(C) 2012 Michael J. Sharkey. This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
For reference, use of the paginated PDF or printed version of this article is recommended.
The species of Thai Zelodia (Hymenoptera: Braconidae: Agathidinae) are revised. Twenty-one species are treated, 19 new species are described, i.e. Zelodia charoeni, Zelodia chongkraii, Zelodia cholathorni, Zelodia idrisi, Zelodia nikomi, Zelodia nopadoli, Zelodia pahangensis, Zelodia panyaii, Zelodia poonsathii, Zelodia ratanae, Zelodia saksiti, Zelodia surachaii, Zelodia suyaneeae, Zelodia toyae, Zelodia uthaii, Zelodia wangi, Zelodia wichaii, Zelodia wirati, Zelodia wirotei. A dichotomous key to species is presented; links to electronic interactive keys and to distribution maps are also included.
Thailand, Insecta, identification key, taxonomy, systematics
Agathidinae is a moderately large subfamily of Braconidae with 1, 061 described species worldwide and 238 in the Oriental Region (
Map showing collection sites in Thailand.
As part of the inventory of Thai insects, we ran three Malaise traps at 30 different localities throughout Thailand from 2007-2010, comprising approximately 90 Malaise trap years. The specimens dealt with here are primarily from these traps.
Species concepts are based on morphological data and cytochrome c oxidase (COI) data. Phenetic and phylogenetic trees, using 558 base pairs of COI data, were constructed using neighbor-joining (NJ), maximum parsimony (MP) and Bayesian methods. MP was performed using TNT (
NJ phylogram of the COI barcode region for 17 of the 21 Zelodia species treated here. Wherever Bayesian and parsimony analyses agreed with the NJ tree branch support values are included in the figure i.e., Bayesian posterior probabilities / parsimony bootstrap (bootstrap values less than 60 are not illustrated).
The dichotomous key, descriptions, and the interactive key (Appendices 1–3) were generated using DELTA Editor
Morphological terms follow
Morphological terms used in this revision were matched to the Hymenoptera Anatomy Ontology (HAO,
All 19 species are treated with a diagnosis and distributional data. They are illustrated with color photos using a JVC digital camera mounted on a Leica MZ16 microscope and Automontage® stacking software. Distributional data are listed for all species and a Google map via Berkeley Mapper is included for all species. The descriptions are of the holotype and variation is given in parentheses.
The source files for the keys, descriptions, illustrations, DNA sequence and distributional data are all freely available to future researchers who may wish to build on this beginning. DNA trace files and primer information are available through the Barcode of Life Data system (BOLD) (
Distribution data, pdf’s of non-copyright references, images, notes, and host and type information can be found by searching TaxaBank (a combined specimen and taxonomic database; http://purl.org/taxabank). Codes beginning with an “H” and followed by numbers are unique identifiers used for specimens in the Sharkey lab at the University of Kentucky, and in the specimen database TaxaBank (e.g., H647).
Abbreviations used for specimen depositories are as follows:BMNH The Natural History Museum, London, England.
FSCA Florida State Collection of Arthropods, Gainesville, Florida, USA.
HIC Hymenoptera Institute Collection, University of Kentucky, Department of Entomology, Lexington, Kentucky, USA.
QSBG Queen Sirikit Botanic Gardens, Chiang Mai, Thailand.
RMNH NCB Naturalis Collection [formerly Rijksmuseum van Natuurlijke Historie], Leiden, Netherlands.
UKM Universiti Kebangsaan, Bangi, Selangor, Malaysia.
ResultsRefer to the tree in Sharkey and Clutts (2010) for generic level placement. The host lepidopterans of the genus are unknown. The neighbor joining branching diagram in Figure 2, based on COI mtDNA, was used to help in determining species limits although we did not devise any cutoff threshold to delimit species. We conducted Bayesian and parsimony analyses and where these agreed with the NJ tree the support values are given in Figure 2. An examination of this figure shows that the NJ diagram and the phylogenetic analyses were in close agreement. We also used morphological differences to delimit species. For example, Zelodia saksiti and Zelodia charoeni are very similar (1/558 bp difference) in COI sequences but are very different morphologically. Contrastingly, Zelodia wangi is a widespread species and COI sequences show some variation (2/558 bp difference), however we could find no morphological differences. The complex may represent several species.
Taxonomy DiagnosisMembers of Zelodia may be distinguished from all other Agathidinae with the following combination of characters: ovipositor short, shorter than length of metasoma; tarsal claws cleft; hind trochantellus with 2 longitudinal carinae; frons lacking lateral carinae between antennae and lateral ocelli.
Key to Thai species of Zelodia(Note: there are often more than 2 alternatives in each couplet)
1 | a. Hind femur all pale | 2 |
b. Hind femur all melanic | 7 | |
c. Hind femur melanic dorsally and ventrally, orange laterally and medially | Zelodia ratanae Sharkey sp.n. | |
2(1) | a. Hind leg including tarsus entirely pale | Zelodia brevifemoralis Achterberg & Long |
b. Hind leg melanic and pale, often mostly pale with tarsus melanic | 3 | |
3(2) | a. Mesoscutum entirely pale | 4 |
b. Mesoscutum mostly melanic except margins pale | Zelodia idrisi Sharkey sp.n. | |
4(3) | a. Stigma entirely melanic | Zelodia nikomi Sharkey sp.n. |
b. Stigma partly or entirely yellow | 5 | |
5(4) | a. Median ocellus distinctly narrower than space between lateral ocelli | 6 |
b. Median ocellus about as widen as space between lateral ocelli | Zelodia toyae Sharkey sp.n. | |
6(5) | a. Hind femur more than 4× longer than wide (average 4.2×) | Zelodia suyaneeae Sharkey sp.n. |
b. Hind femur less than 4× longer than wide (average 3.5×) | Zelodia wichaii Sharkey sp.n. | |
7(1) | a. Mesoscutum entirely pale | 8 |
b. Mesoscutum entirely melanic | 10 | |
c. Mesoscutum mostly pale with melanic spots posteriorly | Zelodia chongkraii Sharkey sp.n. | |
d. Mesoscutum mostly melanic except on margins pale | 12 | |
8(7) | a. Median tergite 1 mostly yellow | 15 |
b. Median tergite 1 all white or cream colored, sometimes with a bit of melanic color subapically | Zelodia wirotei Sharkey sp.n. | |
c. Median tergite 1 all black | 9 | |
9(8) | a. Head (not including mouthparts or antenna) mostly or entirely pale | Zelodia longidorsata (Bhat & Gupta) |
b. Head (not including mouthparts or antenna) mostly or entirely melanic | Zelodia panyaii Sharkey sp.n. | |
10(7) | a. Median tergite 1 all black | Zelodia cholathorni Sharkey sp.n. |
b. Median tergite 1 all white or cream colored or sometimes with a bit of melanic color subapically | 11 | |
11(10) | a. Head (not including mouthparts or antenna) mostly or entirely melanic | Zelodia surachaii Sharkey sp.n. |
b. Head (not including mouthparts or antenna) mostly or entirely pale | Zelodia nopadoli Sharkey sp.n. | |
12(7) | a. Propodeum melanic in anterior half or more | 13 |
b. Propodeum entirely pale | 14 | |
13(12) | a. Hind leg entirely melanic | Zelodia poonsathii Sharkey sp.n. |
b. Hind leg melanic except tibia mostly pale | Zelodia saksiti Sharkey sp.n. | |
14(12) | a. Fore wing not lighter apically than subapically, lacking white setae apically | Zelodia uthaii Sharkey sp.n. |
b. Fore wing lighter apically than subapically, with extensive white setae apically | Zelodia pahangensis Sharkey sp.n. | |
15(8) | a. Hind tibia entirely melanic | Zelodia wangi Sharkey sp.n. |
b. Hind tibia mostly pale | 16 | |
16(15) | a. Median ocellus distinctly narrower than space between lateral ocelli | Zelodia charoeni Sharkey sp.n. |
b. Median ocellus about as wide as space between lateral ocelli | Zelodia wirati Sharkey sp.n. | |
http://species-id.net/wiki/Zelodia_brevifemoralis
Fig. 3Hind leg, including tarsus (except apical tarsomere), entirely pale.
Zelodia brevifemoralisAchterberg and Long a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1–MT2.
TaxaBank#/BOLD Process ID/Genbank Accession: H334/ATRMK197-11/JQ763436.
Recorded from Vietnam and central Thailand.Distribution map of the sole Thai locality can be found at http://purl.org/thaimap/brevifemoralis
H0334 [QSBG] ♀, Thailand, Phu Kradueng NP, 420, 16.844°N, 101.692°E, MT, 1–7.vii.2008.
urn:lsid:zoobank.org:act:85D50740-7E31-4F89-AB31-B8689CC225CF
http://species-id.net/wiki/Zelodia_charoeni
Fig. 4Hind tibia pale except apex black; hind femur melanic; mesoscutum pale; median ocellus distinctly narrower than space between lateral ocelli.
Body length 7.4 mm (7.2–7.4). Head. 41 (39–41) flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells hyaline basally, weakly infuscate apically, veins yellow basally except costa mostly melanic, melanic apically and around parastigma, stigma yellow in basal fifth. Hind tarsal claw bifid. Length/width of hind femur 2.22/0.552 = 3.9 (3.9–4.1). Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 1.257/0.539 = 2.3. Ratio of widest point of MT1 to narrowest point 0.539/0.35 = 1.5 (1.4–1.5). Color. Mostly yellow; black as follows: antenna except scape brown medially, hind leg except basal 4/5 of tibia yellow. Median area of mesopleuron yellow (to mostly brown).
Zelodia charoeni sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1.
TaxaBank#/BOLD Process ID/Genbank Accession: H462/ATRMK214-11/JQ763437.
Found only in northern Thailand.Distribution map can be found at http://purl.org/thaimap/charoeni
Dedicated to Mr. Charoen Wanna, collector at Doi Phuka National Park.
Holotype ♀. H5905 [QSBG], Thailand, Doi Phahompok NP Headquarters, 569, 19.966°N, 99.156°E, MT, 25.vii–1.viii.2007.
Paratypes. Thailand: ♀, Nam Nao NP, Check point, 921m, 16.728°N, 101.563°E, MT, 28.iv–5.v.2007: H0461 [HIC]; ♂, Doi Phahompak Office, 569m, 20.191°N, 99.248°E, MT, 28.v–7.vi.2008: H0462 [QSBG].
urn:lsid:zoobank.org:act:655F182B-8A0E-4EC7-A530-0188F26B0926
http://species-id.net/wiki/Zelodia_cholathorni
Fig. 5Mesoscutum entirely melanic; hind tibia all melanic; head melanic; MT1 black.
Body length 6.0 mm. Head. 38 flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex densely and heavily punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron densely covered with setae. Fore wing cells weakly infuscate, more so apically, veins brown to black. Hind tarsal claw with elongate basal tooth. Length/width of hind femur 1.8/0.7 = 2.7. Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 0.91/0.659 = 1.37. Ratio of widest point of MT1 to narrowest point 0.7/0.4 = 1.5. Color. Black except palpi mostly yellow and metasomal laterotergites and S1–S3.
Zelodia cholathorni sp.n. a lateral habitus b wings c anterodorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1–MT3.
TaxaBank#/BOLD Process ID/Genbank Accession: H035/ATRMK141-09/JQ763438.
Found only at the type locality in northern Thailand.Distribution map can be found at http://purl.org/thaimaps/cholathorni
Dedicated to Mr. Cholathorn Chamnanthip, chief of Doi Phahompok National Park, one of our collecting sites.
Holotype ♂. H0035 [QSBG], Thailand, Doi Chiang Dao WS, Nature Trail, 491m, 19.403°N, 98.922°E, MT, 31.vii–7.viii.2007.
Paratypes. ♂ Thailand, Doi Chiangdao NP, Headquarters, 491m, 19.405°N, 98.922°E, MT, 21–28.viii.2007: H4597 [QSGB], H4593 [HIC].
urn:lsid:zoobank.org:act:9D77F05D-907C-4643-AD41-FFA1E12A74C9
http://species-id.net/wiki/Zelodia_chongkraii
Fig. 6Mesoscutum mostly pale with melanic spots posteriorly.
Description. Body length 7.3 mm (7.1–7.3). Head. 41 (40–41) flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron sparsely covered with setae. Fore wing cells all weakly infuscate, more so apically, veins melanic. Hind tarsal claw bifid. Length/width of hind femur 2.169/0.506 = 4.3. Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.258/0.458 = 2.3 (2.2–2.3). Ratio of widest point of MT1 to narrowest point 0.543/0.405 = 1.3 (1.3–1.4). Color. Yellow, black, and white; metasomal segments 1 and 2, LT3 and S3 all white; yellow as follows: head except antenna, prothorax, most of mesoscutum and scutellar disc, fore leg, mid leg except coxa, trochanter, trochantellus, and base of femur partly melanic; remaining body parts black.
Zelodia chongkraii sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1.
TaxaBank#/BOLD Process ID/Genbank Accession: H905/ATRMK260-11/JQ763439; H996/ATRMK273-11/JQ763440.
Eastern and northeastern Thailand.Distribution map can be found at http://purl.org/thaimap/chongkraii
Dedicated to Mr. Chongkrai Worapongthorn, chief of Doi Inthanon National Park, one of our collecting sites.
Holotype ♀. H0996 [QSBG], Thailand, Huai Nam Dang NP, Thung Buatong, 19.293°N, 98.6°E, MT, 21–28.iv.2008.
Paratype. ♀, Thailand, Kaeng Krachan NP, Panernthung/km27, 950m, 12.8217°N, 99.371°E, MT, 8–15.vi.2009, H0905 [HIC].
urn:lsid:zoobank.org:act:F01DB087-985E-4475-8E3D-D0F50A398CB6
http://species-id.net/wiki/Zelodia_idrisi
Fig. 7Head melanic dorsally and posteriorly, pale ventrally and anteriorly; hind femur all pale.
Body length 9.0 mm (7.6–9.0). Head. 43 (42–43) flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells yellowish basally, infuscate in distal 3rd fourth and clear in apical fourth with white setae, veins yellow basally and apically, brown at midlength, stigma yellow in basal fourth, otherwise dark brown. Hind tarsal claw bifid. Length/width of hind femur 2.63/0.75 = 3.5 (3.5–3.7). Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 1.60/0.82 = 2.0 (1.9–2.0). Ratio of widest point of MT1 to narrowest point 0.82/0.49 = 1.7 (1.5–1.7). Color. Mottled yellow, brown, orange, cream-colored, and black.
Zelodia idrisi sp.n. a lateral habitus b dorsal head c lateral head and mesosoma d wings with buffered light source e wings with normal light source f dorsal head and mesosoma g dorsal propodeum h dorsal MT1–MT3.
Widespread in western Malaysia and likely present in southern Thailand.Distribution map can be found at http://purl.org/thaimap/idrisi
Dedicated to Prof. Idris Abd. Ghani, professor at the School of Environmental and Natural Resource Science, Universiti Kebangsaan, Malaysia.
Holotype ♀. H6733 [UKM], Malaysia, Kuala Lompat, Pahang, 3.695°N, 102.224°E, 3.iii.2009.
Paratypes. ♀, Malaysia: Kampung PETA, Johor, 2.541°N, 103.418°E, viii.2008: H8467 [QSBG]; Kuala Lompat, Pahang, 3.695°N, 102.224°E, 3.iii.2009:H8468 [QSBG]; Wang Kelian, Perlis, 6.678°N, 100.19°E, xi.2008: H8469 [HIC], H8466 [UKM]; Pasoh Forest Reserve, 23.x.2002: H8470 [HIC].
http://species-id.net/wiki/Zelodia_longidorsata
Fig. 8Hind tibia all melanic; mesoscutum entirely pale; stigma entirely melanic; head, except antenna, pale; propodeum mostly or entirely pale; median ocellus diameter equal to the space between lateral ocelli; MT1 long and narrow, not much wider apically than basally
Recorded from southern India and a locality east of Bangkok, undoubtedly more widespread.Distribution map of Thai locality can be found at http://purl.org/thaimap/longidorsata
Zelodia longidorsata a lateral habitus b dorsal head c lateral head and mesosoma d wings e dorsal mesosoma f dorsal propodeum and metasomal terga 1 and 2.
♀, Thailand, Chanda Buri, Pong Nam Ron, 12.908°N, 102.262°E, MT, 20-31.iii.2001: H8471 [RMNH].
urn:lsid:zoobank.org:act:906199C7-511C-4D3E-A709-74059891E100
http://species-id.net/wiki/Zelodia_nikomi
Fig. 9Hind femur all pale; stigma entirely melanic; MT1 long and narrow, only slightly wider apically than basally.
Body length 5.5 mm. Head. 35 flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron sparsely covered with setae. Fore wing cells with a yellowish hue basally, infuscate apically, veins yellow basally, melanic apically, stigma entirely melanic. Hind tarsal claw bifid. Length/width of hind femur 1.83/5.0 = 3.7. Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 0.95/0.585 = 1.6. Ratio of widest point of MT1 to narrowest point 0.585/0.394 = 1.5. Color. Mostly yellow except as melanic as follows: flagellum, lateral surface of scape, hind tarsus, apex of hind tibia.
Zelodia nikomi sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1-MT3.
TaxaBank#/BOLD Process ID/Genbank Accession: H518/ATRMK220-11/JQ763441.
Known only from the type locality in central Thailand.Distribution map can be found at http://purl.org/thaimap/nikomi
Dedicated to Mr. Nikom Wongwan, collector at Phuka National Park.
Holotype ♀. H0518 [QSBG], Thailand, Phu Kradueng NP, 420m, 16.844°N, 101.692°E, MT, 9–16.iii.2009.
urn:lsid:zoobank.org:act:CDC346AF-61D7-4CA3-88A6-33318DA2939E
http://species-id.net/wiki/Zelodia_nopadoli
Fig. 10Mesoscutum entirely melanic; head, except antenna, pale.
Body length 7.7 mm (6.2–7.7). Head. (37–41) flagellomeres (both flagella broken in holotype). Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron with moderately dense setae. Fore wing cells hyaline basally, infuscate distally, veins including stigma melanic. Hind tarsal claw bifid. Length/width of hind femur 2.059/0.551 = 3.8 (3.5–4.8). Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.231/0.583 = 2.1 (2.1–2.3). Ratio of widest point of MT1 to narrowest point 0.583/0.376 = 1.6 (1.2–1.6). Color. Mostly black with yellow, white and cream color; head yellow, scape brown, flagellum black, prothorax yellow; fore leg and mid leg, except parts of coxa, yellow; MT1 and MT2 cream colored; basal laterotergites and sterna white; remainder of body black.
Zelodia nopadoli sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1.
Recorded from southeastern and northwestern Thailand.Distribution map can be found at http://purl.org/thaimap/nopadoli
Dedicated to Mr. Nopadol Nachin, chief of Tad Tone National Park.
Holotype ♀. H0323 [QSBG], Thailand, Khao Khitchakut NP, N/Prabaht Unit, 500m, 12.816°N, 102.152°E, MT, 1–8.ix.2008.
Paratypes. ♀, Thailand: Doi Inthanon NP, Chiang Mai, 1700m, 18.544°N, 98.525°E, MT, 3-12.v.1900: H8491[HIC]; Doi Chiangdao NP, Pha Tang substation, 526m, 19.416°N, 98.9147°E, MT, 28.viii–4.ix.2007: H0948 [QSBG].
urn:lsid:zoobank.org:act:C2307CCB-C04A-46EF-8244-B83DCD4899EE
http://species-id.net/wiki/Zelodia_pahangensis
Fig. 11Mesoscutum mostly melanic except margins pale; hind tibia mostly pale except apex and extreme base black; hind femur melanic; apex of fore wing hyaline with white setae.
Body length 7.8 mm (7.4–7.8). Head. 39 (39–42) flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron sparsely covered with setae. Fore wing cells weakly infuscate, veins mostly brown, lighter basally and apically, stigma yellowish brown in basal 1/3, apex of wing with white setae, making it appear lighter. Hind tarsal claw bifid. Length/width of hind femur 2.32/0.5 = 4.6 (4.5–4.6). Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.3/0.59 = 2.2 (2.1–2.2). Ratio of widest point of MT1 to narrowest point 0.59/0.38 = 1.6 (1.5–1.6). Color. Mottled yellow, cream-colored and black; head yellow except vertex brown and antenna black, hind leg black except most of tibia pale.
Zelodia pahangensis sp.n. a lateral habitus b dorsal head c lateral head and mesosoma d apex of fore wing e wings f dorsal head and mesosoma g dorsal propodeum h dorsal MT1–MT3.
Known only from the type locality in western Malaysia but likely to be found in peninsular Thailand.Distribution map can be found at http://purl.org/thaimap/pahangensis
The name is in reference to the Malaysian state, Pahang, in which the specimens were collected
Holotype ♀, H8489 [UKM], Malaysia, Pahang, Cameron Highland, Parit Falls, 4.493°N, 101.389°E, MT, i.2009.
Paratypes. ♂ Malaysia, Pahang, Cameron Highland: Jasa, 4.493°N, 101.389°E, MT, 28.ii.2009: H8482 [HIC], H8484 [HIC], H8485 [QSBG], H8488 [UKM]; Parit Falls, 4.493°N, 101.389°E, MT, i.2009: H8486 [QSBG], H8487 [QSBG], H8483 [HIC], H8465 [UKM].
urn:lsid:zoobank.org:act:CAE2FCB9-74C6-4B1B-B4A8-35216ACA9154
http://species-id.net/wiki/Zelodia_panyaii
Fig. 12Head (not including mouthparts) melanic; mesoscutum entirely pale.
Body length 7.5 mm. Head. 39 flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron densely covered with setae. Fore wing cells weakly infuscate, more so apically, veins melanic. Hind tarsal claw bifid. Length/width of hind femur 2.117/0.539 = 3.9. Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.234/0.505 = 2.4. Ratio of widest point of MT1 to narrowest point 0.505/0.421 = 1.2. Color. Black, reddish orange, brown, cream colored and white: mostly black, reddish orange as follows: pro and mesothorax, most of fore leg; mid leg mostly brown; fore and mid tarsi yellow; mouthparts cream colored and yellow, anterior laterotergites and sterna white.
Zelodia panyaii sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1 and MT2.
TaxaBank#/BOLD Process ID/Genbank Accession: H918/ATRMK263-11/JQ763442.
Known only from the type locality in northwestern Thailand.Distribution map can be found at http://purl.org/thaimap/panyaii
Dedicated to Mr. Panya Kotesanlee, chief of Phuphan National Park.
Holotype ♀ H0918 [ QSBG], Thailand, Doi Chiangdao NP, Pha Tang substation, 19.416°N, 98.9147°E, MT, 15-21.v.2008.
urn:lsid:zoobank.org:act:B98E3538-0386-430E-9681-50360DD1CD74
http://species-id.net/wiki/Zelodia_poonsathiti
Fig. 13Mesoscutum mostly melanic except margins pale; hind tibia all melanic.
Body length. 7.2 mm. Head. 39 flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron sparsely covered with setae. Fore wing cells weakly infuscate, veins mostly brown to black, M+Cu yellow. Hind tarsal claw bifid. Length/width of hind femur 2.059/0.586 = 3.5. Metasoma. Length/width ratio of MT1, 1.273/0.656 = 1.9. Ratio of widest point of MT1 to narrowest point 0.656/0.436 = 1.5. Color. Mottled yellow, cream-colored and black, head yellow except flagellum black and scape brown, hind leg black, MT1 and MT2 cream colored.
Zelodia poonsathiti sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1.
TaxaBank#/BOLD Process ID/Genbank Accession: H647/ATRMK231-11/JQ763443.
Found only at the type locality in southern peninsular Thailand.Distribution map can be found at http://purl.org/thaimap/poonsathiti
Dedicated to Mr. Poonsathit Wongsawat, chief of Doi Phuka National Park.
Holotype ♂ H0647 [QSBG], Thailand, Khao Sok NP, Bang Huaraed, 122m, 8.909°N, 98.509°E, MT, 11-18.xi.2008.
urn:lsid:zoobank.org:act:2738BC9E-B243-45EE-B970-90AE358BBDB7
http://species-id.net/wiki/Zelodia_ratanae
Fig. 14Hind tibia mostly pale, extreme base cream, apex black, most of dorsal and ventral surfaces black, otherwise orange.
Body length 7.6 mm. Head. 42 flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex densely and heavily punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron sparsely covered with setae. Fore wing cells and veins melanic in basal 3/5, cells hyaline basally with white setae, veins R1, RS, r-m, and M yellow distally, stigma yellow in apical third. Hind tarsal claw bifid. Length/width of hind femur 2.4/0.8 = 3.2. Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 1.112/1.245 = 0.88. Ratio of widest point of MT1 to narrowest point 1.2/0.4. Color. Black except as follows; scape yellow, margins of prothorax cream colored to yellow, legs with extensive orange and/or yellow, scutellar sulcus, lateral and posterior areas of scutellum, and much of metapectal-propodeal complex, yellowish orange, anterior laterotergites and sterna pale brown.
Zelodia ratanae sp.n. a lateral habitus b dorsal head c lateral head and mesosoma d wings e dorsal head and mesosoma f dorsal propodeum g dorsal metasoma.
TaxaBank#/Genbank Accession: H273 /JQ763452.
Found only at the type locality in southern peninsular Thailand.Distribution map can be found at http://purl.org/thaimap/ratanae
Dedicated to Ms. Ratana Luckanawarakul. She is currently the director of the Recreation and Interpretation Division of the Thai Department of National Parks. Ratana was of great assistance to us on numerous field trips and was always the best of company.
Holotype ♂. H0273 [QSBG], Thailand, Khao Pu-Khao Ya NP, 75m, 7.551°N, 99.789°E, MT, 8–10.v.2006.
urn:lsid:zoobank.org:act:AE7C81C4-5E9B-477E-87A0-D80FC221940F
http://species-id.net/wiki/Zelodia_saksiti
Fig. 15Mesoscutum mostly melanic except margins pale; hind tibia mostly pale except apex black; propodeum color mostly or entirely melanic.
Body length 8.4 mm (7.2–8.4). Head. 41 (40–41) flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells weakly infuscate, veins melanic. Hind tarsal claw bifid. Length/width of hind femur 2.007/0.533 = 3.8 (3.8–4.0). Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.318/0.62 = 2.1 (2.0–2.1). Ratio of widest point of MT1 to narrowest point 0.620/0.389 = 1.6 (1.6–1.7). Color. Mottled yellow, orange, cream-colored and black, head yellow except antenna black, hind leg black except most of tibia pale.
Zelodia saksiti sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1-MT3.
TaxaBank#/BOLD Process ID/Genbank Accession: H036/ATRMK142-09/JQ763444.
Widespread throughout western Thailand.Distribution map can be found at http://purl.org/thaimap/saksiti
Dedicated to Saksit Poonsapsiri, one of the chiefs of Phataem National Park.
Holotype ♀. H0324 [QSBG], Thailand, Khao Sok NP, Head quarters, 115m, 8.915°N, 98.53°E, MT, 12-19.v.2009.
Paratypes. ♂, Thailand: Salaeng Luang NP: 557m, 16.844°N, 100.882°E, MT, 25.viii–1.ix.2006: H0036 [QSBG], H8464 [UKM]; 479m, 16.839°N, 100.863°E, MT, 11–18.viii.2006: H5996 [QSBG]; ♀, Khao Sok NP, Headquarters, 155m, 8.915°N, 98.53°E, MT, 9–16.vi.2009: H0663 [HIC]; ♂, evergreen forest, MT, 1988: H8490[BMNH]; ♀, Kaeng Krachan NP, km16/road/stream2, 320m, 12.802°N, 99.444°E, MT, 25.v–1.vi.2009: H0920 [HIC]; ♀, Doi Chiangdao NP: Headquarters, 491m, 19.405°N, 98.922°E, MT, 18–25.ix.2007: H4108 [QSBG]; Pha Tang substation, 526m, 19.416°N, 98.915°E, MT, 28.viii–4.ix.2007: H5967 [UKM].
urn:lsid:zoobank.org:act:D1619B8D-BE53-440D-A637-B1A40338B4A7
http://species-id.net/wiki/Zelodia_surachaii
Fig. 16Mesoscutum entirely melanic; hind tibia all melanic; head (not including mouthparts or antenna) melanic; apex of flagellum pale, contrasting with melanic base.
Median tergite 1 white or cream colored.
Body length 6.6 mm (6.6–8.3). Head. 41 flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron with moderately dense setae. Fore wing cells weakly infuscate, veins and stigma melanic. Hind tarsal claw bifid. Length/width of hind femur 2.019/0.511 = 4.0 (4.0–4.7). Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.117/0.518 = 2.2 (2.2–2.3). Ratio of widest point of MT1 to narrowest point 0.518/0.427 = 1.2 (1.2–1.4). Color. Black except mouthparts yellow, apical 12 flagellomeres yellowish brown, MT1 cream colored, L1 and L2 and basal sterna white.
Zelodia surachaii sp.n. a lateral habitus b wings c dorsal head d dorsolateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1-MT2.
TaxaBank#/BOLD Process ID/Genbank Accession: H389/ATRMK207-11/JQ763445.
Taiwan, peninsular Malaysia and Thailand.Distribution map can be found at http://purl.org/thaimap/surachaii
Dedicated to Mr. Surachai Pransil, chief of Khao Kitchagoot National Park.
Holotype ♀. H0389 [QSBG] Thailand, Kaeng Krachan NP, Panernthung/km27, 950m, 12.819°N, 99.375°E, MT, 18–25.vii.2008.
Paratypes. ♀, Malaysia, Cameron Highlands, Tanah Rata, 4.475°N, 101.38°E, MT, iii–iv.2001: H1122 [UKM]; ♂, Taiwan, Taipei Co., Chuntou, 9.iv.1999: H8492 [FSCA].
urn:lsid:zoobank.org:act:5B1641DD-9A03-4B56-B7B7-8A1749AE17FE
http://species-id.net/wiki/Zelodia_suyaneeae
Fig. 17Hind femur all pale; mesoscutum entirely pale; hind tibia mostly pale except apex and sometimes extreme base black; stigma partly or entirely yellow; median ocellus distinctly narrower than space between lateral ocelli; lateral surface of hind femur sparsely punctate, spaces between punctures wider than diameter of punctures.
Body length 5.7 mm (5.7–6.4). Head. 40 (40–45) flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells hyaline basally, weakly infuscate apically, veins yellow basally, melanic apically and around parastigma, stigma yellow in basal third. Hind tarsal claw bifid. Length/width of hind femur 1.856/0.531 = 3.5 (3.4–4.4). Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 0.87/0.434 = 2.0 (2.0–2.5). Ratio of widest point of MT1 to narrowest point 0.434/0.269 = 1.3 (1.3 – 1.4). Color. Yellow except as follows: flagellum melanic, lateral surface of scape brown, apex of hind tibia and entire tarsus melanic.
Zelodia suyaneeae sp.n. a lateral habitus b wings c dorsal head d dorsolateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and metasoma.
TaxaBank#/BOLD Process ID/Genbank Accession: H1603/ATRMK321-11/JQ763446; H383/ATRMK206-11/JQ763447.
Widespread in northern and central Thailand.Distribution map can be found at http://purl.org/thaimap/suyaneeae
Dedicated to Dr. Suyanee Vessabutr. She was the deputy director of QSBG and was of great assistance to us in providing space and resources at the museum.
Holotype ♀. H1603 [QSBG], Thailand, Chae Son NP, Visitor center, 18.836°N, 99.4739°E, MT, 15-22.ix.2007.
Paratypes. ♀, Thailand, Khuean Srinagarindra NP, 210m, 14.5°N, 98.884°E, MT, 26.iii-2.iv.2009: H0355 [QSBG], H0359 [QSBG]; 22–29.i.2009: H0383 [HIC]; Chae Son NP, Nature Trail; 507m, 18.836°N, 99.473°E, MT, 14–21.x.2007: H4103 [HIC]; 1–8.x.2007: H4119 [HIC]; Doi Chiangdao NP, Headquarters, 491m, 19.405°N, 98.922°E, MT: 18–25.ix.2007: H4113 [HIC]; 19.404°N, 98.9218°E, MT, 21–27.v.2008: H0891 [UKM]; Phu Kradueng NP, 274m, 16.942°N, 101.694°E, MT, 26.iv–3.v.2008: H0572 [UKM]; Kaeng Krachan NP, km16/road/stream2, 320m, 12.802°N, 99.444°E, MT, 25.v–1.vi.2009: H0921 [UKM]; ♂, Phu Phan NP, 526m, 16.81°N, 103.891°E, MT, 16–22.vi.2007: H0293 [QSBG].
urn:lsid:zoobank.org:act:26B4E73C-65B5-4589-AB1D-BBD907DE0FDA
http://species-id.net/wiki/Zelodia_toyae
Fig. 18Hind femur all pale; mesoscutum entirely pale; hind tibia mostly pale except apex black; median ocellus about as wide as or wider than space between lateral ocelli; stigma partly yellow.
Body length 7.6 mm. Head. 41 flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells hyaline basally, weakly infuscate apically, veins yellow basally except costa mostly melanic, veins melanic apically and around parastigma, stigma yellow in basal third. Hind tarsal claw bifid. Length/width of hind femur 2.25/0.54 = 4.2. Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 1.28/0.59 = 2.2. Ratio of widest point of MT1 to narrowest point 0.59/0.325 = 1.8. Color. Yellow except as follows: flagellum melanic, lateral surface of scape brown, apex of hind tibia and entire tarsus melanic.
Zelodia toyae sp.n. a lateral habitus b wings c dorsal head d lateral head and mesosoma e dorsal head and mesosoma f dorsal propodeum g dorsal metasoma.
TaxaBank#/BOLD Process ID/Genbank Accession: H381/ATRMK205-11/JQ763448.
Known only from the type locality in northwestern Thailand.Distribution map can be found at http://purl.org/thaimap/toyae
Dedicated to Ms. Chayanit (Toy) Satatha. Toy was the sorter of Diptera for the TIGER project and is currently a technician at QSBG. The species name reflects her nick-name, Toy.
Holotype ♀. H0381 [QSBG], Thailand, Namtok Mae Surin NP, Car park, 19.36°N, 97.988°E, MT, 27.iv–4.v.2008.
urn:lsid:zoobank.org:act:1BE5192A-344A-4A1D-9959-0255A5A9976A
http://species-id.net/wiki/Zelodia_uthaii
Fig. 19Mesoscutum mostly melanic except margins pale; hind tibia mostly pale except apex and extreme base black; hind femur all melanic; apical half of wing not noticeably more infuscate than base.
Body length. 6.1 mm. Head. 42 flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron sparsely covered with setae. Fore wing cells hyaline basally, weakly infuscate distally, veins yellow basally except costa black, veins melanic distally, stigma melanic. Hind tarsal claw bifid. Length/width of hind femur 2.05/0.54 = 3.8. Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.13/0.61 = 1.9. Ratio of widest point of MT1 to narrowest point 0.61/0.38 = 1.6. Color. Mottled yellow, cream-colored and black, head yellow except antenna black, hind leg black except most of tibia pale.
Zelodia uthaii sp.n. a lateral habitus b wings c anterior head d dorsal head e lateral head and mesosoma f dorsal head and mesosoma g dorsal propodeum h dorsal MT1–MT3.
TaxaBank#/BOLD Process ID/Genbank Accession: H309/ATRMK450-11/JQ763449.
Known only from the type locality in western Thailand.Distribution map can be found at http://purl.org/thaimap/uthaii
Dedicated to Mr. Uthai Promnaree, one of the chiefs of Phataem National Park.
Holotype ♀. H0309 [QSBG], Thailand, Mae Wong NP, Chong Yen, 1306m, 16.087°N, 99.11°E, MT, 24.ix–1.x.2007.
urn:lsid:zoobank.org:act:14DE5D59-9589-46DA-AE41-D0530F98EB1C
http://species-id.net/wiki/Zelodia_wangi
Fig. 20Scutum and MT1 yellow; hind leg, including tibia, melanic.
Body length 6.7 mm (6.5–7.1). Head. 43 (41–45) flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells very weakly infuscate with a yellowish tinge basally, veins yellow basally, melanic in apical half, stigma yellow in basal fourth (or entirely melanic). Hind tarsal claw bifid. Length/width of hind femur 2.0/0.5 = 3.9 (3.7–3.9). Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 1.156/0.481 = 2.4 (2.4–2.5). Ratio of widest point of MT1 to narrowest point 0.481/0.349 = 1.4 (1.4). Color. Yellow except as follows: scape and pedicel pale brown, flagellum brown, hind leg mostly brown in females with areas near joints and most of tibia pale brown to yellow, anterior metasomal laterotergites and sterna cream-colored, median tergites entirely yellow in holotype (but some female paratypes have pale brown markings on apical tergites; males as in females but with hind leg mostly to entirely black, and apical terga and sterna extensively black).
Zelodia wangi sp. n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum h dorsal MT1–MT3.
TaxaBank#/BOLD Process ID/Genbank Accession: H074/JQ763457; H023/JQ763455; H073/JQ763456; H019/JQ763458; H025/JQ763460; H084/JQ763459.
Widespread throughout Thailand. Distribution map can be found at http://purl.org/thaimap/wangi
Dedicated to Mr. Wang Saeyang, who was the sorter for Hymenoptera for the TIGER project.
Holotype ♀. H0057 [QSBG], Thailand, Khoa Chong, Forest Research Stn., 75m, 7.551°N, 99.789°E, MT, vii–xi.2005.
Paratypes. ♀, Thailand: Khoa Chong, Forest Research Stn., 75m, 7.551°N, 99.789°E, MT, xi.2005: H0019 [QSBG], H5990 [UKM], H0084 [HIC]; x.2005: H0023 [QSBG]; viii.2005: H0054 [QSBG], H5906 [UKM], H5988 [UKM], H5991 [UKM], H5994 [UKM], H0061 [UKM]; ♂, x.2005: H5992 [UKM]; viii.2005: H5986 [UKM]; ♀, Thailand, Nayong Khaochong, 75m, 7.561°N, 99.886°E, MT, 24–27.vi.2005: H0025 [QSBG], H0034 [QSBG]; ♀, Nayong Khaochong, 75m, 7.561°N, 99.886°E, MT, 15–18.vi.2005: H0074 [HIC]; 6-8.vii.2005: H0076 [HIC]; ♂, 26–30.vii.2005: H0073 [HIC]; ♀, Khao Sok NP, Headquarters, 115m, 8.915°N, 98.53°E, MT, 12–19.v.2009: H0391 [QSBG]; 17–24.iii.2009: H0475 [QSBG]; 8–15.ix.2008: H0477 [QSBG]; ♂, 28.x–4.xi.2008: H0304 [QSBG]; 25.xi-2.xii.2008: H0310 [QSBG]; 28.iv–5.v.2009: H0332 [QSBG]; 21-28.iv.2009: H0592 [UKM]; 9–16.xii.2008: H0673 [HIC]; 18–25.xi.2008: H0675 [HIC]; 24-31.iii.2009: H0683 [HIC]; 31.viii–8.ix.2008: H0758 [HIC]; 9–16.vi.2009: H0972 [HIC], H0691 [HIC], H0666 [UKM]; ♂, Namtok Yong NP, 95m, 8.174°N, 99.742°E, MT, 9–16.ii.2009: H0351 [QSBG]; Namtok Mae Surin NP, Near Visitor Ctr, 228m, 19.358°N, 97.988°E, MT, 8–15.vii.2007: H5987 [UKM], H0617 [UKM], H0620 [UKM]; Namtok Mae Surin NP, 334m, 19.344°N, 97.988°E, MT, 26.viii–2.ix.2007: H5980 [HIC]; ♂, Namtok Yong NP, 80m, 8.174°N, 99.742°E, MT, 23–30.ix.2008: H0979 [HIC]; Mae Surin near visitor center, 228m, 19.489°N, 98.056°E, MT. 15–22.vii.2007: H0442 [QSBG], H0443 [QSBG]; ♀, Khao Pu-Khao Ya NP, 75m, 7.551°N, 99.789°E, MT, 28–30.v.2006: H5989 [UKM], H5995 [UKM]; 10–13.ii.2006: H5993 [UKM]; Tat Tone NP, 250m, 15.984°N, 102.035°E, litter, 23–26.vi.2006: H5997 [UKM]; Kaeng Krachan NP, km33/helipad, 735m, 12.836°N, 99.345°E, MT, 18–25.ii.2009: H0623 [UKM]; ♂, Doi Chiangdao NP, Pha Tang substation, 526m, 19.4163°N, 98.9147°E, MT, 3–10.xii.2007: H0926 [HIC].
urn:lsid:zoobank.org:act:3FF67B83-DB04-4CB8-A339-47B60BB77C10
http://species-id.net/wiki/Zelodia_wichaii
Fig. 21Hind femur all pale; mesoscutum entirely pale; hind tibia mostly pale except apex black; stigma partly or entirely yellow; ocelli median ocellus distinctly narrower than space between lateral ocelli; lateral surface of hind femur densely punctate to aciculate, spaces between punctures not wider than diameter of punctures.
Body length 6.3 mm (6.0–6.4). Head. 40 flagellomeres. Median ocellus diameter narrower than space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells hyaline basally, weakly infuscate apically, veins yellow basally, melanic apically and around parastigma, stigma yellow in basal third. Hind tarsal claw bifid. Length/width of hind femur 1.027/0.519 = 3.7 (3.4–3.7). Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 1.038/0.49 = 2.1 (2.0–2.1). Ratio of widest point of MT1 to narrowest point 0.49/0.313 = 1.6. Color. Yellow except as follows: antenna melanic, apex of hind tibia and entire tarsus melanic.
Zelodia wichaii sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1–MT3.
TaxaBank#/Genbank Accession: H063/JQ763453.
Distribution. Recorded from two localities in peninsular Thailand. Distribution map can be found at http://purl.org/thaimap/wichaii
Dedicated to Mr. Wichai Srisuka, the curator of insects at QSBG.
Holotype ♀. H0063 [QSBG], Thailand, Khoa Chong, Trang, 75m, 7.551°N, 99.789°E, MT, viii.2005.
Paratype. ♀, Thailand, Namtok Yon NP, Klong Jang Waterfall, 154m, 8.338°N, 99.827°E, MT, 12–19.v.2009: H0594 [HIC].
urn:lsid:zoobank.org:act:73E375A4-5D6C-4668-9A49-95F6BB6B1E5A
http://species-id.net/wiki/Zelodia_wirati
Fig. 22Hind tibia mostly pale except apex black; hind femur all melanic; mesoscutum entirely pale; median ocellus about as wide as or wider than space between lateral ocelli.
Body length 6.9 mm (male 5.8 – 6.7 mm). Head. 41 flagellomeres (41–43 males). Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli partly smooth, otherwise with weak punctures. Metapleuron sparsely covered with setae. Fore wing cells hyaline basally, weakly infuscate apically, veins yellow basally, except costa mostly melanic, melanic apically and around parastigma, stigma yellow in basal fifth. Hind tarsal claw bifid. Length/width of hind femur 2.28/.58 = 3.9 (3.45 – 3.52 males). Lateral surface of hind femur punctate. Metasoma. Length/width ratio of MT1, 1.22/.633 = 1.93. Ratio of widest point of MT1 to narrowest point .633/.385 = 1.6 (1.4–1.6). Color. Mostly yellow, black as follows: antenna except scape brown medially, hind leg except basal 4/5 of tibia yellow.
Zelodia wirati sp.n. a lateral habitus b wings c dorsal head d lateral head e lateral mesosoma f dorsal propodeum g dorsal MT1–MT2.
TaxaBank#/BOLD Process ID/Genbank Accession: H012/JQ763454; H949/ATRMK451-11/JQ763450.
Recorded from three localities in northwestern Thailand.Distribution map can be found at http://purl.org/thaimap/wirati
Dedicated to Mr. Wirat Sukho, collector for the TIGER project at Khao Yai National Park.
Holotype ♀.H0981 [QSBG], Thailand, Doi Phahompok NP, 569m, 19.966°N, 99.156°E, MT, 28.viii–4.ix.2007.
Paratypes. ♂ Thailand: Doi Inthanon NP, campground pond, 1200m, 18.544°N, 98.525°E, MT, 16–24.viii.2006: H0012 [QSBG]; Doi Chiangdao NP, Headquarters, 491m, 19.404°N, 98.9218°E, MT, 15–22.iv.2008: H0909 [HIC]; Chae Son NP, Doi Laan, 1528m, 18.825°N, 99.411°E, MT, 22–28.iii.2008: H949 [UKM].
urn:lsid:zoobank.org:act:8895B38A-E94B-42C3-82E8-77E10978141D
http://species-id.net/wiki/Zelodia_wirotei
Fig. 23Hind tibia all melanic; mesoscutum entirely pale; MT1 all white or cream colored, sometimes with a bit of melanic color subapically.
Body length 5.9 mm. Head. 35 (35–37) flagellomeres. Median ocellus diameter equal to the space between lateral ocelli. Vertex sparsely and weakly punctate. Mesosoma. Notauli mostly or entirely crenulate. Metapleuron densely covered with setae. Fore wing cells all weakly infuscate, veins all melanic. Hind tarsal claw bifid. Length/width of hind femur 1.714/0.475 = 3.5 (3.5–3.7). Lateral surface of hind femur aciculate. Metasoma. Length/width ratio of MT1, 3.978/0.350 = 2.8 (2.8–3.0). Ratio of widest point of MT1 to narrowest point 0.350/0.248 = 1.4. Color. Mostly black with orange, white, brown and cream color; head mostly orange, except area around ocelli slightly darker and antenna black; prothorax orange to brown; mesoscutum orange; scutellum brown, mesopleuron mostly black but orange-brown dorsally; fore leg yellow to cream colored; mid leg yellow to cream colored except coxa mostly melanic; MT1, M2 and part of MT3 cream colored (MT1 sometimes with a tan area posteromedially); anterior laterotergites and sterna white.
Zelodia wirotei sp.n. a lateral habitus b wings c anterior head d lateral head e lateral mesosoma f dorsal mesosoma g dorsal propodeum and MT1–MT2.
TaxaBank#/BOLD Process ID/Genbank Accession: H990/ATRMK272-11/JQ763451.
Recorded only from the type locality south-west of Bangkok. Distribution map can be found at http://purl.org/thaimap/wirotei
Dedicated to Mr. Wirote Naknan, chief of Namnao National Park.
Holotype ♀. H0634 [QSBG], Thailand, Kaeng Krachan NP, km33/helipad, 735m, 12.836°N, 99.345°E, MT, 11–18.v.2009.
Paratype. ♀, Thailand, Kaeng Krachan NP, Panernthung/km27, 950m, 12.822°N, 99.371°E, MT, 8–15.vi.2009: H0990 [HIC].
We thank all of the staff at Queen Sirikit Botanic Gardens in Chiang Mai for sorting the many hundreds of samples and for the Thai park staff for running Malaise traps and other collection devices. Thanks to Dr. van Achterberg for the loan of type specimens. A special thanks to Chaweewan Hutacharern for managing the Thai end of the TIGER project. Funding was provided by NSF grants DEB-0542864 and EF-0337220.
DELTA data matrix, images, and other files to the dichotomous key for Zelodia (Hymenoptera: Braconidae: Agathidinae) from Thailand. doi: 10.3897/JHR.26.2527.app1
Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
DELTA data matrix, images, and other files to species descriptions for Zelodia (Hymenoptera: Braconidae: Agathidinae) from Thailand. doi: 10.3897/JHR.26.2527.app2
Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Interactive key, in IntKey format, to Zelodia (Hymenoptera: Braconidae: Agathidinae) from Thailand. doi: 10.3897/JHR.26.2527.app3
Explanation note: To run the identification key, you will need Windows 95/NT or a later version.
You also need to download Intkey software and reboot your computer, if it is not already installed. The software package, Intkey, can be downloaded from http://delta-intkey.com/www/programs.htm. Once Intkey is installed you need only click on the .ink fi le (below) and the key will open. Click on any character on the left to begin.
More details on how to use Intkey efficiently are found at http://fl orabase.calm.wa.gov.au/help/keys/intkey_tutorial.pdf
Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same freedom for others, provided that the original source and author(s) are credited.
Morphological terms matched to the Hymenoptera Anatomy Ontology. Identifiers (URIs) represent anatomical concepts in HAO version http://purl.obolibrary.org/obo/hao/2011-05-18/hao.owl