitner (2011), MNHN).

Type material examined. Lectotype: ♀, MNHN Paris EY0000001710 // Terrain sable Propriano 20/6 99 [20.VI.1899] // Museum Paris Coll. Ferton // Lectotypus Megachile argentata fossoria Ferton M. Schwarz des. 2009 // = Megachile leachella fossoria Ferton, 1909 Le Divelec det. 2020; Paralectotypes: ♀, MNHN Paris EY0000001711 // Bonifacio 4/6 99 [04.VI.1899] dans nid page 286 Megachile // Museum Paris Coll. Ferton // Paralectotypus Megachile argentata fossoria Ferton M. Schwarz des. 2009 // = Megachile leachella fossoria Ferton, 1909 Le Divelec det. 2020; ♀, MNHN Paris EY0000001712 // Bonifacio 18/7 97 [18.VII.1897] dans nid trou sable feuilles collées-miel liquide œuf baignant page 190 argentata // Museum Paris Coll. Ferton // Paralectotypus Megachile argentata fossoria Ferton M. Schwarz des. 2009 // = Megachile leachella fossoria Ferton, 1909 Le Divelec det. 2020.

Figures 35–38.

Lectotype of Megachile argentata var. fossoria Ferton 35 dorsal view 36 lateral view 37 head in front view 38 labels. Scale bars: 1 mm.

Current status. Megachile (Eutricharaea) leachella Curtis, 1828.

Megachile lucidifrons Ferton, 1905

Megachile (Chalicodoma) lucidifrons Ferton, 1905: p.57, ♀. Corsica: Bonifacio (Type lost).

Remark. Megachile lucidifrons has been consistently listed in the works on French Megachile (Benoist 1935, 1940; Liongo Li Enkulu 1988; Rasmont et al. 1995) despite its uncertain identity (Ghisbain et al. 2023). It was described based on a single worn female from Bonifacio and belongs to the lefebvrei species-group (Benoist 1935). The holotype was last examined by Benoist (1940) but despite exhaustive research in MNHN collections and consultation of the loan logs, its whereabouts remain unknown. In a correspondence with B. Tkalců, who sought to borrow the holotype, J. Kelner-Pillault (former curator of the Hymenoptera collections) mentioned her inability to locate it. The type is also not present in B. Tkalců’s collection (M. Schwarz, pers. comm.) and can be considered lost. A series of entirely black individuals of Megachile belonging to the lefebvrei species-group and matching the descriptions of Ferton (1905) and Benoist (1935) were collected in Corsica (Figs 39–42). No morphological differences were observed between these specimens and M. albocristata. Aside from the darker pilosity, the sculpture of the tegument, sterna, and genitalia is similar. The colour of the pilosity is highly variable in this subgenus and cannot be relied upon for species differentiation. Consequently, we propose considering M. lucidifrons as a new junior synonym for M. albocristata.

Figures 39–42.

Megachile albocristata Smith (Corsica, Mausoléo) 39 dorsal view 40 lateral view 41 head in front view 42 sternum 6. Black Scale bars: 5 mm. White Scale bars: 0.5 mm.

Current status. – Megachile (Chalicodoma) albocristata Smith, 1853.

Megachile sicula var. corsica Benoist, 1935

Figs 43–46

Megachile sicula f. corsica Benoist, 1935: p.103. Locality not indicated (Lectotype here designated, MNHN).

Type material examined. Lectotype: ♀, MNHN, Paris EY24221 // Bonif° [Bonifacio, Pérez’s handwritting] // Muséum Paris Ex. Coll. C. Ferton in J. Pérez // corsica [Benoist’s handwritting] // Lectotype // Megachile sícula f. corsica Benoist Lectotype des. Le Divelec 2021 // = Megachile sícula corsica Benoist, 1935 Le Divelec det. 2020; Paralectotype: ♂, MNHN, Paris EY24164 // Bonif° [Bonifacio, Pérez’s handwritting] // Ex. Coll. J. Pérez MNHN // Paralectotype // Megachile sícula f. corsica Benoist Paralectotype des. Le Divelec 2021 // = Megachile sícula corsica Benoist, 1935 Le Divelec det. 2020.

Figures 43–46.

Lectotype of Megachile sicula var. corsica Benoist 43 dorsal view 44 lateral view 45 head in front view 46 labels. Black Scale bars: 5 mm. White Scale bars: 1 mm.

Remark. Benoist (1935) did not specify a locality in the original description but the taxon’s name suggests it originates from Corsica. Benoist (1940) clarified the distribution of this taxon, indicating it is found in Bonifacio. Within Benoist’s reference collection, there is a female and a male from Bonifacio, both labelled as “corsica” by Benoist himself. These specimens were acquired by Benoist from Pérez’s collection, who, in turn, received them from C. Ferton. Other M. sicula corsica specimens in Benoist’s collection originate from Ajaccio or Vivario. Notably, three specimens of Benoist’s collection, borrowed by B. Tkalců in 1965, have not been returned according to the loans log. However, they could not be located in Tkalců’s collection (M. Schwarz pers. comm.). Consequently, we designate the female from Bonifacio, labelled by Benoist, as the lectotype. Despite partial damage to the left eye, left mandible, and left leg from pests, the female is in overall good condition and perfectly matches the original description. The associated male from Bonifacio is chosen as paralectotype.

Current status. Megachile (Chalicodoma) sícula corsica Benoist, 1935.

Megachile xanthopyga Pérez, 1895

Figs 47–50

Megachile xanthopyga Pérez, 1895: p.25, ♀♂. Locality not indicated (Lectotype designated by Praz and Bénon (2023), MNHN).

