Research Article |
Corresponding author: Andrew M.R. Bennett ( andrew.bennett@canada.ca ) Academic editor: Gavin Broad
© 2019 Andrew M.R. Bennett, Sophie Cardinal, Ian D. Gauld, David B. Wahl.
This is an open access article distributed under the terms of the CC0 Public Domain Dedication.
Citation:
Bennett AMR, Cardinal S, Gauld ID, Wahl DB (2019) Phylogeny of the subfamilies of Ichneumonidae (Hymenoptera). Journal of Hymenoptera Research 71: 1-156. https://doi.org/10.3897/jhr.71.32375
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A combined morphological and molecular phylogenetic analysis was performed to evaluate the subfamily relationships of the parasitoid wasp family Ichneumonidae (Hymenoptera). Data were obtained by coding 135 morphological and 6 biological characters for 131 exemplar species of ichneumonids and 3 species of Braconidae (the latter as outgroups). The species of ichneumonids represent all of the 42 currently recognized subfamilies. In addition, molecular sequence data (cytochrome oxidase I “DNA barcoding” region, the D2 region of 28S rDNA and part of the F2 copy of elongation factor 1-alpha) were obtained from specimens of the same species that were coded for morphology (1309 base pairs total). The data were analyzed using parsimony and Bayesian analyses. The parsimony analysis using all data recovered previously recognized informal subfamily groupings (Pimpliformes, Ophioniformes, Ichneumoniformes), although the relationships of these three groups to each other differed from previous studies and some of the subfamily relationships within these groupings had not previously been suggested. Specifically, Ophioniformes was the sister group to (Ichneumoniformes + Pimplformes), and Labeninae was placed near Ichneumoniformes, not as sister group to all Ichneumonidae except Xoridinae. The parsimony analysis using only morphological characters was poorly resolved and did not recover any of the three informal subfamily groupings and very few of the relationships were similar to the total-evidence parsimony analysis. The molecular-only parsimony analysis and both Bayesian analyses (total-evidence and molecular-only) recovered Pimpliformes, a restricted Ichneumoniformes grouping and many of the subfamily groupings recovered in the total-evidence parsimony analysis. A comparison and discussion of the results obtained by each phylogenetic method and different data sets is provided. It is concluded that the molecular characters produced results that were relatively consistent with traditional, non-phylogenetic concepts of relationships between the ichneumonid subfamilies, whereas the morphological characters did not (at least not by themselves). The inclusion of both molecular and morphological characters using parsimony produced a topology that was the closest to the traditional subfamily relationships. The method of analysis did not greatly affect the overall topology for the molecular-only analyses, but there were differences between Bayesian and parsimony results for the total-evidence analyses (especially near the root of the tree). The Bayesian results did not seem to be altered very much by the inclusion of morphological characters, unlike in the parsimony analysis. In summary, the following groups were supported in multiple analyses regardless of the characters used or method of tree-building: Pimpliformes, higher Ophioniformes, higher Pimpliformes, (Claseinae + Pedunculinae), (Banchinae + Stilbopinae), Campopleginae, Cremastinae, Diplazontinae, Ichneumoninae (including Alomya), Labeninae, Ophioninae, Poemeniinae, Rhyssinae, and Tersilochinae sensu stricto. Conversely, Ctenopelmatinae and Tryphoninae were never recovered without inclusion of other taxa. Based on the hypothesis of relationships obtained by the total-evidence parsimony analysis, the following formal taxonomic changes are proposed: Alomyinae Förster (= Alomya Panzer and Megalomya Uchida) is once again synonymized with Ichneumoninae and is now considered a tribe (Alomyini rev. stat.); and Notostilbops Townes is transferred from Stilbopinae to Banchinae, tribe Atrophini.
Ichneumonidae, phylogeny, parsimony, Bayesian, classification, taxonomy
The catalogue of
Comparison of extant subfamilies of Ichneumonidae recognized by recent studies. Column 1: subfamilies recognized by morphological studies alone (up to 2002). Column 2: most recent analysis of all subfamily relationships using both morphological and molecular data. Column 3: subfamilies recognized following current study. *Brachyscleromatinae is now known as Sisyrostolinae (
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Current study (after formal changes) |
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Acaenitinae | Acaenitinae | Acaenitinae |
Adelognathinae | Adelognathinae | Adelognathinae |
Agriotypinae | Agriotypinae | Agriotypinae |
Alomyinae (previously part of Ichneumoninae) | ||
Anomaloninae | Anomaloninae | Anomaloninae |
Ateleutinae (previously part of Cryptinae) | ||
Banchinae | Banchinae | Banchinae (including Notostilbops) |
Brachycyrtinae | Brachycyrtinae | Brachycyrtinae |
Brachyscleromatinae* (previously part of Phrudinae) | ||
Campopleginae | Campopleginae | Campopleginae |
Claseinae | Claseinae | Claseinae |
Collyriinae | Collyriinae | Collyriinae |
Cremastinae | Cremastinae | Cremastinae |
Cryptinae | Cryptinae | Cryptinae |
Ctenopelmatinae | Ctenopelmatinae | Ctenopelmatinae |
Cylloceriinae | Cylloceriinae | Cylloceriinae |
Diacritinae | Diacritinae | Diacritinae |
Diplazontinae | Diplazontinae | Diplazontinae |
Eucerotinae | Eucerotinae | Eucerotinae |
Hybrizontinae (previously Paxylommatinae) | Hybrizontinae (previously Paxylommatinae) | Hybrizontinae |
Ichneumoninae | Ichneumoninae | Ichneumoninae (including Alomyini) |
Labeninae | Labeninae | Labeninae |
Lycorininae | Lycorininae | Lycorininae |
Mesochorinae | Mesochorinae | Mesochorinae |
Metopiinae | Metopiinae | Metopiinae |
Microleptinae | Microleptinae | Microleptinae |
Neorhacodinae | Neorhacodinae | |
Nesomesochorinae (including Nonninae of |
Nesomesochorinae | |
Ophioninae | Ophioninae | Ophioninae |
Orthocentrinae | Orthocentrinae | Orthocentrinae |
Orthopelmatinae | Orthopelmatinae | Orthopelmatinae |
Oxytorinae | Oxytorinae | Oxytorinae |
Pedunculinae | Pedunculinae | Pedunculinae |
Phrudinae | ||
Phygadeuontinae (previously part of Cryptinae) | ||
Pimplinae | Pimplinae | Pimplinae |
Poemeniinae | Poemeniinae | Poemeniinae |
Rhyssinae | Rhyssinae | Rhyssinae |
Sisyrostolinae* | ||
Stilbopinae | Stilbopinae | Stilbopinae (excluding Notostilbops) |
Tatogastrinae | Tatogastrinae | Tatogastrinae |
Tersilochinae | Tersilochinae (including Neorhacodinae + part of Phrudinae) | Tersilochinae |
Tryphoninae | Tryphoninae | Tryphoninae |
Xoridinae | Xoridinae | Xoridinae |
From 1998–2009, several studies used single gene molecular evidence (the D2–D3 region of 28S ribosomal DNA) to examine ichneumonid subfamily relationships, either combined with morphological characters (
More recently,
In terms of the relationships of the subfamilies, prior to about 1990, relationships could only be inferred by the their relative arrangement in Henry Townes’s Genera of Ichneumonidae (
For the purposes of this study, the following definitions for the subfamily groupings are used:
1) Ophioniformes includes the following 18 subfamilies: Anomaloninae, Banchinae, Campopleginae, Cremastinae, Ctenopelmatinae, Hybrizontinae, Lycorininae, Mesochorinae, Metopiinae, Oxytorinae, Neorhacodinae, Nesomesochorinae, Ophioninae, Sisyrostolinae, Stilbopinae, Tatogastrinae, Tersilochinae and Tryphoninae. This concept is the same as that of
2) Pimpliformes includes the following nine subfamilies: Acaenitinae, Cylloceriinae, Diacritinae, Diplazontinae, Orthocentrinae, Pimplinae, Poemeniinae, Rhyssinae and in addition, Collyriinae. This definition corresponds to the concept of
3) Ichneumoniformes is considered in two senses: a) in the strict sense (Ichneumoniformes sensu stricto) as originally proposed by
In addition,
4) Xoridiformes comprised of Xoridinae
5) Labeniformes comprised of Labeninae
6) Orthopelmatiformes comprised of Orthopelmatinae
7) Brachycyrtiformes comprised of Brachycyrtinae, Claseinae and Pedunculinae.
See the Results section on Support/ relationships of subfamilies (below) for discussion on the support/ placement of these groups in the current study.
There is very strong evidence that Braconidae is the sister group of Ichneumonidae based on morphology (
Ingroup taxa were chosen in order to provide complete coverage of all currently recognized subfamilies based on the most recent study of ichneumonid relationships (
Data matrix for morphological character states of Ichneumonidae and outgroup taxa. Key to the letters in matrix below: a=0/1; b=0/2; c=0/3; d=1/2; e=1/3; f=1/4; g=1/5; h=2/3; k=2/4; m=2/6; n=3/4; p=3/7; q=5/6; r=6/7; s=0/1/3; t=1/5/6; u=2/6/7; v=5/6/7; w=2/5/6/7; x=3/5/6/7.
Taxon | 1 | 2 | 3 | 4 | 5 | 6 | 7 | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | |
Doryctes erythromelas | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 3 | ? | 3 | ? | ? | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | ? | ? | 6 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Rhysipolis sp. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | ? | ? | 6 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Aleiodes terminalis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 3 | ? | 3 | ? | ? | 1 | 1 | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | ? | ? | 6 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Spilopteron occiputale | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | h | 0 | ? | 1 | 0 | 0 | 2 | b | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Coleocentrus rufus | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 3 | ? | 1 | 0 | 1 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Adelognathus sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Agriotypus armatus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Anomalon picticorne | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 2 | 2 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Therion texanum | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Lissonota scutellaris | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 2 | 3 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 3 | ? | 0 | 0 | 1 | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Exetastes bioculatus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 2 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | ? | 0 | 0 | 3 | ? | 0 | 0 | 1 | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Apophua simplicipes | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 3 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | ? | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Sphelodon phoxopteridis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Brachycyrtus wardae | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | ? | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Bathyplectes infernalis | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 |
Campoletis sonorensis | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 |
Campoplex sp. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | c | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Casinaria grandis | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | 1 | 0 | ? | 1 | 1 | 1 | 0 | 3 | ? | ? | 0 | 0 | 2 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Dusona egregia | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | ? | 0 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Hyposoter sp. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 |
Olesicampe sp. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 |
Rhimphoctona macrocephala | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 |
Clasis sp. nov. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 0 | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Collyria catoptron | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | d | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 3 | ? | 3 | ? | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Eiphosoma pyralidis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Xiphosomella setoni | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 |
Agonocryptus chichimecus | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 1 | 0 | ? | 0 | 1 | 0 | 1 | 2 | 0 | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 2 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Ateleute sp. nov. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 4 | 1 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 1 | 3 | ? | 1 | ? | ? | 2 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 6 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | a | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Baryceros texanus | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 3 | ? | ? | 0 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Diapetimorpha brunnea | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 2 | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Echthrus reluctator | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 3 | ? | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Lymeon orbus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 2 | 1 | 2 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | ? | ? | 0 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 0 |
Polytribax contiguus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | a | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 |
Rhytura pendens | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 1 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | h | ? | b | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 0 |
Acrolyta sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 0 |
Endasys patulus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 0 |
Mastrus sp. | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 0 |
Ctenopelma sanguineum | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | s | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Xenoschesis limata | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Euryproctus sentinis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | 0 |
Mesoleptidea decens | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 3 | ? | 3 | ? | ? | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Barytarbes honestus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Himerta luteofacia | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | b | d | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Lathrolestes asperatus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | a | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Perilissus concolor | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | b | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Rhorus bartelti | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Sympherta fucata | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Seleucus cuneiformis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Onarion sp. | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 2 | 3 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | ? | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Westwoodia sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 0 | 1 | 1 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Ctenopelmatinae Genus NZ | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 2 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 |
Cylloceria melancholica | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | ? | 0 | 0 | d | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Diacritus incompletus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Diplazon laetatorius | 0 | 0 | 0 | 0 | 4 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 3 | ? | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Woldstedtius flavolineatus | 0 | 0 | 0 | 0 | 4 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 2 | 3 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Euceros sp. nov. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 3 | ? | 0 | 1 | 0 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Hybrizon rileyi | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 2 | 3 | 0 | 3 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 3 | 1 | 0 | 1 | ? | ? | 3 | 0 | ? | 1 | 0 | 0 | 2 | 1 | 1 | 2 | 1 | ? | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Alomya debellator | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | ? | 0 | 0 | 1 | 2 | 2 | 2 | a | a | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Centeterus euryptychiae | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Phaeogenes hebrus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 3 | 0 | d | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Stenodontus sp. nov. | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | d | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Coelichneumon eximius | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | d | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | b | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Protichneumon grandis | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | e | 0 | 0 | 0 | 0 | 0 | 0 | 0 | a | a | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | 1 |
Barichneumon neosorex | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Cratichneumon w-album | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Orgichneumon calcatorius | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | b | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Patrocloides montanus | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Plagiotrypes concinnus | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Dilopharius otomitus | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Cyclolabus impressus | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | a | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 1 |
Linycus exhortator | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 1 |
Grotea anguina | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 2 | 3 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 7 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Labium sp. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | b | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 7 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Apechoneura sp. | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 3 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Labena grallator | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | a | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 7 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Poecilocryptus nigromaculatus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 7 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Lycorina glaucomata | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 3 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | ? | 0 | 1 | 3 | ? | c | 0 | 1 | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Astiphromma sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | h | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | a | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Chineater masneri | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 |
Cidaphus paniscoides | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 2 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 |
Lepidura collaris | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 2 | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 |
Mesochorus sp. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 |
Exochus semirufus | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Metopius pollinctorius | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 3 | 0 | d | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 3 | ? | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 7 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Scolomus sp. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 2 | 3 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | ? | 0 | 2 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Seticornuta terminalis | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 2 | 3 | 0 | 2 | 2 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 3 | ? | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Microleptes sp. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | d | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 |
Neorhacodes enslini | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 2 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | 4 | ? | ? | ? | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Chriodes sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Nonnus sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 1 | 1 | 3 | ? | 0 | 1 | 1 | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 7 | 0 | 0 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Enicospilus flavostigma | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 3 | ? | ? | 2 | 0 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | ? | 1 | 0 | 1 | 2 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Hellwigia obscura | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 2 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | ? | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | ? | 1 | 0 | 0 | 2 | 2 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | 1 |
Ophion sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 2 | 0 | ? | 2 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | ? | 1 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Skiapus sp. | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 2 | 3 | 0 | 1 | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 1 | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 0 | ? | 1 | 0 | 0 | 2 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | 1 |
Thyreodon sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | ? | 1 | 0 | 0 | 2 | 1 | 1 | 1 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Megastylus sp. nov. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 3 | 1 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Orthocentrus sp. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | ? | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Proclitus speciosus | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | ? | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | n | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Orthopelma mediator | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 3 | 0 | ? | 1 | 1 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Oxytorus albopleuralis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Pedunculus sp. nov. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 5 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 1 |
Perithous divinator | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | ? | 0 | 1 | 1 | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Acrotaphus wiltii | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | ? | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 3 | 0 | 0 | 0 | 1 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Clistopyga recurva | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Dolichomitus irritator | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 3 | ? | 3 | ? | ? | b | b | 2 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Zaglyptus pictilis | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Pimpla annulipes | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Theronia bicincta | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Neoxorides caryae | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 3 | ? | 0 | 0 | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Poemenia albipes | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 2 | 0 | 4 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | a | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Megarhyssa greenei | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Rhyssa crevieri | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Rhyssella nitida | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Brachyscleroma sp. | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | a | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 |
Erythrodolius calamitosus | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 1 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Notostilbops sp. nov. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Stilbops vetulus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | a | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Tatogaster nigra | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | ? | ? | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 2 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Allophrys sp. | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 3 | 1 | ? | 1 | 1 | 0 | 2 | 1 | 1 | 2 | 1 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 |
Peucobius fulvus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | a | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Phrudus sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 0 | 1 | 2 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 |
Stethantyx nearctica | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 1 | 1 | 2 | 1 | 1 | 0 | a | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 |
Tersilochus sp. | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 3 | 1 | ? | 1 | 1 | 0 | 2 | 1 | 1 | 2 | 1 | 1 | 2 | a | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 |
Eclytus sp. | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 3 | ? | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Idiogramma longicauda | 0 | 0 | 0 | 1 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | ? | 3 | ? | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Zagryphus nasutus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 |
Netelia sp. | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 |
Phytodietus vulgaris | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | b | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 1 | 3 | ? | 3 | ? | ? | 2 | 2 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 |
Cteniscus sp. | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 |
Cycasis rubiginosa | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 |
Polyblastus sp. | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 |
Aplomerus sp. | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | ? | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 2 | 2 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Odontocolon albotibiale | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
Xorides stigmapterus | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 2 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 |
(Continued) Data matrix for morphological character states of Ichneumonidae and outgroup taxa. Key to the letters in matrix below: a=0/1; b=0/2; c=0/3; d=1/2; e=1/3; f=1/4; g=1/5; h=2/3; k=2/4; m=2/6; n=3/4; p=3/7; q=5/6; r=6/7; s=0/1/3; t=1/5/6; u=2/6/7; v=5/6/7; w=2/5/6/7; x=3/5/6/7.
Taxon | 1 | 1 | 1 | 1 | 1 | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|---|
8 | 9 | 0 | 1 | 2 | 3 | 4 | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 0 | 1 | |
Doryctes erythromelas | 0 | 1 | 2 | ? | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 2 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | ? | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 1 |
Rhysipolis sp. | 0 | 0 | 2 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 3 | 1 | 0 | 0 | 1 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | ? | ? | ? | ? | ? | ? | 0 | 0 | 0 | 1 | 0 | 2 | 1 | 5 |
Aleiodes terminalis | 0 | 0 | 2 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | 5 |
Spilopteron occiputale | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | ? | ? | 0 | 0 | 0 | ? | ? | ? | 3 | 0 | ? |
Coleocentrus rufus | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 3 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 0 | 0 | 1 | 1 | 0 | 0 | 0 | ? | ? | ? | 3 | 0 | ? |
Adelognathus sp. | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | k | 1 |
Agriotypus armatus | 1 | 2 | 2 | ? | 0 | 0 | 0 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 6 | 6 | 2 |
Anomalon picticorne | 1 | 2 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | ? | ? | 0 | 1 | 1 | h | k | 6 |
Therion texanum | 1 | 2 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 2 | 0 | 0 | 1 | 1 | 2 | 4 | 6 |
Lissonota scutellaris | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 2 | 5 |
Exetastes bioculatus | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | k | 5 |
Apophua simplicipes | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 2 | 5 |
Sphelodon phoxopteridis | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | 1 | 1 | 2 | 2 | 5 |
Brachycyrtus wardae | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 1 | ? | ? | ? | ? | 0 | 7 | ? | w |
Bathyplectes infernalis | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 3 | 4 | q |
Campoletis sonorensis | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | 1 |
Campoplex sp. | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 2 | g |
Casinaria grandis | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | 1 |
Dusona egregia | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | q |
Hyposoter sp. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | 1 |
Olesicampe sp. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 4 | q |
Rhimphoctona macrocephala | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | ? | ? | 0 | 1 | 1 | 3 | 0 | g |
Clasis sp. nov. | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 3 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Collyria catoptron | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 4 | 0 | 0 | 0 | ? | 2 | 5 | 2 | 0 | ? | 0 | ? | ? | ? | 0 | 0 | ? | 0 | 0 | 2 | 1 | ? | 0 | 0 | 0 | 1 | 1 | 1 | 1 | 3 |
Eiphosoma pyralidis | 1 | 2 | 2 | ? | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 2 | 5 |
Xiphosomella setoni | 0 | 1 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | ? | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 1 | 1 | ? | ? | ? | ? | ? | 2 | 2 | ? |
Agonocryptus chichimecus | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | ? | ? | 3 | a | ? |
Ateleute sp. nov. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | 2 | 2 | ? |
Baryceros texanus | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 2 | 4 | ? |
Diapetimorpha brunnea | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | ? | 2 | 4 | 2 |
Echthrus reluctator | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 3 | 0 | ? |
Lymeon orbus | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | ? | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | ? | ? | ? | ? | d | ? | ? |
Polytribax contiguus | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 0 | 2 | ? | m |
Rhytura pendens | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Acrolyta sp. | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | ? | 0 | 0 | 0 | 1 | 4 | 2 |
Endasys patulus | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 4 | 2 |
Mastrus sp. | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | d | f | 2 |
Ctenopelma sanguineum | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | h | p |
Xenoschesis limata | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 3 | t |
Euryproctus sentinis | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 4 | ? |
Mesoleptidea decens | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | ? | ? | ? | 1 | 1 | 1 | 4 | ? |
Barytarbes honestus | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | 0 | 0 | ? | ? | 1 | 4 | ? |
Himerta luteofacia | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 4 | ? |
Lathrolestes asperatus | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | d | 1 | q |
Perilissus concolor | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 4 | ? |
Rhorus bartelti | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | ? | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 4 | 6 |
Sympherta fucata | 1 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 4 | ? |
Seleucus cuneiformis | 0 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 1 | 2 | ? |
Onarion sp. | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Westwoodia sp. | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | 0 | 0 | ? | 1 | 1 | 4 | t |
Ctenopelmatinae Genus NZ | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | 0 | ? | ? | 1 | 2 | ? | u |
Cylloceria melancholica | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | ? | ? | ? | ? | 1 | 4 | ? | ? |
Diacritus incompletus | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Diplazon laetatorius | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 1 | 3 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 2 | ? | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 4 | 4 | 7 |
Woldstedtius flavolineatus | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | ? | 0 | 0 | 1 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 2 | ? | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 2 | ? | ? | ? | ? | ? | ? | ? | 4 | 4 | ? |
Euceros sp. nov. | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 4 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 3 | 2 | 0 | 1 | 1 | 1 | 8 | 8 |
Hybrizon rileyi | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 1 | 1 | 4 | q |
Alomya debellator | 0 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | ? | 2 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 1 | ? | ? | 0 | 1 | 1 | 2 | 4 | 5 |
Centeterus euryptychiae | 1 | 1 | 2 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 0 | 0 | 2 | ? | 1 | 1 | ? | ? | ? | ? | 1 | 2 | ? | ? |
Phaeogenes hebrus | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 0 | 0 | 2 | 1 | 1 | 1 | 0 | 0 | 0 | ? | 1 | 2 | k | u |
Stenodontus sp. nov. | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 2 | ? | ? |
Coelichneumon eximius | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | 0 | 1 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 1 | 0 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | ? | 1 | 2 | 4 | u |
Protichneumon grandis | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 1 | 0 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | ? | 1 | 2 | 4 | u |
Barichneumon neosorex | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 0 | 0 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | ? | 1 | 2 | 4 | u |
Cratichneumon w-album | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 0 | 0 | 1 | ? | 1 | 1 | 0 | 0 | 0 | 0 | 1 | 2 | 4 | 2 |
Orgichneumon calcatorius | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 3 | 0 | 1 | 0 | 0 | ? | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | ? | 0 | 1 | ? | 1 | 1 | ? | ? | 0 | 1 | 1 | 2 | 4 | r |
Patrocloides montanus | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | ? | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | ? | 0 | 1 | ? | 1 | 1 | ? | ? | 0 | 1 | 1 | 2 | 4 | r |
Plagiotrypes concinnus | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Dilopharius otomitus | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 3 | ? | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Cyclolabus impressus | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 0 | 0 | 1 | ? | 1 | 1 | 0 | 0 | 0 | 1 | 1 | ? | 4 | 6 |
Linycus exhortator | 1 | 1 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | 0 | 0 | 0 | 2 | 2 | ? | ? | 0 | 0 | 0 | 1 | ? | 1 | 1 | ? | ? | 0 | ? | 1 | 2 | 4 | ? |
Grotea anguina | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 |
Labium sp. | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 4 | 0 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | ? | ? | ? | ? | 0 | 1 | 5 | ? |
Apechoneura sp. | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 0 | ? | ? | ? | 0 | ? |
Labena grallator | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | ? | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | 3 | 0 | 1 |
Poecilocryptus nigromaculatus | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | ? | 2 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 1 | 0 | ? | ? | ? | 0 | 0 | 0 | 1 | ? |
Lycorina glaucomata | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 2 | 0 | 0 | 3 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 1 | 1 | 1 | 0 | 1 | 2 | 2 | 4 | 6 |
Astiphromma sp. nov. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 1 | 1 | ? | ? | 0 | 1 | 1 | 1 | 9 | v |
Chineater masneri | 0 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Cidaphus paniscoides | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | ? | ? | 0 | 1 | 1 | 1 | 9 | v |
Lepidura collaris | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Mesochorus sp. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 1 | 9 | 5 |
Exochus semirufus | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 0 | 0 | 0 | 0 | ? | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | ? | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | k | 6 |
Metopius pollinctorius | 0 | 0 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 2 | 1 | 0 | 0 | 0 | 1 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | ? | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | 6 |
Scolomus sp. | 0 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 2 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Seticornuta terminalis | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | ? | ? | 0 | 1 | 1 | 2 | 2 | u |
Microleptes sp. | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 1 | ? | ? | ? | ? | 1 | 4 | ? | ? |
Neorhacodes enslini | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | ? | 0 | 0 | ? | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 1 | ? | ? | ? | ? | 1 | 1 | ? | ? |
Chriodes sp. | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Nonnus sp. | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Enicospilus flavostigma | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 2 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | 4 | q |
Hellwigia obscura | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 1 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 2 | ? | ? |
Ophion sp. | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 2 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | h | 4 | q |
Skiapus sp. | 1 | 2 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Thyreodon sp. | 1 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 2 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | 2 | n | 5 |
Megastylus sp. nov. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 0 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 1 | 4 | ? | ? | 4 | ? | 0 | 1 | 0 | 0 | 3 | 2 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | ? | ? | 0 | 1 | 1 | 4 | f | x |
Orthocentrus sp. | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 2 | 0 | 0 | 2 | 0 | 0 | 3 | 2 | ? | 0 | 0 | ? | 1 | 0 | ? | 0 | ? | 0 | 0 | ? | 0 | 0 | ? | ? | ? | ? | 1 | 4 | ? | ? |
Proclitus speciosus | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 4 | ? | ? |
Orthopelma mediator | 1 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 0 | ? | 2 | ? | 2 | ? | 0 | 0 | ? | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 0 | ? | ? | 0 | ? | 1 | 1 | 1 | ? |
Oxytorus albopleuralis | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Pedunculus sp. nov. | 1 | 2 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 3 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Perithous divinator | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 2 |
Acrotaphus wiltii | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | 1 | 0 | 5 | 4 | 4 |
Clistopyga recurva | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | 0 | 0 | 8 | 3 | 0 |
Dolichomitus irritator | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 1 |
Zaglyptus pictilis | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | 0 | 0 | 0 | 8 | 3 | 0 |
Pimpla annulipes | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 2 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 4 | 2 |
Theronia bicincta | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 2 | 4 | 2 |
Neoxorides caryae | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | ? | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 4 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | 0 | 0 | 0 | e | 0 | 1 |
Poemenia albipes | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | ? | ? | 0 | 0 | 0 | 1 | a | 1 |
Megarhyssa greenei | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 2 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 |
Rhyssa crevieri | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 |
Rhyssella nitida | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | ? | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 |
Brachyscleroma sp. | 0 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 3 | 2 | ? |
Erythrodolius calamitosus | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Notostilbops sp. nov. | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Stilbops vetulus | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 1 | ? | ? | 0 | 1 | 1 | 2 | 2 | 7 |
Tatogaster nigra | 1 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Allophrys divaricata | 1 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 3 | 1 | ? |
Peucobius fulvus | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 0 | ? | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Phrudus sp. | 1 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | 0 | ? | 1 | 0 | 0 | 2 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | ? | ? | ? | 1 | ? | ? | ? | ? | ? | 3 | ? | ? |
Stethantyx nearctica | 0 | 1 | 0 | 1 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 1 | 1 | ? | ? | 0 | 1 | 1 | 3 | 1 | 6 |
Tersilochus sp. | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 2 | 1 | 0 | 1 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 1 | h | 1 | 6 |
Eclytus sp. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 4 | 6 |
Idiogramma longicauda | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | ? | 0 | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 1 | 6 |
Zagryphus nasutus | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 1 | ? | 1 | ? | ? | ? | ? | ? |
Netelia sp. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 1 | 1 | 0 | 1 | 1 | 0 | 2 | 4 | 6 |
Phytodietus vulgaris | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 2 | 0 | 0 | 3 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 1 | 0 | 1 | 1 | 0 | 2 | 2 | 6 |
Cteniscus sp. | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | ? | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 4 | 6 |
Cycasis rubiginosa | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | 1 | 1 | 1 | ? | ? | 1 | 4 | ? |
Polyblastus sp. | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 1 | 1 | 1 | 0 | 1 | 4 | 6 |
Aplomerus sp. | 0 | 0 | 2 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? | ? |
Odontocolon albotibiale | 1 | 1 | 2 | ? | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 1 |
Xorides stigmapterus | 0 | 1 | 2 | ? | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 2 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 1 | 0 | 1 | 1 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 0 | 1 | 0 | 0 | 0 | 0 | 0 | 3 | 0 | 1 |
Voucher specimens and larval slides are deposited at the Australian National Insect Collection, Canberra, Australia (
We used a species exemplar approach in this study, rather than coding morphological characters at the tribal or subfamily level as has been done in other studies on family and subfamily relationships within Hymenoptera (
All terms of ichneumonid morphology follow
The carinae on the propodeum (see Figs
1) In cases in which there is ambiguity between the anterior and posterior transverse carinae, the posterior transverse carina was given precedence.
2) In cases in which there is ambiguity between the medial and lateral longitudinal carinae, the medial longitudinal carinae were given precedence.
3) In cases in which longitudinal and transverse carinae run obliquely together across the propodeum, the transverse carina was given precedence over the longitudinal carinae.
4) In cases in which an abscissa of a carina is incomplete (e.g., the costula of many specimens of Campopleginae), the abscissa was considered present if it extended half the way across the area in which it is located or greater, or if this could not be determined (e.g., no other carinae are present on the propodeum), one sixth of the length (for longitudinal carinae) or width (for tranverse carinae) of the propodeum.
Digital photos at the CNC were taken using a Leica MZ16 stereomicroscope with motorized focus drive attached to a Leica DFC420 digital camera. Photos were combined and edited using Leica Application Suites Montage Multifocus software V3.8, Auto-Montage Pro 5.01 and Adobe Photoshop CS4. Photos taken at EMUS were taken with an EntoVision micro-imaging system. This system consists of a Leica M16 zoom lens attached to a JVC KY-75U 3-CCD digital video camera that feeds image data to a desktop computer. The program Archimed 5.3.1 was used to merge an image series (representing typically 15–30 focal planes) into a single in-focus image. Lighting was provided by an EntoVision dome light.
Most sequences in this study were obtained by sequencing specimens at the Canadian National Collection of Insects, Arachnids and Nematodes (CNCI), except as noted in Appendix
Molecular extracts were obtained using standard protocols outlined in DNeasy Blood and Tissue extraction kit instructions (Qiagen, Gaithersburg, MD, USA). A single mid leg was macerated except for small specimens in which a hind and mid leg were used. PCRs were carried out using 50 μl reactions containing 4 μl DNA extract, 1 μl of each primer, 5 μl PCR buffer, 1 μl DNTPs, 1 μl MgCl2, 0.25 μl Taq and 36.75 μl RNase-free water. Primers were as follows: Cytochrome oxidase I (COI) (F) (LCO1490): GGTCAACAAATCATAA AGATATTGG; (R) (HCO2198): TAAACTTCAGGGTGACCAAAAAATCA (
Preliminary alignment of coding genes (COI and EF1-a) was done using ClustalW (
Aligned sequences are available in Suppl. materials
Parsimony analysis was performed using TNT v. 1.1 (
Bayesian analyses were conducted using MrBayes 3.1.2 (
In total, 141 morphological and biological characters were included in the analyses: 104 adult, 28 larval, 3 egg and 5 biological.
1. Clypeus: 0) comprised of one part (Fig.
2. Clypeus and supraclypeal area: 0) separated by groove or indentation (Fig.
3. Clypeal margin in anterior view: 0) simple, truncate to slightly convex or slightly concave (Fig.
4. Clypeal margin vestiture: 0) with small scattered setae or lacking setae; 1) with regularly spaced strong setae (Fig.
5. Mandibles: 0) bidentate, gradually to strongly tapering (Fig.
6. Subocular sulcus: 0) absent to indistinct (Fig.
7. Apical flagellomere of female: 0) simple; 1) with projections arising at outgrowth from surface (see
8. Antennal color of female: 0) more or less unicolorous; 1) with distinct light coloured median band.
9. Flagellum of male: 0) central flagellomeres lacking tyloids; 1) central flagellomeres with elliptical (Fig.
10. Inner margin of eye: 0) not or only weakly emarginate (Figs
11. Lateral ocellus: 0) small, separated from eye by 0.5× its diameter or greater; 1) enlarged, touching, or almost touching eye.
12. Gena: 0) simple; 1) denticulate (Fig.
13. Occiput with medial notch near foramen magnum: 0) absent; 1) present (see
14. Occipital carina: (0) joining hypostomal carina at a distance from mandible that is less than basal width of mandible (Fig.
15. Foramen magnum: 0) simple (Fig.
16. Maxillary palpus: 0) 5-segmented; 1) 4-segmented; 2) 3-segmented.
17. Maxillary palpus: 0) normal; 1) elongate, reaching to or beyond middle coxa.
18. Labial palpus: 0) 4-segmented; 1) 3-segmented.
19. Propleuron: 0) without lateroventral posteriorly-projecting lobe (Fig.
20. Epomia: 0) present as ridge ventro-anteriorly that crosses furrow dorsally (Fig.
21. Mesoscutum: 0) smooth (Fig.
22. Notaulus: 0) shallow or vestigial; 1) deeply impressed anteriorly (Fig.
23. Notaulus: 0) extending to 0.5× length of mesoscutum or less; 1) extending posteriorly past centre of mesoscutum but not joining other notaulus; 2) extending posteriorly past centre and joining either other notaulus or rugose medial area (Fig.
24. Notaular crest or carina anterolateral to notaulus: 0) absent; 1) present (arrow in Fig.
25. Epicnemial carina: 0) not curving anteriorly, vertical to around middle of pronotum; 1) curving to anterior of mesopleuron, around middle of pronotum (arrow in Fig.
26. Sternaulus length: 0) present, less than 0.7× length of mesopleuron; 1) present, greater than or equal to 0.7× length of mesopleuron (Figs
27. Sternaulus curvature: 0) short, not reaching posterior end of mesopleuron (posterior curvature not scoreable); 1) posterior end ending dorsal to posterolateral corner of mesopleuron (Fig.
28. Foveate groove of mesopleuron: 0) absent; 1) present (arrow in Fig.
29. Mesopleural groove: 0) absent to incomplete; 1) complete to posterior margin of mesopleuron (“mg” in Fig.
30. Posterior transverse carina of mesothoracic venter: 0) absent; 1) interrupted near anterior of middle coxa (gap indicated by arrow in Fig.
31. Metapostnotum: 0) posterolateral triangles present (Figs
32. Propodeum base: 0) without median tubercle; 1) with small, median tubercle (arrow in Fig.
33. Propodeal spiracles: 0) separated from pleural carina by about minimum diameter of spiracle or more (Fig.
34. Propodeal spiracles: 0) round to sub-circular (less than 1.5× as long as high) (Figs
35. Lateral profile of propodeum: 0) angulate with separate dorsal and posterior faces (Fig.
36. Anterior transverse carina of propodeum: 0) complete (medially and sublaterally) (Fig.
37. Anterior transverse carina of propodeum: 0) medially angled (Fig.
38. Posterior transverse carina of propodeum: 0) complete (medially and sublaterally) (Figs
39. Posterior transverse carina of propodeum: 0) as strongly developed as other carinae; 1) conspicuously stronger than other carinae (Fig.
40. Posterior transverse carina: 0) angled (Fig.
41. Anterior abscissa of medial longitudinal carina: 0) present, not fused (Fig.
42. Median abscissa of medial longitudinal carina: 0) present, not fused (Fig.
43. Posterior abscissa of medial longitudinal carina: 0) present, not fused (Fig.
44. Anterior abscissa of lateral longitudinal carina: 0) present (Fig.
45. Median abscissa of lateral longitudinal carina: 0) present (Fig.
46. Posterior abscissa of lateral longitudinal carina: 0) present (Fig.
47. Propodeal surface reticulation: 0) smooth, without reticulation (Fig.
48. Submetapleural carina: 0) complete, anterior section unmodified to slightly broadened (arrow in Fig.
49. Height of ventral edge of metasomal foramen (“mf” in Figs
50. Metacoxal cavity posteromedially: 0) separated from metasomal insertion by sclerotized bridge (Fig.
51. Fore wing vein 2m-cu: 0) present (Fig.
52. Fore wing vein 2m-cu posteriorly: 0) vertical (joining vein Cu at right angle) to slightly reclivous (Fig.
53. Fore wing vein 2m-cu: 0) with two discrete bullae separated by small length of tubular vein (arrows in Fig.
54. Fore wing cell 1+2Rs (areolet): 0) obliquely rhombic to subtriangular (Fig.
55. Fore wing cell 1+2Rs (areolet) veins: 0) smilar thickness to other fore wing veins (Fig.
56. Fore wing cell 1+2Rs (areolet): 0) sessile anteriorly (Figs
57. Antero-medial fore wing flexion line basally: 0) splitting anterior to M and anterior fold running anterior to M (forming bulla indicated by arrow) (Fig.
58. Fore wing cell 2R1 (radial cell) posterior angle: 0) greater than 100 degrees (Fig.
59. Fore wing cell 1M+1R1: 0) uniformly hirsute; 1) with small to large glabrous area that may have free sclerites (Fig.
60. Fore wing vein Rs+M (ramellus): 0) complete (extending across all of cell 1M + 1R1) (Fig.
61. Fore wing vein 1cu-a: 0) opposite, slightly proximal or slightly distad vein M (Fig.
62. Hind wing vein 1rs-m: 0) basal to separation of veins R1 and Rs (Fig.
63. Hind wing vein 2/Cu: 0) equidistant between A and M, closer to A or slightly closer to M (Fig.
64. Hind wing vein M+Cu: 0) complete (Fig.
65. Hind wing vein M+Cu: 0) straight to weakly arched (Fig.
66. Hind wing basal hamuli: 0) distant from wing base on membrane or spectral vein; 1) close to wing base on spur of tubular vein; 2) absent.
67. Hind wing distal hamuli: 0) 1–3; 1) 4 or greater.
68. Apex of fore tibia: 0) without tooth; 1) with distinct tooth on dorsal margin (Fig.
69. Fore tibia dorsal surface: 0) covered with uniform thickness of setae; 1) with uniform setae and sparse, much stouter spines (Fig.
70. Middle tibial spurs: 0) two; 1) one.
71. Apex of middle and hind tibiae: 0) with common area of insertion for spurs and basitarsus; 1) with sclerotized bridge separating insertion areas (Fig.
72. Female hind coxa: 0) simple, without furrow; 1) with inner surface near base with vertical basal furrow (Fig.
73. Hind tibia: 0) covered with uniform thickness of setae; 1) with uniform setae and sparse, much stouter spines (Fig.
74. Hind tibia: 0) with posterior face simple (not smooth and shining) and with moderately sparse fringe of setae (Fig.
75. Hind tibial spurs: 0) present; 1) absent.
76. T1 petiolar cross-section (measured at petiolar midpoint): 0) not petiolate or if petiolate not wider than high (height/ width = 1–1.2×); 1) wider than high (height/ width = 0.6–0.7×).
77. T1 spiracle location: 0) at or anterior to 0.6× length of segment; 1) posterior to 0.6× length of segment.
78. T1 glymma: 0) present and shallow (Fig.
79. First metasomal segment shape: 0) non-petiolate: more or less evenly broadened from near base to posterior end or, if more or less parallel-sided throughout, then clearly depressed and less than 2.0× as long as posteriorly wide (Fig.
80. S1 length: 0) 0.6× length of T1 or less; 1) longer than 0.6× length of T1.
81. S1 fusion apically: 0) not fused to T1 (Fig.
82. Thyridium shape: 0) present, ovoid (Fig.
83. Thyridium location: 0) less than one length or diameter distant from anterior edge of T1 (Fig.
84. Gastrocoelus: 0) absent; 1) present (“gs” in Fig.
85. Pseudothyridium of T2: 0) present (“ps” in Fig.
86. T2 sculpture: 0) smooth to granulate; 1) with clearly defined longitudinal rugulae.
87. T2 and T3 fusion: 0) separate, with flexion line allowing movement (Fig.
88. MS2 laterotergites: 0) creased and curved mesad under metasoma (“L2” in Fig.
89. MS3 laterotergite: 0) separated by crease, often partially turned under (“L3” in Fig.
90. MS4 laterotergite: 0) separated by crease (at least basally), often turned under; 1) not separated by crease (Fig.
91. T2–T4 sculpture: 0) each tergite uniformly sculptured; 1) posterior 0.2 of each tergite sculptured differently than anterior 0.8 (arrows in Fig.
92. Apical segment of female metasoma: 0) short; 1) elongate with horn or boss (“h” in Fig.
93. Posterior sternites of female: 0) without ovipositor guides; 1) with tuberculate ovipositor guides (arrows in Fig.
94. Female hypopygium in lateral profile: 0) inconspicuous and uniformly sclerotized; 1) moderate length, regularly triangular in profile, uniformly sclerotized (Fig.
95. Hind margin of female hypopygium: 0) simple; 1) with median apical notch (arrow in Fig.
96. Ovipositor length: 0) longer than apical height of metasoma but shorter than metasoma; 1) shorter than or equal to apical height of metasoma; 2) longer than length of metasoma.
97. Ovipositor ventral valve: 0) with teeth apically (Fig.
98. Ovipositor ventral valve: 0) not enclosing dorsal valve; 1) enclosing dorsal valve.
99. Ovipositor dorsal valve, apically: 0) simply tapered (Fig.
100. T8 or T8+9 of male: 0) medially longitudinally divided (Fig.
101. Hypopygium of male: 0) short and apically truncate; 1) elongate and scoop-like (Fig.
102. Gonoforceps: 0) simple (“gon” in Fig.
103. Apical 0.5 of aedeagus: 0) subcylindrical and slightly to strongly clubbed (Fig.
104. Ovaries: 0) bohrertypus of
105. Larval epistomal suture: 0) unsclerotized (Fig.
106. Larval hypostoma and pleurostoma: 0) not laterally expanded (Fig.
107. Larval pleurostoma location: 0) inferior mandibular process dorsad or opposite to dorsal margin of labial sclerite (Fig.
108. Larval labral sclerite: 0) present (Figs
109. Larval mandible shape: 0) triangular, auxiliary tooth present or absent (Figs
110. Larval mandible with accessory teeth: 0) absent (Fig.
111. Larval mandible sclerotization: 0) uniformly sclerotized (Fig.
112. Larval mandibular blade denticles: 0) present on entire dorsal and ventral margins (Fig.
113. Larval mandible, spines at base of blades: 0) juncture of base and blade without long, horizontal spines (Figs
114. Larval posterior struts of inferior mandibular processes: 0) short (as long as dorsal struts) and not connected by band (Fig.
115. Larval hypostoma length: 0) long (>2 × as long as hypostomal spur) (Fig.
116. Larval hypostoma, lateral end: 0) simple/ undivided (Fig.
117. Larval hypostomal spur: 0) normal, about 2× as long as basal width or longer (Fig.
118. Larval hypostomal spur meeting stipital sclerite: 0) near middle to lateral end of stipital sclerite (Fig.
119. Larval stipital sclerite orientation: 0) oriented so that lateral end at about right angle to labial sclerite (Figs
120. Larval stipital sclerite lateral end: 0) unmodified (Fig.
121. Larval cardo: 0) unsclerotized (Fig.
122. Larval maxillary apex: 0) unsclerotized (Fig.
123. Larval maxillary and labial palpi sensilla: 0) maxillary bearing one to two, labial bearing one to three (Fig.
124. Larval labial sclerite shape: 0) quadrate (Fig.
125. Larval labial sclerite dimensions: 0) about as long as wide, length of ventral portion/ total length = 0.2–0.3 (Fig.
126. Larval labial sclerite ventral margin: 0) relatively unmodified (may be lobes, minor scalloping, etc.) (Fig.
127. Larval prelabial sclerite: 0) absent (Fig.
128. Larval prelabium, number of sensilla: 0) 6 (Fig.
129. Larval sclerotized plate ventral to labial sclerite: 0) absent (Fig.
130. Larval clypeolabral plate location: 0) absent (Fig.
131. Larval antenna: 0) present and with central papillus present (Figs
132. Larval spiracles: 0) present, closing apparatus separated from atrium by section of trachea (Fig.
133. Larval salivary orifice: 0) transverse or ovoid (Figs
134. Egg: 0) without stalk; 1) with chorionic stalk (Fig.
135. Egg stalk anchor: 0) anchor absent or entire stalk absent; 1) with tryphonine-like anchor (Fig.
136. Exit of egg from body: 0) egg travels down lumen of ovipositor; 1) egg exits from hole ventral to ovipositor, stalk goes down ovipositor; 2) entire egg exits from hole ventral to ovipositor.
137. Biological mode (timing of larval maturation): 0) idiobiont; 1) koinobiont.
138. Biological mode (location of development): 0) ectoparasitoid; 1) endoparasitoid; 2) inside hind gut with final ectoparasitoid phase AND pupate inside host cocoon.
139. Host/ source of larval nutrition: 0) phytophagous (at least facultatively after consuming insect host); 1) Hymenoptera; 2) Lepidoptera; 3) Coleoptera; 4) Diptera; 5) Araeneae; 6) Trichoptera; 7) Neuroptera; 8) egg predators.
140. Oviposition location: 0) through lignified plant tissue; 1) through non-lignified plant or gall tissue; 2) in leaf rolls, cases and other plant tissues held by silk; 3) in silken bags or sacs; 4) into or onto exposed larval/ pupal hosts; 5) into ground nests; 6) into trichopteran tubes; 7) into exposed eggs; 8) on to leaf; 9) into host in host.
141. Oviposition/emergence stages: 0) eggs (predators); 1) larval-larval (not last instar); 2) pupa-pupa; 3) egg-larval; 4) immature spider - adult spider; 5) larval-larval (last instar); 6) larval-pupal; 7) egg-pupal; 8) leaf-pupa.
Total-evidence
The total-evidence parsimony analysis found 1728 equally parsimonious trees of length 9917 (C.I. = 0.15, R.I. = 0.44). In the strict consensus tree (Fig.
Comparison of the monophyly of selected taxa with different phylogenetic analyses and datasets. See footnotes and introduction for composition of selected taxa. “Yes” indicates that all taxa were supported in all equally parsimonious trees or had a support value of 100 in the Bayesian majority credibility tree. “No” indicates lack of monophyly in at least one most parsimonious tree or a support value of less than 100 in the Bayesian analysis. Pars. = parsimony; all = all characters; morph. = morphological and biological characters only; mol. = molecular characters only; Bayes = Bayesian analysis.
Taxon | Pars. (all) | Pars. (morph.) | Pars. (mol.) | Bayes (all) | Bayes (mol.) |
---|---|---|---|---|---|
Pimpliformes | Yes | No | Yes | Yes | Yes |
Ichneumoniformes | No1 | No | No | No | No |
Ophioniformes | Yes | No | No | No | No |
Acaenitinae | No | Yes | No | No | No |
Anomaloninae | Yes | Yes | No | No | No |
Banchinae | No2 | Yes | No2 | No2 | No2 |
Campopleginae | Yes | No | Yes | Yes | Yes |
Cremastinae | Yes | Yes | Yes | Yes | Yes |
Cryptinae | Yes | No | No | Yes | Yes |
Ctenopelmatinae | No3 | No | No | No | No |
Diplazontinae | Yes | Yes | Yes | Yes | Yes |
Ichneumoninae | Yes | Yes | Yes9 | Yes | Yes9 |
Labeninae | Yes | No | Yes | Yes | Yes |
Mesochorinae | No4 | Yes | No | No4 | No4 |
Metopiinae | No5 | Yes | No5 | No5 | No5 |
Nesomesochorinae | Yes | No | No | No | No |
Ophioninae | Yes | Yes | No | Yes | No |
Orthocentrinae | No | Yes | No | No | No |
Phrudus group | No6 | No | No6 | No6 | No6 |
Phygadeuontinae | Yes | Yes | No | No | No |
Pimplinae | No | No | No | No | No |
Poemeniinae | Yes | Yes | Yes | Yes | No |
Rhyssinae | Yes | Yes | Yes | Yes | Yes |
Sisyrostolinae | No | No | No | No | No |
Stilbopinae | No | No | No | No | No |
Tersilochinae s.l. | No7 | No | No | No7 | No |
Tersilochinae s.s. | Yes | Yes | Yes | Yes | Yes |
Tryphoninae | No8 | No | No | No | No |
Xoridinae | No | Yes | No | Yes | No |
14–15 Head, anterior view 14 Skiapus sp 15 Hybrizon rileyi Arrow indicates reduced mandible 16–17 Head, lateral view 16 Diplazon laetatorius 17 Orthocentrus sp. Arrow indicates subocular groove 18–20 Antennae 18 Labena grallator, apical flagellomere 19–20 Flagellum, lateral view, arrows indicate longitudinal tyloids 19 Protichneumon grandis 20 Lymeon orbus (Say).
21 Podoschistus vittifrons (Cresson), head, lateral view 22 Venturia sokanakiakorum (Viereck), head and mesosoma, lateral view, showing ventral lobe of propleuron 23 Labena grallator, pronotum, lateral view. Arrow indicates epomia 24 Rhyssella perfulva Porter, head and mesosoma, lateral view 25–26 Diacritus incompletus 25 Mesosoma, dorsal view 26 Mesosoma, lateral view. Arrow indicates notaular crest.
27–30 Mesosoma, lateral view 27 Rhimphoctona macrocephala. ec = epicnemial carina 28 Diapetimorpha brunnea Townes. Arrow indicates sternaulus curving ventrally anterior to posterolateral corner 29 Stethantyx nearctica. Arrow indicates foveate groove 30 Agriotypus armatus. mg = mesopleural groove, st = sternaulus 31–32 Mesosternum, ventral view 31 Therion longipes (Provancher). Arrow indicates incomplete posterior transverse carina 32 Dusona egregia. Arrow indicates complete posterior transverse carina.
Propodeum. 33 Polytribax contiguous (Cresson), dorsolateral view. Arrow indicates posterolateral triangle of metanotum 34 Centeterus euryptychiae, dorsal view. Arrow indicates median tubercle at base of propodeum 35 Sphelodon phoxopteridis, lateral view 36–38 Dorsal view (re-drawn after
39–42 Propodeum, dorsal view 39 Pyracmon hyalinus (re-drawn after
45 Dolichomitus irritator, mesopleuron and metapleuron, lateral view. Arrow indicates submetapleural carina 46–48 Mesosoma and metasoma, ventroposterior view. Line above cf = dorsal edge of coxal foramen, line below mf = ventral edge of metasomal foramen 46 Lissonota scutellaris 47 Apechoneura sp. 48 Polyblastus pedalis (Cresson). Arrow in 48 indicates unsclerotized region joining metasomal and coxal foramina 49 Wings, Helcon sp. (Braconidae) (modified from
62 Ateleute tsiriria, hind wing. 63–64 Fore tibia, lateral view 63 Euryproctus sentinis Davis (apex with distinct tooth) 64 Phytodietus vulgaris. Arrows indicate sparse, stout spines 65 Eiphosoma pyralidis, hind tibia, apical view. Arrow indicates sclerotized bridge between insertion points of spurs and basitarsus 66 Labena grallator, inner surface of hind coxa of female, lateral view. Arrow indicates furrow for bracing ovipositor during oviposition 67 Phytodietus vulgaris, lateral view. Arrows indicate sparse, stout spines.
68–69 Hind tibial apex, apical view. sp = spur, tar = tarsus, tib apex = tibial apex 68 Polyblastus pedalis (simple apex with sparse fringe of stout setae) 69 Pedunculus sp. (apex with apical face smooth and enlarged with thick fringe of fine setae) 70–71 Tergite 1 of metasoma, lateral view 70 Phytodietus vulgaris Arrow points to glymma 71 Netelia sp. Arrow points to deep glymma (sides of glymma separated medially by only a thin, translucent sclerite) 72–73 Tergite 1, dorsal view 72 Pimpla aequalis 73 Campoletis sonorensis (Cameron).
74–75 Metasomal segment 1, ventral view. S1 = sternite 1, T1 = tergite 1 74 Rhyssa lineolata (Kirby) 75 Megarhyssa macrura (Linnaeus). Arrow indicates S1 fused to T1 posteriorly 76–79 Anterior tergites of metasoma 76 Lateral view, Olesicampe sp. Arrow points to thyridium 77 Dorsal view, Patrocloides montanus (Cresson). gs = gastrocoelus (delineated by dotted line), th = thyridium (linear, posterior part of gastrocoelus), ps = pseudothyridium, T2 = tergite 2 78–79 Lateral view 78 Patrocloides montanus sp = spiracle. T3 = tergite 3 79 Netelia sp. Arrow indicates crease separating tergite 3 and laterotergite 3. L2 = laterotergite 2, L3 = laterotergite 3.
Metasoma. 80 Lateral view, Allophrys divaricata MS2, etc. = metasomal segment 2, etc. 81 Tergites 2 to 4, dorsal view, Pimpla aequalis. Arrows indicate different sculpture on posterior 0.2 of tergites 82–83 Posterior segments of female, lateral view 82 Megarhyssa greenei. c = cercus , h = horn, o = ovipositor 83 Odontocolon albotibiale. Arrow indicates lack of horn 84 Sternites of female, ventral view, Rhyssa lineolata. Arrows indicate tuberculate ovipositor guides 85 Posterior segments of female, lateral view, Astiphromma splenium.
Posterior of female metasoma 86 Ventroposterior view, Lycorina albomarginata (Cresson). Arrow indicates medial, membranous area of hypopygium 87 Lateral view, Coleocentrus occidentalis Cresson 88 Ventral view, Lissonota scutellaris. Arrow indicates medial apical notch of hypopygium 89–91 Ovipositor, lateral view: 89 Pimpla aquilonia Cresson 90 Exetastes sp. nov. Arrow indicates dorsal, subapical notch 91 Phytodietus burgessi (Cresson). Arrow indicates dorsal, subapical nodus.
Posterior of male metasoma. 92 Dorsal view, Phytodietus vulgaris. Arrow showing longitudinal division of tergite 8. T7 = tergite 7, T8 = tergite 8, gon = gonoforceps 93 Ventrolateral view, Pimpla sp. 94 Ventrolateral view, Mesochorus sp. 95–96 Gonoforceps and aedeagus, lateral view 95 Thyreodon sp. 96 Rhyssa crevieri.
Cephalic sclerites of final larval instar 103 Lissonota occidentalis (Cresson) (copied from
111 Cephalic sclerites of final larval instar, Phrudus sp. Figures in upper left corner is whole mandible (upper) and close-up of apex of mandible (lower). Scale bar: 0.1 mm. Mandible and spiracle not same scale as main figure (spiracle is 27 µm in length) 112 Final larval instar, anterior of whole larva, Collyria catoptron, lateroventral view of head and thorax (stained with acid fuschin).
113 Mature egg of Ctenochira sanguinatoria (Ratzeburg) (re-drawn after
Part 1. Total-evidence parsimony analysis. Strict consensus cladogram (14 nodes collapse). Number of most parsimonious cladograms = 1728, length = 9917, CI = 0.15, RI = 0.44. Morphological characters optimized on tree using ACCTRAN. Open boxes are homoplasiously derived character states. Closed boxes are uniquely derived character states. Small numbers above boxes are character numbers. Small numbers below boxes are character states. Large numbers in squares above characters are total number of supporting characters (morphological and molecular). Large numbers in squares below characters are Bremer support values. Taxon names reflect classification prior to study (taxa with asterisks after name are formally re-classified in current study). “A” continued on part 2 of Fig.
Part 4. Total-evidence parsimony analysis strict consensus cladogram continued from part 1 of Fig.
Part 1. Parsimony analysis (morphological and biological characters only: Chs 1-141). Strict consensus cladogram (75 nodes collapse). Characters optimized on tree using ACCTRAN. Number of most parsimonious cladograms = 3872, length = 1527, CI = 0.15, RI = 0.60. See Figure
Part 1. Parsimony analysis (molecular characters only). Strict consensus cladogram (10 nodes collapse). Number of most parsimonious cladograms = 104, length = 8126, CI = 0.16, RI = 0.42. Numbers above branches are number of substitutions supporting each node or taxon. Taxon names reflect classification prior to study (taxa with asterisks after name are formally re-classified in current study). “A” continued on part 2 of Fig.
Part 1. Total-evidence Bayesian analysis maximum clade credibility tree. Numbers to the right of nodes are posterior probabilities. “EN” = change from ectoparasitoid to endoparasitoid (character 138); “EC” = change from endoparasitoid to ectoparasitoid; “E2” = change from ectoparasitoid to development inside hind gut with final ectoparasitoid phase; “I” = change from koinobiont to idiobiont (character 137); “K” = change from idiobiont to koinobiont. Taxon names reflect classification prior to study (taxa with asterisks after name are formally re-classified in current study). “B” is continued on part 2 of Fig.
Part 1. Bayesian analysis (molecular characters only) maximum clade credibility tree. Numbers to the right of nodes are posterior probabilities. Taxon names reflect classification prior to study (taxa with asterisks after name are formally re-classified in current study). “B” is continued on part 2 of Fig.
Morphological and biological characters only
The parsimony analysis using only the 141 morphological and biological characters produced 3872 equally parsimonious trees of 1527 steps (C.I. = 0.15, R.I. = 0.60). The strict consensus had 75 nodes collapsed (Fig.
Molecular characters only
The parsimony analysis with molecular characters found 104 trees of 8126 steps (C.I. = 0.16, R.I. of 0.42). Ten nodes collapsed in the strict consensus cladogram (Fig.
Total-evidence
The total-evidence Bayesian analysis majority credibility tree is shown in Figure
Molecular characters only
Similar to the total-evidence Bayesian analysis and the parsimony analysis with only molecular characters, the base of the tree for the Bayesian analysis with only molecular characters (Fig.
It was not the purpose of this study to examine relationships outside of Ichneumonidae, therefore only minimal outgroup sampling was used; however, all analyses did recover a monophyletic Ichneumonidae (Figs
The following taxa were sister group to all other Ichneumonidae depending on the analysis:
Aplomerus sp. (Xoridinae): total-evidence parsimony analysis (Fig.
Agriotypus armatus (Agriotypinae): morphology-only parsimony analysis (Fig.
Lycorina glaucomata (Lycorininae): molecular-only parsimony analysis (Fig.
Neorhacodes enslini (Neorhacodinae): Bayesian analyses (Figs
Our total-evidence parsimony analysis found the three Xoridinae species at the base of Ichneumonidae; however, they were not monophyletic with Aplomerus Provancher sister group to all other ichneumonids and (Odontocolon Cushman + Xorides Latreille) sister to all the rest (Fig.
In our morphological parsimony analysis, the clade comprising all ichneumonids except Agriotypus armatus was supported by 11 synapomorphies, of which one was uniquely derived: character 87(0) (T2 and T3 separate, with flexion line allowing movement) (Fig.
The evidence that Lycorininae is sister group to all other ichneumonids is based on 36 uniquely derived molecular substitutions that support the clade comprising all ichneumonids except Lycorina glaucomata in the molecular parsimony analysis (Fig.
The biology of Lycorina is not completely known, but species for which oviposition and development have been observed are koinobionts (
Neorhacodinae was sister group to all other Ichneumonidae in both Bayesian analyses on the basis of posterior probabilities of 100 supporting the grouping comprised of all ichneumonids except Neorhacodes enslini (Figs
In terms of previous studies,
Ophioniformes
Ophioniformes (including Tryphoninae) was supported in the total-evidence parsimony analysis by 20 total characters, of which 5 were morphological: 36(0) anterior transverse carina of propodeum complete (Fig.
In terms of the arrangement within Ophioniformes in the total-evidence parsimony analysis, Tryphoninae (including Neorhacodes enslini) was sister to all other taxa (part 4 of Fig.
The topology within Ophioniformes in the current total-evidence parsimony analysis is similar to that of
The anchored enrichment study by
Pimpliformes
Pimpliformes was monophyletic in all analyses with the exception of parsimony using only morphological characters (Table
In the current study, the topology within Pimpliformes was equivocal, depending on the method of analysis. The total-evidence parsimony and molecular parsimony analyses recovered Diplazontinae as sister to all other taxa, followed by the orthocentrine genera (not clustering together) and the rest of the taxa (part 4 of Fig.
In terms of previous hypotheses of internal Pimpliformes relationships,
Ichneumoniformes
Ichneumoniformes sensu stricto (of
Apart from the study of
The total-evidence parsimony analysis was the only one that recovered Pimpliformes, Ophioniformes and Ichneumoniformes s.l. as monophyletic (Table
The study of
Acaenitinae
In the total-evidence analyses (parsimony and Bayesian), the two exemplars of Acaenitinae (Spilopteron occiputale (Cresson) and Coleocentrus rufus Provancher) did not cluster together. Coleocentrus rufus was sister to Collyria catoptron Wahl (Collyriinae), whereas S. occiputale had various placements within Pimpliformes, such as sister to Cylloceria melancholica (Gravenhorst) (Cylloceriinae) in the total-evidence parsimony analysis (part 2 of Fig.
The current analysis and
Adelognathinae
The total-evidence parsimony analysis found Adelognathinae to be the sister group to ((Microleptinae + Ateleutinae) + ((Aptesini + Cryptini including Echthrus reluctator (Linnaeus)) + (Phygadeuontinae + (Alomya debellator (Fabricius) + Ichneumoninae))) (part 3 of Fig.
Agriotypinae
The total-evidence parsimony analysis found that Agriotypus armatus was sister species to Euceros sp. nov. (Eucerotinae) and these two species were sister group of the clade listed above under Adelognathinae , (i.e., Adelognathinae… to Ichneumoninae) (part 3 of Fig.
The monotypic Agriotypinae is morphologically aberrant in that it has strongly sclerotized posterior metasomal tergites, which is an autapomorphy within Ichneumonoidea. They are also biologically unusual in that they are aquatic idiobiont ectoparasitoids of prepupae and pupae of Trichoptera in fast-running streams (
With respect to the putative relationship of Agriotypinae and Eucerotinae, of the eight morphological synapomorphies, five of them undergo transformations/ reversals in one taxon or the other. For example, the uniquely derived character 140(6) (oviposition into Trichoptera cases) which supports both taxa, changes to character 140(8) (oviposition on to leaves) in Eucerotinae. Given the extremely different morphology and biology of these two subfamilies, it appears likely that these two highly derived taxa have clustered together because of coincidental homoplasious traits and the lack of any phylogenetic signal linking them to another group. Having said that, both the parsimony and Bayesian total-evidence analyses placed Agriotypinae in relatively the same part of the tree (i.e. somewhere within Ichneumoniformes s.l.), which is similar to the conclusions of
Alomyinae
In the total-evidence parsimony and Bayesian analyses (Figs
Since the beginning of the 20th century, Alomya has been recognized as closely associated with Ichneumoninae. It has been treated variously as a separate subfamily by
Our study, with three genes and a large morphological data set, generally places Alomya as the sister group of the Ichneumoninae (both total-evidence analyses and parsimony with only morphological characters). Based on these results, it would be possible to expand Ichneumoninae to encompass Alomya (as a tribe), although maintenance of Alomyinae is equally acceptable. We prefer to classify Alomya and the closely related genus Megalomya Uchida as a tribe within Ichneumoninae, and therefore formally synonymize Alomyinae with Ichneumoninae. Since both classifications are equally acceptable, there is no need for a discussion of similarities and differences between Ichneumoninae and Alomya as this would not justify the rank of Alomyini/ Alomyinae one way or the other. A detailed re-description of the larva of Alomya is presented in Appendix
Anomaloninae
The two Anomaloninae exemplars (Anomalon picticorne (Viereck) and Therion texanum (Ashmead)) clustered together in the total-evidence and morphology-only parsimony analyses (Figs
Anomaloninae was the sister group of Ophioninae in the total-evidence parsimony analysis, supported by 33 characters (13 morphological) including the uniquely derived character 54(2): cell 1 +2Rs (areolet) of fore wing with vein Rs absent so that the only cross-vein is distad vein 2m-cu (Fig.
Previous morphological analyses (e.g.,
Ateleutinae
Ateleute sp. nov. was placed wihin Ichneumoniformes s.l. in all of our analyses, except parsimony with only morphological characters in which it was sister species to Nonnus sp. in a portion of the tree that lacked resolution (part 1 of Fig.
Previously,
Banchinae
The four exemplar species of Banchinae were closely related in all analyses. The two Glyptini species (Apophua simplicipes (Cresson) and Sphelodon phoxopteridis (Weed)) were always sister species and Atrophini (Lissonota scutellaris (Cresson)) was generally sister to Banchini (Exetastes bioculatus Cresson). In the total-evidence parsimony analysis, Glyptini, (Atrophini + Banchini) and Notostilbops sp. (Stilbopinae) were found in a clade that lacked internal resolution (part 6 of Fig.
Brachycyrtinae
The placement of Brachycyrtus wardae Bennett varied somewhat between our analyses. Total-evidence parsimony placed it as the sister to Labeninae, supported by 32 characters (6 homoplasious morphological) with a Bremer support of 10 (part 3 of Fig.
As noted in the Introduction,
Campopleginae
Both total-evidence analyses strongly supported the monophyly of the eight exemplar species of Campopleginae. In the parsimony analysis, the subfamily was supported by 43 synapomorphies (5 morphological, none of which were uniquely derived) with a Bremer support value of 10+ (part 6 of Fig.
The current results strongly support the monophyly of Campopleginae (i.e., the group comprised of the genera that were previously classified in Townes’s Campoplegini and Porizontini). They also support the removal of Hellwigiini and Nesomesochorini from Campopleginae. In terms of the sister-group relationship, both total-evidence analyses support Cremastinae as sister to Campopleginae with Nesomesochorinae and (Ophioninae + Anomaloninae) also related, albeit more distantly. With respect to internal relationships, Townes’s Campoplegini and Porizontini were not supported (Campoplex sp. was sister to all other genera, but the other Campoplegini exemplar, Casinaria grandis, clustered within the six Porizontini species. Likewise, there was no support for the genus groups of
Claseinae
Claseinae (Clasis sp. nov.) was sister group to Pedunculinae (Pedunculus sp. nov.) in all analyses except the morphology-only parsimony analysis in which it was unresolved near the base of Ichneumonidae (part 1 of Fig.
Clasis Townes was originally placed within the cryptine tribe Phygadeuontini (
The fact that our total-evidence parsimony analysis supports the monophyly of Labeninae as it was defined by
Collyriinae
Both total-evidence analyses (Figs
Historically, the taxonomic placement of Collyria has been contentious.
More recently,
Cremastinae
The two species of Cremastinae, Eiphosoma pyralidis Ashmead and Xiphosomella setoni Johnson, clustered together in all analyses with very strong support. The total-evidence parsimony analysis supported this grouping with 61 synapomorphies (5 morphological) and a Bremer support value of 10+ (part 6 of Fig.
The monophyly of Cremastinae has generally gone unquestioned, supported largely by the unique synapomorphy (within Ichneumonidae) of the sclerotized bridge of the middle and hind tibiae separating the insertion points of the tarsus and tibial spurs (
Cryptinae sensu stricto and Cryptinae sensu lato
The seven species of Cryptinae sensu stricto, i.e., members of the tribes Cryptini and Aptesini (formerly Hemigastrini), grouped together in both of the total-evidence analyses and the Bayesian (molecular-only) analysis (Table
In the analyses in which Cryptinae s.s. was monophyletic, it was always related to species in the subfamilies Ateleutinae, Microleptinae, Phygadeuontinae, Alomyinae, Ichneumoninae and Adelognathinae. The specific relationships of each of these subfamilies are discussed more fully in the section above on Ichneumoniformes, as well as in the respective subfamily sections.
The 11 species of Cryptinae sensu lato (species of Ateleutinae, Phygadeuontinae and Cryptinae s.s.) never shared a unique, common ancestor, regardless of the analysis. This was caused by: 1) placement of Ateleute sp. nov. (Ateleutinae) away from the other species; 2) paraphyly of the other 10 species of Cryptinae s.l. with respect to Alomyinae + Ichneumoninae. In summary, we found no evidence to support the monophyly of Cryptinae s.l. (= Gelinae) of
Regarding Phygadeuontinae, in the total-evidence parsimony analysis, the three species (Endasys patulus (Viereck) + (Acrolyta sp. + Mastrus sp.)) grouped together with weak support (13 synapomorphies and a Bremer support value of 1) (part 3 of Fig.
With respect to the relationship between Cryptinae s.s. and Ichneumoninae,
In terms of more recent, sequence-based studies, some of the analyses of
Finally, in terms of support for the tribes of Cryptinae s.s., the total-evidence parsimony analysis had strong support for Cryptini (including Echthrus reluctator) with 30 synapomorphies and a Bremer support of 6, and Aptesini had 25 synapomorphies and a Bremer support of 9. The Bayesian total-evidence analysis also supported both tribes (BPP = 100 for each) (part 2 of Fig.
Ctenopelmatinae
Ctenopelmatinae was never recovered as monophyletic (Table
In terms of support for the ctenopelmatine tribes, of the five for which multiple species were included, only two (Perilissini and Mesoleiini) had their species clustering together in the total-evidence parsimony analysis. The other three large tribes (Pionini, Euryproctini and Ctenopelmatini) were not recovered as monophyletic (part 5 of Fig.
Previous studies have questioned the monophyly of Ctenopelmatinae.
Our total-evidence analyses indicate that the taxa currently comprising Ctenopelmatinae belong within Ophioniformes and yet not within the “higher Ophioniformes” (Anomaloninae, Campopleginae, Cremastinae, Nesomesochorinae, Ophioninae). Ctenopelmatinae may be related to Mesochorinae (part 4 of Fig.
Cylloceriinae
In the total-evidence parsimony analysis, Cylloceriinae (Cylloceria melancholica) was sister taxon to Spilopteron occiputale (Acaenitinae) in the middle of our Pimpliformes grouping (part 2 of Fig.
The combined analyses of
In our study, all the exemplars of Diacritinae, Acaenitinae, Collyriinae, Cylloceriinae, Diplazontinae and Orthocentrinae clustered together in only one analysis: the total-evidence Bayesian analysis (part 1 of Fig.
Diacritinae
The total-evidence parsimony analysis recovered our single exemplar of Diacritinae (Diacritus incompletus) in a clade within Pimpliformes with equivocal relationships as follows: Diacritus/ Coleocentrus rufus + Collyria catoptron/ all species of Pimplinae, Poemeniinae and Rhyssinae (part 2 of Fig.
Previous analyses have found Diacritinae as either sister to all Pimpliformes (
Diplazontinae
The monophyly of Diplazontinae (Diplazon laetatorius (Fabricius) and Woldstedtius flavolineatus (Gravenhorst)) was supported in all of our analyses, regardless of the data used or the method of analysis (Table
As described above in the section on Cylloceriinae, all previous studies have placed Diplazontinae within Pimpliformes, generally as sister group to Orthocentrinae (Gauld and Wahl 1998;
Eucerotinae
As described in the section above on Agriotypinae, in the total-evidence parsimony analysis part 3 of (Fig.
Euceros Gravenhorst is unique in Ichneumonidae in that species lay eggs on vegetation which hatch into planidial larvae (
Previous phylogenetic studies have attempted to ascertain the relationships of Eucerotinae within Ichneumonidae. A morphological analysis by
Hybrizontinae
Hybrizon rileyi, the exemplar of Hybrizontinae in our study, was sister to Lycorina glaucomata (Lycorininae) in the total-evidence parsimony analysis and was nested within the Ctenopelmatinae and relatives clade (part 5 of Fig.
Similar to Agriotypus, Hybrizon Fallén and its relatives have had a varied placement over time: included in Braconidae (
In terms of proposed placement within Ichneumonidae,
Ichneumoninae
The following discussion pertains to the 13 exemplar species of Ichneumoninae excluding Alomyini. See the section on Alomyinae (above) for the rationale for moving Alomya and Megalomya within Ichneumoninae (as Alomyini).
The 13 species of Ichneumoninae were monophyletic in all analyses (Table
In terms of the tribes, all analyses recovered the two species of Platylabini together (Cyclolabus impressus (Provancher) + Linycus exhortator (Fabricius)) and the two species of Heresiarchini (Coelichneumon eximius (Stephens) + Protichneumon grandis (Brullé)) were also always monophyletic, although the latter pair was always nested within Ichneumonini (5 species), as was the single exemplar of Listrodromini (Dilopharius otomitus (Cresson)). All analyses except parsimony with only morphological characters recovered Phaeogenini as monophyletic (Stenodontus sp. nov. + (Centeterus euryptychiae (Ashmead) + Phaeogenes hebrus (Cresson))), and (Platylabini + Phaeogenini) was sister to all other species in the total-evidence and molecular-only parsimony analyses (Figs
In terms of tribal relationships,
With respect to monophyly and relationships of the other tribes of Ichneumoninae, none of the studies of
Labeninae
Labeninae was monophyletic in all analyses except parsimony using only morphology (Table
Some previous studies have suggested that Labeninae may be sister group to all ichneumonids except Xoridinae, for example, most analyses in
With respect to tribal relationships within Labeninae, our analysis only included exemplars of three tribes (no Xenothyrini exemplars). In the consensus tree of the total-evidence parsimony analysis, the tribes Labenini (Labena grallator (Say) + Apechoneura sp.), Orthognathelini (= Groteini) (Grotea anguina Cresson + Labium sp.) and Poecilocryptini (Poecilocryptus nigromaculatus) had equivocal relationships (part 3 of Fig.
Lycorininae
The placement of Lycorininae (Lycorina glaucomata) was one of the least stable of any taxon in our analyses. The total-evidence parsimony analysis placed L. glaucomata within the clade of Ctenopelmatinae and relatives as sister to Hybrizontinae (see section on Hybrizontinae for details on support). In the parsimony analysis with only molecular characters, L. glaucomata was sister to the rest of Ichneumonidae (part 1 of Fig.
What is known of the biology of Lycorina is that they are koinobionts (
In summary, the majority of our analyses agreed with
Mesochorinae
The five species of Mesochorinae only clustered together in the morphology-only parsimony analysis (part 3 of Fig.
Chineater masneri does have a large, rhombic areolet (character 54(1)) (Fig.
With respect to the four other mesochorine species, the total-evidence parsimony analysis placed them in a clade with equivocal relationships within Ophioniformes as follows: Mesochorinae except Chineater/ Ctenopelmatinae and relatives/ Metopiinae to Campopleginae (part 4 of Fig.
There have been some previous studies that suggested that Mesochorinae may render Ctenopelmatinae paraphyletic (
Metopiinae
Similar to Mesochorinae, all four species of Metopiinae only clustered together in the morphology-only parsimony analysis (part 3 of Fig.
In terms of the placement of Scolomus,
The current total-evidence parsimony analysis placed Scolomus within the Ctenopelmatinae and relatives grouping (part 5 of Fig.
Unfortunately, we were not able to include any other of the problematic genera of Metopiinae in our analysis: Bremiella Dalla Torre, Ischyrocnemis Holmgren, and Lapton Nees; therefore we cannot comment on their relatedness to the four exemplar metopiines we analyzed.
Microleptinae
Microleptinae (Microleptes sp.) was sister to Ateleute sp. nov. (Ateleutinae) in the parsimony total-evidence analysis, supported by 37 synapomorphies (7 morphological, none of which were uniquely derived) and a Bremer support of 6. These two taxa were sister group of (Cryptinae + (Phygadeuontinae + (Alomyinae + Ichneumoninae))) (part 3 of Fig.
Previous studies were equivocal in their placement of Microleptinae.
Our study provides support for the placement of Microleptinae near the base of Ichneumoniformes s.l. In terms of morphology, there are no compelling characters for this placement – the only uniquely derived character in this region of the tree is Character 9 (state 1): central flagellomeres of male with elliptical or longitudinal ridge-like tyloids which supports Adelognathinae + ((Microleptinae + Ateleutinae) + (Cryptinae + (Phygadeuontinae + (Alomyinae + Ichneumoninae)))). This character; however, has a reversal in Microleptinae + Ateleutinae (part 3 of Fig.
Neorhacodinae
Neorhacodinae (Neorhacodes enslini) was perhaps the most unstable taxon in our analyses. The total-evidence parsimony analysis placed N. enslini within Tryphoninae (as sister to Phytodietini) (part 4 of Fig.
Originally,
Given the equivocal placement of Neorhacodes enslini in the current study, we are not able to make any precise statements with respect to the relationships of Neorhacodinae. There are no compelling morphological or biological characters in our study that link Neorhacodinae to Tryphoninae. The egg of Neorhacodinae is not known, and determining whether it bears a stalk or not would help greatly in determining whether the sister-group relationship of Neorhacodinae and Phytodietini is artefactual or not. The current study coded the host of Neorhacodinae as Hymenoptera, which is similar to most Tryphoninae, although Phytodietini are parasitoids of Lepidoptera. Furthermore, one could argue that coding the hosts for Neorhacodes (aculeate Hymenoptera) (
Nesomesochorinae
Nesomesochorinae (Nonnus sp. and Chriodes sp.) was only recovered as monophyletic in the total-evidence parsimony analysis (part 6 of Fig.
Nesomesochorini was proposed by
Ophioninae
Ophioninae (Enicospilus flavostigma Hooker, Hellwigia obscura, Ophion sp. Skiapus sp. and Thyreodon sp.) was supported in both total-evidence analyses and the parsimony analysis with only morphological characters (Table
The monophyly of the majority of the genera of Ophioninae has long been established on the basis of the relatively rare (in Ichneumonidae) fore wing areolet lacking vein Rs so that the only cross vein is distad vein 2m-cu (character 54, state 2) (Fig.
Orthocentrinae
Orthocentrinae (Megastylus sp. nov., Orthocentrus sp. and Proclitus speciosus Dasch) was only recovered as monophyletic in the parsimony analysis using only morphological characters (part 2 of Fig.
Previous definitions of Orthocentrinae have differed depending on whether authors considered Orthocentrus Gravenhorst and relatives (the Orthocentrus group of genera) to be related to Helictes Haliday and relatives (the Helictes group of genera).
Given the strong support of Orthocentrinae in other studies, it is likely that the lack of monophyly of Orthocentrinae is artefactual, perhaps because of low taxon sampling and, as discussed by
Orthopelmatinae
The total-evidence parsimony analysis placed Orthopelmatinae as sister group to all other Ichneumonidae except the exemplars of Xoridinae (part 1 of Fig.
Previous analyses have had trouble discerning the precise relationships of Orthopelmatinae.
None of our analyses recovered a sister-group relationship of Orthopelmatinae and Ophioniformes, although it is noted that moving O. mediator as sister to Ophioniformes only lengthened the total-evidence parsimony tree by 9 steps (9926). The parsimony analysis with only morphological characters (part 3 of Fig.
Oxytorinae
Our analyses generally placed Oxytorinae (Oxytorus albopleuralis) with exemplars of Ophioniformes; however, its placement within this group changed depending on the analysis. In our total-evidence parsimony analysis, Oxytorinae clustered within a clade comprised of all 14 Ctenopelmatinae species as well as Hybrizontinae, Lycorininae, Tatogastrinae, Chineater masneri (Mesochorinae) and Scolomus sp. (Metopiinae) (part 5 of Fig.
Prior to
Pedunculinae
Pedunculinae (Pedunculus sp. nov.) was strongly supported as sister group to Claseinae (Clasis sp. nov.) in all analyses except the parsimony analysis with only morphological characters in which its relationships were unclear. In both total-evidence analyses, these two subfamilies were placed near the base of Ichneumoniformes s.l. See the sections on Brachycyrtinae, Claseinae and Labeninae (above) for discussion of support of this node and further relationships of Pedunculinae.
Phygadeuontinae
As discussed above in the section on Cryptinae, Phygadeuontinae (Acrolyta sp., Endasys patulus and Mastrus sp.) was supported in both total-evidence analyses (part 3 of Fig.
Our total-evidence analyses results, albeit with very limited sampling, concur with that of
Pimplinae
The seven species of Pimplinae (Perithous divinator, Acrotaphus wiltii (Cresson), Clistopyga recurva (Say), Dolichomitus irritator (Fabricius), Zaglyptus pictilis Townes, Pimpla annulipes Brullé and Theronia bicincta) never clustered together with unequivocal support in any of our analyses (Table
Historically, Pimplinae in the broad sense (i.e., including related taxa such as Poemeniinae and Rhyssinae) was one of the five traditional subfamilies of Ichneumonidae (
With respect to the placement of Pimplinae in the current study, when there was some support for the family (in 87% of the trees in the morphology-only parsimony analysis and the Bayesian total-evidence analysis with BPP = 93), Pimplinae was sister to (Poemeniinae + Rhyssinae). This is the same as the relationship postulated by the morphological analysis of
Concerning tribal monophyly and their relationships, the following previous hypotheses have been postulated:
1) (Delomeristini (not including Perithous Holmgren) + (Ephialtini + (Perithous + Pimplini including the Theronia group)) (morphological parsimony analysis of
2) (Pimplini + (Delomeristini including Perithous) + Ephialtini)) (morphological parsimony analysis of
3) (Delomeristini (including Perithous) + (Pimplini + Ephialtini)) (combined morphological and molecular parsimony analysis of
4) (Delomeristini (including Perithous and Pseudorhyssa) + Theroniini) + (Pimplini + Ephialtini), with Xanthopimpla clustering outside Pimplinae) (anchored enrichment analysis using amino acids of
The current study supported monophyly of Ephialtini in all studies except parsimony with only morphological data. For example, the Bayesian total-evidence analysis had the following topology: (Perithous divinator + Theronia bicinta) + (Pimpla annulipes + (Dolichomitus irritans + (Acrotaphus wiltii/ Clistopyga recurva/ Zaglyptus pictilis))). (part 2 of Fig.
Poemeniinae
Poemeniinae (Neoxorides caryae (Harrington) and Poemenia albipes) was supported unequivocally in all analyses except the Bayesian analysis using only molecular characters (Table
In terms of relationships, there was strong support for Poemeniinae being the sister group to Rhyssinae (e.g., Bayesian total-evidence, BPP = 99) (part 2 of Fig.
Historically, Poemenia Holmgren was placed within the traditional “Pimplinae” of early authors, e.g.,
More recently, Pseudorhyssa clustered within Pimplinae in all of the full-data-set analyses of
Rhyssinae
Similar to Poemeniinae, Rhyssinae (Megarhyssa greenei Viereck, Rhyssa crevieri (Provancher) and Rhyssella nitida (Cresson) was well-supported – it was unequivocally monophyletic in all of our analyses (Table
As described above for Poemeniinae, Rhyssa Gravenhorst and its relatives were also historically placed in the traditional Pimplinae (
Sisyrostolinae
Sisyrostolinae (Brachyscleroma sp. and Erythrodolius calamitosus Seyrig) was not recovered as monophyletic in any of our analyses (Table
The genera to which our two exemplars of Sisyrostolinae belong were, until recently, included in the subfamily Phrudinae (e.g.,
More recent phylogenetic studies investigated the monophyly of Phrudinae.
Our analyses do not uphold the monophyly of Sisyrostolinae, nor its separate status from Tersilochinae (including the Phrudus group). It is likely that the Erythrodolius group is monophyletic (Melanodolius Saussure and Icariomimus Seyrig are very closely related based on morphology, for example, the frons in all three genera bears a longitudinal ridge, and the latter may even be paraphyletic with respect to Erythrodolius (
Stilbopinae
The two species of Stilbopinae (Stibops vetulus and Notostilbops sp. nov.) were never sister taxa in any of our analyses (Table
Tatogastrinae
Tatogastrinae (Tatogastra nigra) was one of the small subfamilies that clustered within the “Ctenopelmatinae and related subfamilies” clade in the total-evidence parsimony analysis (part 5 of Fig.
In terms of previous hypotheses concerning the relationships of Tatogastrinae,
The morphological analysis of
Tersilochinae
As discussed in Sisyrostolinae (above), the five species of Tersilochinae sensu lato (Allophrys divaricata, