Diagnosis.

Cryptopimpla can be distinguished from all other banchine genera by the following combination of characters: lower end of occipital carina joining hypostomal carina above base of mandible; apical 0.3–0.4 of flagellum tapered towards apex; epomia absent; lower half of mesopleuron weakly convex or flat; posterior transverse carina of propodeum complete, or sometimes partly weak; pleural carina present, sometimes weak; areolet present, its petiole short or absent; 2m-cu with two bullae or a single wide bulla; hind wing with distal abscissa of Cu1 much closer to 1A than to M; first metasomal tergite with glymma, its spiracle before the middle, its dorsal profile strongly convex before the spiracle; median dorsal carina on first metasomal tergite absent; ovipositor shorter than hind tibia, sometimes upcurved.

Remarks.

Keys to species of Cryptopimpla have been published for the following regions: Oriental (Chandra and Gupta 1977), Russian Far East (Kuslitzky 2007), China (Sheng and Zheng 2005; Sheng 2011), Japan (Momoi 1970), Europe (Aubert 1978; Schwarz 2003 treating only species with black metasomal tergites) and the Nearctic (Townes and Townes 1978). All the described species with black metasomal tergites, except for the Afrotropical Cryptopimpla rubrithorax Morley, 1916, are included in these keys. Although the species collected from Sulawesi possesses a black metasoma, it could not be assigned to any described species by the keys or by comparison with the description of Cryptopimpla rubrithorax, and most described species are very unlikely to be found in Sulawesi. None of the described species would be expected to match this species from Sulawesi, considering differences in climate and geography, because most described species are distributed in the temperate or subtropical regions, especially in the northern hemisphere.

Diagnosis.

Cryptopimpla sulawesiensis can be distinguished from all other known species by the following combination of characters: frons without horn or tubercle; malar space 0.9–1.0 times as long as basal width of mandible; antenna uniformly cylindrical, not moniliform, apical part slender; antenna with 40–44 flagellomeres; palpi whitish yellow; posterior transverse carina and pleural carina on propodeum weak, not distinctly defined; apical part of tarsal claws bent at right angle; hind femur short and robust; areolet pentagonal, receiving 2m-cu medially; metasomal tergite I 1.7–2.0 (female) or 2.0–2.2 (male) times as long as maximum width; base of metasomal tergite I marked yellow to brown; ovipositor sheath 0.5–0.6 times as long as hind tibia; mesosoma and metasoma in female largely black, without reddish parts or apical yellow bands on metasomal tergites.

Female (Figs 1, 3, 5, 6). Body 8.5–9.5 (HT: 8.6) mm long, fore wing 6.2–7.0 (HT: 6.4) mm long. Head polished, punctate, 0.6 times as long as wide in dorsal view; clypeus 0.5 times as long as wide, strongly convex in lateral view, punctures sparser than on face; face 0.8–1.0 (HT: 1.0) times as long as wide, median part with longitudinal ridge (Fig. 3); punctures on frons partly tend towards striation; malar space 1.0 times as long as basal width of mandible; basal portion of mandible convex; vertex and gena minutely and finely punctate; OOL/OD 1.0–1.1 (HT: 1.0); POL/OD 0.9–1.0 (HT: 1.1); antenna with 40–44 (HT: 40) flagellomeres; flagellomere I 3.0–3.8 (HT: 3.0) times as long as apical depth and 1.4–1.6 (HT: 1.6) times as long as flagellomere II.

Mesosoma matt, densely punctate (Figs 5, 6), distance between punctures shorter than puncture diameter, mesosoma 2.3–2.4 (HT: 2.3) times as long as width between tegulae; scutellum slightly more sparsely punctate than mesoscutum; lower half of mesopleuron slightly convex; upper end of epicnemial carina reaching lower 1/4 of pronotum; mesopleural suture straight, foveolate (Fig. 5); episternal scrobe small, slightly larger than propodeal spiracle (Fig. 5); propodeum punctate-reticulate with median portion reticulate rugose on anterior half, rugose on posterior half; posterior transverse carina present, interrupted by rugosity, pleural carina weak and other carinae absent (Fig. 6); border between propodeum and metapleuron indicated by furrow (Fig. 5); area posteroexterna short, about 1/4 total length of propodeum (Fig. 6); propodeal spiracle round (Fig. 5). Legs: hind femur 5.2–5.8 (HT: 5.7) times as long as deep; hind tibia 8.0 times as long as wide; hind tarsomere I 2.4–2.6 (HT: 2.6) times as long as hind tarsomere II and 2.0–2.1 (HT: 2.1) times as long as longer hind tibial spur, respectively; tarsal claws 1.2 times as long as arolium, pectinate, their apical part bent at right angle. Wings (Fig. 7): fore wing with Cu-a distad Rs+M by 0.5–0.8 (HT: 0.5) times length of Cu-a, with pentagonal areolet receiving 2m-cu medially; hind wing with distal abscissa of Cu1 much closer to 1A than to M, basal abscissa of Cu1 2.0 times as long as cu-a.

Metasoma polished, punctate; tergite I 1.7–2.0 (HT: 1.7) times as long as maximum width, 1.3–1.5 (HT: 1.3) times as long as tergite II, densely punctate except for smooth basal part, narrow median longitudinal smooth line and posterior margin, punctures partly tend towards striation, each side of basal part weakly convex; tergites II and III punctate, punctures along anterior and posterior margins sparse and shallow, or partly absent; tergite IV and following tergites almost smooth; ovipositor 1.6–2.2 (HT: 1.6) mm, weakly up-curved, sheath 0.5–0.6 (HT: 0.6) times as long as hind tibia.

Coloration (Figs 1, 3, 5–7). Black; scape, pedicel, anterior part of clypeus, mandible base, fore and mid legs excluding black coxal base, apex of hind trochantellus, base of hind tibia, apices of hind tarsomeres I-IV, tibial spurs, tarsal claws excluding brown apex, base of metasomal tergite I and apical part of ovipositor sheath yellowish brown to brown; flagellomeres VIII-XVIII, posterior spot of tegula, basal sclerites of fore wing, posterior spot of metasomal tergite VI, and metasomal tergite VII white; palpi, and mandible excluding base and apex whitish yellow; ovipositor reddish brown; wings greyish hyaline.

Male (Figs 2, 4, 8–12). Similar to female, except following characters: body 7.2–9.7 mm long, fore wing 6.0–7.7 mm long; face slightly wider, 0.7–0.8 times as long as wide; malar space 0.9–1.0 times as long as basal width of mandible; OOL/OD 0.9–1.0; POL/OD 1.0–1.2; antenna with 44–46 flagellomeres; flagellomere I 4.3–5.0 times as long as apical width, 1.3 times as long as flagellomere II. Mesosoma 2.3–2.5 times as long as width between tegulae. Metasoma polished, punctate; tergite I 2.0–2.2 times as long as maximum width, 1.3–1.4 times as long as tergite II; hind tibia 8.0–8.4 times as long as wide; 7–9 distal hamuli.

Male terminalia: subgenital plate pentagonal with median portion of apical margin weakly concave (Fig. 12); apex of paramere not projected beyond apex of aedeagus (Figs 8, 9), apical margin round (Fig. 11); aedeagus with basal apodeme about half as long as penis valve (Fig. 10).

Colorationmarkedly differs from female by presence of several yellowish markings (Figs 1–4). Body black; spots on scape and pedicel, mandible excluding brownish apex, clypeus, longitudinal stripe along facial orbit, palpi, collar, pair of anterolateral spots on mesoscutum, V-shaped marking on scutellum, tegula, subalar prominense, spots on posterodorsal corner of mesopleuron and metapleuron, basal sclerites of fore and hind wing, fore and mid coxae, trochanters, trochantelli, apex of hind trochanter and hind tarsomeres I-IV excluding black dorsal surface of tarsomere I whitish yellow; fore and mid legs except whitish yellow parts, base of hind coxa, hind trochanter excluding apex, hind femur excluding blackish base and apex, basal part of hind tibia yellowish to brownish red; flagellomeres XII-XIX, basal 1/3 and posterior margin of metasomal tergite I and posterior margin of metasomal tergite II white. Wings and tarsal claws similar to female. Brown areas on hind femur and tibia sometimes darker

Type series.

Holotype: ♀, Mts. Tilongkabila, Bogani Nani Warta Bone National Park, Prov. Gorontalo, North Sulawesi, INDONESIA, 100m alt., 20.Feb.–3.Mar.2009, Keizo Takasuka leg. (Malaise trap, 0°33.10'N, 123°10.34'E) (EUM). Paratypes: same locality and collector as holotype, 2♂, 100-500m alt., 6–9.Feb.2009 (NIAES); 2♂, 500–800m alt., 6–9.Feb.2009 (BMNH); 1♀, 900m alt., 7–18.Feb.2009 (Malaise trap, 0°34.32'N, 123°11.41'E) (BMNH); 1♀, 1200m alt., 7–19.Feb.2009 (Malaise trap, 0°34.36'N, 123°11.58'E) (NIAES); 1♀, 500m alt., 19.Feb.-2.Mar.2009 (Malaise trap, 0°34.04'N, 123°11.15'E) (EUM); 2♂, 100–500m alt., 2.Mar.2009 (NIAES); 12♂1♀, 500–800m alt., 2.Mar.2009 (EUM).

Distribution.

Indonesia (Sulawesi).

Etymology.

The specific name is derived from Sulawesi, the type locality.

Biological note.

Host unknown. Female wasps were, as a whole, caught in Malaise traps except for one individual caught by sweep net, whilst male wasps were caught exclusively by sweep net. On 2nd March, many male wasps were flying around one (unidentified) shrub growing between 500 and 800m.