Type material examined. Lectotype: ♀, MNHN, Paris EY33616 // Sassari [Pérez’s handwritting] // dark blue circle [May] // Museum Paris Coll. J. Pérez 1915 // Lectotype ♀ Megachile xanthopyga Pérez, 1896 D. B. Baker des. 1991 // Megachile schmiedeknechti Costa det. G. v. d. Zanden 1995; Paralectotypes: ♀, MNHN, Paris EY33617 // Sassari [Pérez’s handwritting] // Museum Paris Coll. J. Pérez 1915 // Paralectotype ♀ Megachile xanthopyga Pér., 1896 D. B. Baker des. 1991 // Megachile schmiedeknechti Costa det. G. v. d. Zanden 1995; ♀, MNHN, Paris EY33618 // Sassari [Pérez’s handwritting] // Museum Paris Coll. J. Pérez 1915 // Paralectotype ♀ Megachile xanthopyga Pér., 1896 D. B. Baker des. 1991 // Megachile schmiedeknechti Costa det. G. v. d. Zanden 1995; ♀, MNHN, Paris EY23891 // Bonifacio 15/8 [15.VIII., Ferton’s label partly cut out] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Megachile xanthopyga J.P. Paralectotype des. Le Divelec 2021 // Megachile schmiedeknechti Costa det. G. v. d. Zanden 1995; ♀, MNHN, Paris EY23892 // Bonifacio 10/8 [10.VIII., Ferton’s label partly cut out] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Megachile xanthopyga J.P. Paralectotype des. Le Divelec 2021 // Megachile schmiedeknechti Costa det. G. v. d. Zanden 1995; ♂, MNHN, Paris EY23894 // Bonifacio 30/7 [30.VII., Ferton’s label partly cut out] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Megachile xanthopyga J.P. Le Divelec des. 2021.

Figures 47–50.

Lectotype of Megachile xanthopyga Pérez 47 dorsal view 48 lateral view 49 head in front view 50 labels. Scale bars: 1 mm.

Remark. No locality is specified in the original description. Pérez, in his original manuscript under M. xanthopyga (catalogue number 1732), mentions “Bonifacio, Sassari, ♀ mai-août, ♂ mai-juillet” (https://science.mnhn.fr/catalogue/ey-bib-perez1/). Praz and Bénon (2023) accepted and designated the Sardinian lectotype, along with two paralectotypes of M. xanthopyga labelled by D. Baker in 1991. Additionally, we have labelled as paralectotypes the three other specimens present under the xanthopyga head label in Pérez’s collection.

Current status. Megachile (Eutricharaea) argentata schmiedeknechti Costa, 1884.

Osmia corsica Ferton, 1901

Figs 51–54

Osmia corsica Ferton, 1901: p.61, ♀♂. Corsica: Bonifacio, Monte Renoso (Lectotype here designated, MNHN).

Type material examined. Lectotype: ♀, MNHN, Paris EY23992 // Vizzavona 19/7 99 [19.VII.1899] cote 1550 [altitude] Osmia corsica Fert ♀♂ [the mention “pallicornis Friese” crossed out] // Muséum Paris Coll. C. Ferton // Lectotype Le Divelec des. 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; Paralectotypes: ♀, MNHN, Paris EY23964 // Bonifacio 24/6 1900 [24.VI.1900] corsica Ferton ♀ // Muséum Paris Coll. C. Ferton // Paralectotype vdZ-1987 // Paralectotype valide Le Divelec 2021 // Hoplitis bihamata (Costa det. G. v. d. Zanden 1987 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♀, MNHN, Paris EY23968 // Osmia corsica Ferton ♀ Trinité (Bonifacio) 24/6 1900 [24.VI.1900] // Muséum Paris Coll. J. Vachal 1911 // Ex. Coll. C. Ferton // Paralectotype vdZ-1986 // Paralectotype valide Le Divelec 2021 // Hoplitis bihamata (Costa det. G. v. d. Zanden 1986 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♂, MNHN, Paris EY23972 // Vizzavona 19/7 1899 [19.VII.1899] cote 1550 [altitude] Osmia corsica ♂ // Muséum Paris Coll. C. Ferton // Paralectotype // Osmia corsica Ferton Paralectotype des. Le Divelec 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♂, MNHN, Paris EY23982 // Bonifacio (Trinité) 17/6 00 [17.VI.1900] ♂ corsica [the mention “pallicornis” crossed out] // Muséum Paris Coll. C. Ferton // Paralectotype // Osmia corsica Fert. Paralectotype des. Le Divelec 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♀, MNHN, Paris EY23994 // Fermant nid 19/7 99 [19.VII.1899] Vizzavona 1550 [altitude] Osmia corsica Fert. [the mention “pallicornis Friese” crossed out, four legs glued on the label] // Muséum Paris Coll. C. Ferton // Paralectotype // Osmia corsica Ferton Paralectotype des. Le Divelec 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♀, MNHN, Paris EY23999 // Muséum Paris Bonifacio (Trinité) 17-6-1900 [17.VI.1900] Coll. C. Ferton // Paralectotype // Osmia corsica Ferton Paralectotype des. Le Divelec 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♀, MNHN, Paris EY24001 // Muséum Paris Bonifacio (Trinité) 17-6-1900 [17.VI.1900] Coll. C. Ferton // Paralectotype // Osmia corsica Ferton Paralectotype des. Le Divelec 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021; ♀, MNHN, Paris EY24002 // Muséum Paris Bonifacio (Trinité) 17-6-1900 [17.VI.1900] Coll. C. Ferton // Paralectotype // Osmia corsica Ferton Paralectotype des. Le Divelec 2021 // Hoplitis bihamata (Costa, 1885) Le Divelec det. 2021.

Figures 51–54.

Lectotype of Osmia corsica Ferton 51 dorsal view 52 lateral view 53 head in front view 54 labels. Scale bars: 1 mm.

Remark. An unpublished lectotype and some paralectotypes were labelled by G. v. d. Zanden in the eighties. He randomly selected specimens of H. bihamata in Ferton’s collection. Therefore, his lectotype is not part of the type series and cannot be accepted. We have designated one female in good condition as lectotype and selected 8 specimens collected by C. Ferton in the granitic part of Bonifacio (e.g. La Trinité) and in the Monte Renoso as paralectotypes, in accordance with the original description information.

Current status. Hoplitis (Hoplitis) bihamata (Costa, 1885).

Osmia erythrogastra Ferton, 1905

Figs 55–58

Osmia erythrogastra Ferton, 1905: p.57, ♀♂. Corsica: Bonifacio, Santa–Manza (Lectotype here designated, MNHN).

Type material examined. Lectotype: ♂, Museum Paris EY0000002285 // Bonifacio 26/6 04 [26.VI.1904] erythrogastra Ferton ♂ // Museum Paris Corse Bonifacio C. Ferton 1902 // Lectotype Le Divelec des. // Os. Erythrogastra Ferton, 1905 Lectotype des. Le Divelec 2021 // Osmia erythrogastra Ferton, 1905 Le Divelec det. 2021; Paralectotypes: ♀, Museum Paris EY0000002284 // Bonifacio 26/6 04 [26.VI.1904] erythrogastra Ferton ♀ // Museum Paris Corse Bonifacio C. Ferton 1902 // Anthocopa erythrogastra (Ferton) ♀ det. G. v. d. Zanden 1987 // Paralectotype // Os. Erythrogastra Ferton, 1905 Paralectotype des. Le Divelec 2021 // Osmia erythrogastra Ferton, 1905 Le Divelec det. 2021; ♂, MNHN, Paris EY23930 // Bonifacio 26/6 04 [26.VI.1904] // Museum Paris Corse Bonifacio C. Ferton 1902 // Paralectotype // Os. Erythrogastra Ferton, 1905 Paralectotype des. Le Divelec 2021 // Osmia erythrogastra Ferton, 1905 Le Divelec det. 2021.

Figures 55–58.

Lectotype of Osmia erythrogastra Ferton 55 dorsal view 56 lateral view 57 head in front view 58 labels. Scale bars: 1 mm.

Remark. Ferton described this species based on four females and six males collected in the granitic area of Santa Manza on the June 26, 1904. Only two males and one female of this type series have been located. We designate as lectotype one male in good condition and bearing an original label.

Current status. Osmia (Erythrosmia) erythrogastra Ferton, 1905.

Osmia lanosa Pérez, 1879

Figs 59–62

Osmia lanosa Pérez, 1879: p.194, ♀. Algeria and Southern France (Lectotype here designated, MNHN).

Type material examined. Lectotype: ♀, MNHN, Paris EY33613 // M^tpel^r [Montpellier] // green disc [June] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Lectotype Le Divelec des. 2021 // Osmia lanosa Pérez Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; Paralectotypes: ♀, MNHN, Paris EY24179 // Bonifacio [cut out Ferton’s label] // green disc [June] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY24180 // Bonifacio [cut out Ferton’s label] // green disc [June] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33605 // Alg. [Algeria] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021; ♀, MNHN, Paris EY33606 // Marsr^le [Marseille] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33607 // Marsr^le [Marseille] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33608 // Marsr^le [Marseille] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33609 // Marsr^le [Marseille] // green disc [June] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33610 // Marsr^le [Marseille] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33611 // Marsr^le [Marseille] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021; ♀, MNHN, Paris EY33612 // Marsr.^le [Marseille] // green disc [June] // 1850 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lanosa Pérez Paralectotype des. Le Divelec des. 2021 // Hoplitis bisulca (Gerstäcker, 1869) Le Divelec det. 2021.

Figures 59–62.

Lectotype of Osmia lanosa Pérez 59 dorsal view 60 lateral view 61 head in front view 62 labels. Scale bars: 1 mm.

Remark. More information about the type localities can be found in Pérez’s manuscript catalogue (https://science.mnhn.fr/catalogue/ey-bib-perez1/): “Connue dans le midi oriental de la France et en Barbarie, en Corse, en Espagne. Montpellier. Marseille. Bonifacio. Barcelone. Vole en juin”. In accordance with this information, we have selected the lectotype and 11 paralectotypes among the specimens under the lanosa head label in Pérez’s collection. The female lectotype here designated is in good condition and matches with the current species concept of H. bisulca.

Current status. Hoplitis (Anthocopa) bisulca (Gerstäcker, 1869).

Osmia lineola Pérez, 1895

Figs 63–67

Osmia lineola Pérez, 1895: p.16, ♀♂. Locality not indicated (Lectotype here designated, MNHN).

Type material examined. Lectotype: ♂, MNHN, Paris EY24173 // S^taManza [Bonifacio, Santa–Manza] // green disc [June] // 1662 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Museum Paris Ex. Coll. C. Ferton // Lectotype Le Divelec des. 2021 // Osmia lineola J.P. Lectotype // Hoplitis praestans (Mor.) ♀ det. G. v. d. Zanden 1987 // Hoplitis praestans (Morawitz, 1893) Le Divelec det. 2021; Paralectotypes: ♀, MNHN, Paris EY24172 // Bonif.° [Bonifacio] // green disc [June] // 1662 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Museum Paris Ex. Coll. C. Ferton // Paralectotype // Osmia lineola J.P. Paralectotype des. Le Divelec 2021 // Hoplitis praestans Mor. ♀ det. G. v. d. Zanden 1986 // Hoplitis praestans (Morawitz, 1893) Le Divelec det. 2021; ♂, MNHN, Paris EY33604 // Mars^le [Marseille] // 439 // 1662 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lineola J.P. Paralectotype des. Le Divelec 2021 // Hoplitis praestans Mor. ♂ det. G. v. d. Zanden 1986 // Hoplitis praestans (Morawitz, 1893) Le Divelec det. 2021; ♀, MNHN, Paris EY33614 // Tibériade // modesta Ab. Tib. N°467 // 1662 [Pérez’s catalogue number] // Museum Paris Coll. J. Pérez 1915 // Paralectotype // Osmia lineola J.P. Paralectotype des. Le Divelec 2021 // Hoplitis praestans (Mor.) ♀ det. G. v. d. Zanden 1986 // Hoplitis praestans (Morawitz, 1893) Le Divelec det. 2021.

Figures 63–67.

Lectotype of Osmia lineola Pérez 63 dorsal view 64 lateral view 65 head in front view 66 gaster in ventral view 67 labels. Scale bars: 1 mm.

Remark. Additional details regarding the type localities can be accessed in Pérez’s manuscript catalogue (https://science.mnhn.fr/catalogue/ey-bib-perez1/): “♀ Tibériade (Abeille de Perrin). ♀ Bonifacio, Juin (Ferton), butinant sur le Centaurea suaveolens (Ferton). ♂ Marseille et Corse (Santa–Manza)”. Within Pérez’s collection, only four specimens are located under the “lineola” head label, with two originating from Corsica (Ferton leg.). These specimens align with both the manuscript information and the original description. The male from Corsica in the best condition is designated as the lectotype, while the three remaining individuals are labelled as paralectotypes.

Current status. Hoplitis (Alcidamea) praestans (Morawitz, 1893).

Osmia nasoproducta Ferton, 1909

Figs 68–71

Osmia nasoproducta Ferton, 1909b: p.406, ♀. Corsica: Bonifacio (Holotype, MNHN).

Type material examined. Holotype: ♀, MNHN, Paris EY24008 // Osmia ♀ Fert. [the mention “bonifaciensis” crossed out] Bonifacio 24/06 02 [24.VI.1902], page 24 nasoproducta // Museum Paris Corse Bonifacio C. Ferton 1902 // Holotype.

Figures 68–71.

Lectotype of Osmia nasoproducta Ferton 68 dorsal view 69 lateral view 70 head in front view 71 labels. Scale bars: 1 mm.

Current status. Osmia (Helicosmia) nasoproducta (Ferton, 1909).

Acknowledgements

We are grateful to Andreas Müller, Gérard Le Goff, Eric Dufrêne and Paul Vignac for sharing their data with us. Many thanks to Erwin Scheuchl and Maurizio Cornalba for our helpful exchanges. Thanks to Martin Schwarz for his help with the material preserved in Linz collections and to Rémi Santerre for his help with Mavromoustakis collection. We are grateful to Christophe Praz for improving this manuscript. Quentin Rome (Patrinat) kindly supported us with the uploading of our data on the INPN data portal. Special thanks to Claire Villemant, Agnièle Touret-Alby, Laurent Albenga, and Antoine Mantilleri for access to MNHN collections and to the photography setup of the department of terrestrial Arthropod’s collections management (MNHN, Paris). Collecting specimens carried out in 2016–2017 was part of a program funded by the Labex BCDiv of MNHN, with the help of OCIC, OEC and the Conservatoire du Littoral de Corse du Sud. Thanks to Claire Villemant, Colin Fontaine, Marianne Elias, Benjamin Yguel, Marie-Cécile Ruiz and Viviane Sorba for joining or helping with the collecting in the campaigns of 2016 and 2017. The “Our Planet Revisited–Corsica 2019–2022” expedition was organized by MNHN (Paris) and funded by the Office Français de la Biodiversité (OFB) and the Collectivité de Corse (CdC). Many thanks to Julien Touroult, François Dusoulier and Jean Ichter for organizing these wonderful expeditions. Thanks to Claire Villemant, Bernardo F. Santos and Quentin Rome for collecting Megachilid bees during these expeditions. The present catalogue was realized within the “Inventaire national du patrimoine naturel (inpn.mnhn.fr)” project with funding from PatriNat (OFB, CNRS, MNHN).

References

Aguib S, Louadi K, Schwarz M (2010) Le genre Stelis PANZER 1806 (Hymenoptera, Apoidea, Megachilidae) de l’Algérien avec une espèce nouvelle pour la faune de ce pays. Entomofauna 35(26): 553–572.

Alfken JD (1923) Zur Kenntnis einiger Arten der Megachile argentata-Gruppe. Acta Musaei Dzieduszckiani 9: 4–8.

Baker DB (1999) On new stelidine bees from S. W. Asia and N. W. Africa, with a list of the Old-World taxa assigned to the genus Stelis Panzer, 1806 (Hymenoptera, Apoidea, Megachilidae). Deutsche Entomologische Zeitschrift 46(2): 231–242. https://doi.org/10.1002/mmnd.4800460209

Baldock D, Wood TJ, Cross I, Smit J (2018) The bees of Portugal (Hymenoptera: Apoidea: Anthophila). Entomofauna Supplement 22: 1–164.

Benoist R (1929) Les Heriades de la faune française (Hym. Apidae). Annales de la Société Entomologique de France 98: 131–141. https://doi.org/10.1080/21686351.1929.12280329

Benoist R (1931) Les Osmies de la faune française (Hymenopt. Apidae). Annales de la Société Entomologique de France 100: 23–60. https://doi.org/10.1080/21686351.1931.12280120

Benoist R (1935) Remarques sur quelques espèces du genre Megachile [Hymen. Apidae]. Annales de la Société Entomologique de France 104: 98–108.

Benoist R (1940) Remarques sur quelques espèces de Mégachiles principalement de la faune française. Annales de la Société Entomologique de France 109: 41–88. https://doi.org/10.1080/21686351.1940.12279433

Blüthgen P (1930) Stelis phaeoptera K. subsp. nov. franconica (Hym. Apidae). Archiv für Insektenkunde des Oberrheingebietes u. der Angrenzenden Länder 2: 277–278.

Bogusch P (2023) European cuckoo bees of the tribe Dioxyini (Hymenoptera, Megachilidae): distribution, annotated checklist and identification key. Journal of Hymenoptera Research 96: 599–628. https://doi.org/10.3897/jhr.96.104957

Brullé GA (1839) Insectes. In: Barker-Webb P, Berthelot S (Eds) Histoire Naturelle des Îles Canaries (Tome deux). Béthune, Paris, 54–95.

Bürgis H (1995) Leucopsis gigas (Chalcidoidea: Leucopsidae) als Parasit der Mörtelbiene Megachile sicula (Apoidea: Megachilidae). bembiX 5: 27–32.

Canovai R, Giannotti P, Giannetti S, Loni A, Raspi A, Santini L, Dellacasa M, Generani M, Pagliano G, Strumia F, Scaramozzino PL, Zuffi M, Baldaccini NE, Puglisi L, Battesti MJ, Brocard E (2000) Biodiversita: Compilazione delle specie dell’Entomofauna e dei piccoli vertebrati della Corsica e della Toscana Maritima. Progretto interreg II. Toscana-Corsica, 1997–1999. L’attività scientifica delle Università di Pisa e Corte: 75–86.

Chao A (1984) Nonparametric estimation of the number of classes in a population. Scandinavian Journal of statistics 11(4): 265–270.

Chao A, Lee SM (1992) Estimating the number of classes via sample coverage. Journal of the American statistical Association 87(417): 210–217. https://doi.org/10.1080/01621459.1992.10475194

Colwell RK, Coddington JA (1994) Estimating terrestrial biodiversity through extrapolation. Philosophical Transactions: Biological Sciences 345: 101–118. https://doi.org/10.1098/rstb.1994.0091

Comba M (2019) Hymenoptera: Apoidea: Antophila of Italy. Biogeographic Checklist of Italian Wild Bees with Notes on Taxonomy, Biology, and Distribution. https://digilander.libero.it/mario.comba [accessed on 12.10.2022]

Cornuel-Willermoz A, Andrei-Ruiz MC (2021) Plan Territorial d’Actions en faveur des pollinisateurs sauvages et de l’abeille mellifère –Corse : 2021–2030. Office de l’Environnement de la Corse, Corte, 43 pp.

Ducke A (1900) Die Bienengattung Osmia Panz. als Ergänzung zu Schmiedeknecht’s “Apidae europaeae” Vol. II in ihren palaearctischen Arten monographisch bearbeitet. Bericht des Naturwissenschaftlich-Medizinischen Vereins in Innsbruck 25: 1–323.

Dorchin A, Praz CJ (2018) Taxonomic revision of the Western Palaearctic bees of the subgenus Pseudomegachile (Hymenoptera, Apiformes, Megachilidae, Megachile). Zootaxa 4524(3): 251–207. https://doi.org/10.11646/zootaxa.4524.3.1

DREAL (2017) Le Profil Environnemental de la Corse, édition 2016. DREAL Corse, Corte, 236 pp.

Dufrêne E, Genoud D, Bourlet P (2016) Sur la distribution en France de Lithurgus cornutus Fabricius 1827 (Hymenoptera – Megachilidae -Lithurgini): Apport de données récentes. OSMIA 6: 16–21. https://doi.org/10.47446/OSMIA6.5

Ebmer AW (1997) Hymenopterologische Notizen aus Österreich – 7 (Insecta: Hymenoptera: Apoidea). Linzer Biologische Beiträge 29: 45–62.

Ferton C (1897) Nouvelles observations sur l’instinct des hyménoptères gastrilégides de France et de Corsica. Actes de la Société Linnéenne de Bordeaux 52: 37–50.

Ferton C (1899) Sur les mœurs du Chrysis dichroa Dahlbom [Hymén.]. Bulletin de la Société Entomologique de France 1899: 70–73. https://doi.org/10.3406/bsef.1899.22301

Ferton C (1901a) Description de l’Osmia corsica, n. sp. et observations sur la faune Corsica (Hymén.). Bulletin de la Société Entomologique de France 1901: 61–66. https://doi.org/10.3406/bsef.1901.22776

Ferton C (1901b) Notes détachées sur l’instinct des Hyménoptères mellifères et ravisseurs avec la description de quelques espèces. Annales de la Société entomologique de France 70: 83–148.

Ferton C (1905) Notes détachées sur l’instinct des hyménoptères mellifères et ravisseurs (3^e série) avec la description de quelques espèces. Annales de la Société Entomologique de France 74: 56–104. [pl. 3–4.]

Ferton C (1909a) Notes détachées sur l’instinct des hyménoptères mellifères et ravisseurs (4^e série) avec la description de quelques espèces. Annales de la Société Entomologique de France 77: 535–586.

Ferton C (1909b) Notes détachées sur l’instinct des hyménoptères mellifères et ravisseurs (5^e série) avec la description d’une espèce nouvelle. Annales de la Société Entomologique de France 78: 401–422.

Ferton C (1911) Notes détachées sur l’instinct des hyménoptères mellifères et ravisseurs (7^e série) avec la description de quatre espèces nouvelles. Annales de la Société Entomologique de France 80: 351–412.

Ferton C (1920) Notes détachées sur l’instinct des hyménoptères mellifères et ravisseurs (9ème série) avec la description de deux espèces nouvelles. Annales de la Société Entomologique de France 89: 329–375.

Ferton C (1923) La vie des Abeilles et des Guêpes. Chiron, Paris, 376 pp.

Floris I, Satta A, Lentini A (2000) Monitoring of insect pollinators in two different agricultural landscapes (Sardinia, Italy). Insect Social Life 3: 115–118.

Friese H (1895) Die Bienen Europa‘s (Apidae europaeae) Theil I. Schmarotzerbienen. Friedländer und Sohn, Berlin, 218 pp.

Friese H (1899) Die Bienen Europa‘s (Apidae europaeae) nach ihren Gattungen, Arten und Varietäten auf vergleichend morphologisch-biologischer Grundlage. Theil V: Solitäre Apiden: Genus Lithurgus, Genus Megachile (Chalicodoma). C. Lampe, Innsbruck und Imst, 228 pp.

Friese H (1911) Apidae I. Megachilinae. Das Tierreich 28: 1–440.

Ghisbain G, Rosa P, Bogusch P, Flaminio S, Le Divelec R, Dorchin A, Kasparek M, Kuhlmann M, Litman J, Mignot M, Müller A, Praz C, Radchenko VG, Rasmont P, Risch S, Roberts SPM, Smit J, Wood TJ, Michez D, reverté S (2023) The new annotated checklist of the wild bees of Europe (Hymenoptera: Anthophila). Zootaxa 5327(1): 001–147. https://doi.org/10.11646/zootaxa.5327.1.1

Gonzalez VH, Engel MS, Hinojosa-Díaz IA (2010) A new species of Megachile from Pakistan, with taxonomic notes on the subgenus Eutricharaea (Hymenoptera: Megachilidae). Journal of the Kansas Entomological Society 83: 58–67. https://doi.org/10.2317/JKES0905.16.1

Kasparek M (2015) The cuckoo bees of the genus Stelis Panzer, 1906 in Europe, North Africa and in the Middle East. A review and identification guide. Entomofauna Supplement 18: 1–144.

Kasparek M (2022) The Resin and Wool Carder Bees (Anthidiini) of Europe. Chimaira, Frankfurt am Main, 290 pp.

Kasparek M, Lhomme P (2019) Revision of the taxonomic status of Rhodanthidium stictitum ordonezi (Dusmet, 1915) an anthidiine bee endemic to Morocco (Apoidea: Anthidiini). Turkish Journal of Zoology 43: 43–51. https://doi.org/10.3906/zoo-1809-22

Kusdas K (1974) Beitrag zur Kenntnis der Insektenfauna von Korsika. Zeitschrift der Arbeitsgemeinschaft Österreichischer Entomologen 24: 153–166.

Le Divelec R, Dufrêne E (2020) Sur l’identité du mâle de Coelioxys coturnix PÉREZ 1884 (Hymenoptera – Megachilidae). OSMIA 8: 5–10. https://doi.org/10.47446/OSMIA8.1

Le Goff G (2004) Hoplitis (Hoplitis) manicata Morice, espèce nouvelle pour la faune des France et la peu commune Megachile (Pseudomegachile) ericetorum melaleuca Van der Zanden en Corse (Apoidea; Megachilidae; Megachilinae; Osmiini; Megachilini). L’Entomologiste 60: 9–12.

Le Goff G (2010) Note sur la nidification dans le sol de quatre Hoplitis Klug du sous-genre Annosmia Warncke (Apoidea, Megachilidae, Osmiini). OSMIA 4: 5–10. https://doi.org/10.47446/OSMIA4.2

Le Goff G (2016) Description du mâle d’Osmia (Helicosmia) nasoproducta Ferton 1909 (Apoidea – Megachilidae – Osmiini). OSMIA 6: 4–5. https://doi.org/10.47446/OSMIA6.1

Liongo li Enkulu M (1988) Les Mégachiles (Hymenoptera, Apoidea) d’Europe et d’Afrique. Une étude écologique et agronomique. PhD thesis. Faculté Universitaire des Sciences Agronomiques, Gembloux, 247 pp.

Litman JR, Fateryga AV, Griswold TL, Aubert M, Proshchalykin MY, Le Divelec R, Burrows S, Praz CJ (2021) Paraphyly and low levels of genetic divergence in morphologically distinct taxa: revision of the Pseudoanthidium scapulare complex of carder bees (Apoidea: Megachilidae: Anthidiini). Zoological Journal of the Linnean Society 195: 1287–1337. https://doi.org/10.1093/zoolinnean/zlab062

Marchal P, Chardonnet G (2001) Sortie entomologique en Corse en mai 1998, liste des insectes récoltés, 2e note. Bulletin Mensuel de la Société Linnéenne de Lyon 70(10): 201–204. https://doi.org/10.3406/linly.2001.11403

Meunier JY, Geslin B, Issertes M, Mahé G, Vyghen F, Labrique H, Dutour Y, Poncet V, Migliore J, Neve G (2023) Apoidea of the collections of Lyon, Aix-en-Provence, Marseille and Toulon Museums of Natural History (France). Biodiversity Data Journal 11: e99650. https://doi.org/10.3897/BDJ.11.e99650

Morawitz F (1871) Neue suedeuropaeische Bienen. Horae Societatis Entomologicae Rossicae 8: 201–231.

Müller A (2018) Palaearctic Osmia bees of the subgenus Hoplosmia (Megachilidae, Osmiini): biology, taxonomy and key to species. Zootaxa 4415(2): 297–329. https://doi.org/10.11646/zootaxa.4415.2.4

Müller A (2020) Palaearctic Osmia bees of the subgenera Hemiosmia, Tergosmia and Erythrosmia (Megachilidae, Osmiini): biology, taxonomy and key to species. Zootaxa 4778(2): 201–236. https://doi.org/10.11646/zootaxa.4778.2.1

Müller A (2022a) Palaearctic Osmiine Bees. http://blogs.ethz.ch/osmiini [accessed on 09.09.2022]

Müller A (2022b) Palaearctic Osmia bees of the subgenera Allosmia and Neosmia (Megachilidae, Osmiini): biology, taxonomy and key to species. Zootaxa 5188(3): 201–232. https://doi.org/10.11646/zootaxa.5188.3.1

Nadig A, Nadig A (1933) Beitrag zur Kenntnis der Hymenopterenfauna von Marokko und Westalgerien – Erster Teil: Apidae, Sphegidae, Vespidae. Jahresbericht der Naturforschenden Gesellschaft Graubündens 71: 37–107.

Nadig A, Nadig A (1934) Beitrag zur Kenntnis der Orthopteren- und Hymenopterenfauna von Sardinien und Korsika. Jahresbericht der Naturforschenden Gesellschaft Graubündens 72: 3–40.

Nobile V, Catania R, Niolu P, Pusceddu M, Satta A, Floris I, Flaminio S, Bella S, Quaranta M (2021) Twenty New Records of Bees (Hymenoptera, Apoidea) for Sardinia (Italy). Insects 12(7). https://doi.org/10.3390/insects12070627

Ornosa C, Ortiz-Sánchez FJ, Torres F (2007) Catalogo de los Megachilidae del Mediterraneo occidental (Hymenoptera, Apoidea). II. Lithugini y Megachilini. Graellsia 63: 111–134. https://doi.org/10.3989/graellsia.2007.v63.i1.85

Orr MC, Hughes AC, Chesters D, Pickering J, Zhu C-D, Ascher JS (2021) Global Patterns and Drivers of Bee Distribution. Current Biology 31: 451–458. https://doi.org/10.1016/j.cub.2020.10.053

Pagliano G (1994) Catalogo degli Imenotteri italiani. IV. Apoidea: Colletidae, Andrenidae, Megachilidae, Anthophoridae, Apidae). Memorie della Società Entomologica Italiana 72: 331–467.

Peters DS (1977) Systematik und Zoogeographie der west-paläarktischen Arten von Osmia Panzer, 1806 s.str., Monosmia Tkalců, 1974 und Orientosmia n. subgen. (Insecta: Hymenoptera: Megachilidae). Senckenbergiana Biologica 58: 287–346.

Pérez J (1879) Contribution à la faune des apiaires de France. Actes de la Société Linnéenne de Bordeaux 33: 119–229.

Pérez J (1895) Espèces Nouvelles de Mellifères de Barbarie. (Diagnoses préliminaires). Gounouilhou, Bordeaux, 64 pp.

Praz CJ (2017) Subgeneric classification and biology of the leafcutter and dauber bees (genus Megachile Latreille) of the western Palearctic (Hymenoptera, Apoidea, Megachilidae). Journal of Hymenoptera Research 55: 1–54. https://doi.org/10.3897/jhr.55.11255

Praz CJ, Bénon D (2023) Revision of the leachella group of Megachile subgenus Eutricharaea in the Western Palaearctic (Hymenoptera, Apoidea, Megachilidae): A renewed plea for DNA barcoding type material. Journal of Hymenoptera Research 95: 143–198. https://doi.org/10.3897/jhr.95.96796

Přidal A (2004) Checklist of the bees in the Czech Republic and Slovakia with comments on their distribution and taxonomy (Insecta: Hymenoptera: Apoidea). Acta Universitatis Agriculturae et Silviculturae Mendelianae Brunensis 52(1): 29–66. https://doi.org/10.11118/actaun200452010029

Quaranta M, Ambroselli S, Barro P, Bella S, Carini A, Celli G, Cogoi P, Comba L, Comoli R, Felicioli A, Floris I, Intoppa F, Longo S, Maini S, Manino A, Mazzeo G, Medrzycki P, Nardi E, Niccolini L, Palmieri N, Patetta A, Piatti C, Piazza MG, Pinzauti M, Porporato M, Porrini C, Ricciardelli D’Albore G, Romagnoli F, Ruiu L, Satta A, Zandigiacomo P (2004) Wild bees in agroecosystems and semi-natural landscapes. 1997–2000 collection period in Italy. Bulletin of Insectology 57(1): 11–61.

Radoszkowsi O (1887) Sur quelques Osmia russes. Horae Societatis Entomologicae Rossicae 21: 275–293.

Rasmont P, Ebmer P, Banaszak J, Zanden G (1995) Hymenoptera Apoidea Gallica. Liste taxonomique des abeilles de France, de Belgique, de Suisse et du Grand-Duché de Luxembourg. Bulletin de la Société Entomologique de France 100: 1–98.

Rasmont P, Adamski A (1995) Les bourdons de la Corse (Hymenoptera, Apoidea, Bombinae). Notes Fauniques de Gembloux 31: 3–87. https://doi.org/10.1080/21686351.1995.12277716

Rasmont P, Genoud D, Gadoum S, Aubert M, Dufrêne E, Le Goff G, Mahé G, Michez D, Pauly A (2017) Hymenoptera Apoidea Gallica: liste des abeilles sauvages de Belgique, France, Luxembourg et Suisse. http://www.atlashymenoptera.net/biblio/01500/414_Rasmont_et_al_2017_Hymenoptera_Apoidea_Gallica_2017_02_16.pdf [accessed on 27.06.2023]

Rebmann O (1968) 3. Beitrag zur Kenntnis der Gattung Megachile Latr. (Hym. Apidae): subgenus Eutricharaea und seine bisher bekanntgewordenen Arten. Deutsche Entomologische Zeitschrift 15: 21–48. https://doi.org/10.1002/mmnd.19680150105

Ruzzier E, Menchetti M, Bortolotti L, Selis M, Monterastelli E, Forbicioni L (2020) Updated distribution of the invasive Megachile sculpturalis (Hymenoptera: Megachilidae) in Italy and its first record on a Mediterranean island. Biodiversity Data Journal 8: e57783. https://doi.org/10.3897/BDJ.8.e57783

Sabiani T (2004) Encyclopaedia Corsicae (Vol. 1): Géologie – La mer – la terre. Edition Dumane, Corse, 812 pp.

Scheuchl E, Willner W (2016) Taschenlexikon der Wildbienen Mitteleuropas. Quelle and Meyer, Wiebelsheim, 917 pp.

Schmiedeknecht O (1885–1886) Genus Osmia Panz. Apidae Europaeae (Die Bienen Europa’s) per Genera, Species et Varietates. Vol. 2. Friedländer, Berlin, 205 pp.

Schwarz M, Gusenleitner F (2011) Beitrag zur Kenntnis der Gattung Megachile (Hymenoptera, Apoidea, Megachilinae). Entomofauna 32(15): 249–260.

Soltani GG, Bénon D, Alvarez N, Praz CJ (2017) When different contact zones tell different stories: putative ring species in the Megachile concinna species complex (Hymenoptera: Megachilidae). Biological Journal of the Linnean Society 121: 815–822. https://doi.org/10.1093/biolinnean/blx023

Stanek E (1969) Neue oder wenig bekannte Osmia-Arten aus dem Mittelmeergebiet (Hymenoptera, Apoidea, Megachilidae). Nachrichten des Naturwissenschaftlichen Museums der Stadt Aschaffenburg 78: 1–40. [pl. 1.]

Stöckl P (2000) Synopsis der Megachilinae Nord -und Südtirols (Österreich, Italien) (Hymenoptera: Apidae). Berichte des Naturwissenschaftlich-medizinischen Vereins in Innsbruck 87: 273–306.

Terzo M, Iserbyt S, Rasmont P (2007) Révision des Xylocopinae (Hymenoptera : Apidae) de France et de Belgique. Annales de la Société Entomologique de France 43: 445–491. https://doi.org/10.1080/00379271.2007.10697537

Tkalců B (1969) Beiträge zur Kenntnis der Fauna Afghanistans. (Sammelergebnisse von O. Jakeš 1963–64, D. Povolný 1965, D. Povolný and Fr. Tenora 1966, J. Šimek, 1965–66, D. Povolný, J. Gaisler, Z. Šebek and Fr. Tenora 1967). Osmiini, Megachilidae, Apoidea, Hym. Casopis Moravského Musea Acta Musei Moraviae 54: 327–346.

Tkalců B (1970) Typenrevision der von J.C. Fabricius beschriebenen paläarktischen Arten der Tribus Osmiini (Hymenoptera, Apoidea, Megachilidae). Annotationes Zoologicae et Botanicae 62: 1–15.

Tkalců B (1971) Zur Identität zweier Osmia-Arten (Hymenoptera, Apoidea, Megachilidae). Acta Entomologica Bohemoslovaca 68: 222–230.

Tkalců B (1975a) Revision der Europäischen Osmia (Chalcosmia)-Arten der fulviventris-Gruppe (Hymenoptera, Apoidea, Megachilidae). Věstník Československé Společnosti Zoologické 39: 297–317.

Tkalců B (1975b) Sammelergebnisse der von RNDr. A. Hoffer geleiteten Algerien-Expeditionen in den Jahren 1971 und 1972 (Hymenoptera: Apoidea) 1. Teil: Megachilidae. Zbornik Slovenského Národného Mùzea 21: 165–190.

Tkalců B (1977) Taxonomische Notizen zu einigen paläarktischen Osmiini-Arten (Hymenoptera, Apoidea). Acta Musei Moraviae 62: 87–98.

Tkalců B (1993) Neue Taxa der Bienen von den Kanarischen Inseln. Mit Bemerkungen zu einigen bereits bekannten Arten (Insecta, Hymenoptera, Apoidea). Veröffentlichungen des Übersee-Museum Bremen 12: 791–858.

Ungricht S, Müller A, Dorn S (2008) A taxonomic catalogue of the Palaearctic bees of the tribe Osmiini (Hymenoptera: Apoidea: Megachilidae). Zootaxa 1865: 1–253. https://doi.org/10.11646/zootaxa.1865.1.1

Varnava AI, Roberts SP, Michez D, Ascher JS, Petanidou T, Dimitriou S, Devalez J, Pittara M, Stavrinides MC (2020) The wild bees (Hymenoptera, Apoidea) of the island of Cyprus. ZooKeys 924: 1–114. https://doi.org/10.3897/zookeys.924.38328

Vereecken NJ, Barbier E (2009) Premières données sur la présence de l’abeille asiatique Megachile (Callomegachile) sculpturalis Smith (Hymenoptera, Megachilidae) en Europe. OSMIA 3: 4–6. https://doi.org/10.47446/OSMIA3.3

Warncke K (1981) Die Bienen des Kagenfurter Beckens. Carinthia II 171(91): 275–348.

Warncke K (1988) Isolierte Bienenvorkommen auf dem Olymp in Griechenland (Hymenoptera, Apidae). Linzer Biologische Beiträge 20: 83–117.

Warncke K (1991a) Die Bienengattung Osmia Panzer, 1806, ihre Systematik in der Westpaläarktis und ihre Verbreitung in der Türkei. 9. Die Untergattung Annosmia subg. n. Linzer Biologische Beiträge 23: 307–336.

Warncke K (1991b) Die Bienengattung Osmia Panzer, 1806, ihre Systematik in der Westpaläarktis und ihre Verbreitung in der Türkei. 10. Die Untergattung Alcidamea Cress. Linzer Biologische Beiträge 23: 701–751.

Warncke K (1992a) Die westpaläarktischen Arten der Bienengattung Coelioxys Lep. (Hymenoptera, Apoidea). Bericht der Naturforschenden Gesellschaft Augsburg 53: 31–77.

Warncke K (1992b) Die westpaläarktischen Arten der Bienengattung Stelis Panzer, 1806 (Hymenoptera, Apidae, Megachilinae). Entomofauna 13(22): 341–376.

Warncke K (1992c) Die westmediterranen Arten der Bienen Osmia subg. Hoplitis Klug, 1807. Linzer Biologische Beiträge 24: 103–121.

Warncke K (1992d) Die Bienengattung Osmia Panzer, 1806 ihre Systematik in der Westpaläarktis und ihre Verbreitung in der Türkei. 11. Die Untergattung Pyrosmia Tkalců, 1975. Linzer Biologische Beiträge 24: 893–921.

Willmer P (2011) Pollination and Floral Ecology. Princeton University Press, Princeton and Oxford, 778 pp. https://doi.org/10.23943/princeton/9780691128610.001.0001

Zanden G (1984) Neue paläarktische Taxa der Familie Megachilidae (Hymenoptera, Apoidea, Megachilidae). Reichenbachia 22: 175–191.

Zanden G (1989) Neue oder wenig bekannte Arten und Unterarten der palaearktischen Megachiliden (Insecta, Hymenoptera, Apoidea: Megachilidae). Entomologische Abhandlungen Staatliches Museum für Tierkunde Dresden 53: 71–86.

Zanden G (1991) Systematik und Verbreitung der paläarktischen Arten der Untergattung Caerulosmia van der Zanden, 1989 (Hymenoptera, Apoidea, Megachilidae). Linzer Biologische Beiträge 23: 37–78.

﻿Abstract

Keywords

﻿Introduction

﻿Material and methods

﻿Study of the collections housed at MNHN

﻿Recent sampling

﻿Additional data gathering

﻿Conventions of the checklist

﻿Results and discussion

﻿Checklist of the Corsican Megachilidae

﻿Anthidiini Ashmead, 1899

﻿Dioxyini Cockerell, 1902

﻿Lithurgini Newman, 1834

﻿Megachilini Latreille, 1802

﻿Osmiini Newman, 1834

﻿Dubious records

﻿Erroneous records

﻿Types of Megachilid bees described from Corsica and housed at MNHN

﻿Acknowledgements

References

Abstract

Introduction

Material and methods

Study of the collections housed at MNHN

Recent sampling

Additional data gathering

Conventions of the checklist

Results and discussion

Checklist of the Corsican Megachilidae

Anthidiini Ashmead, 1899

Dioxyini Cockerell, 1902

Lithurgini Newman, 1834

Megachilini Latreille, 1802

Osmiini Newman, 1834

Dubious records

Erroneous records

Types of Megachilid bees described from Corsica and housed at MNHN

Acknowledgements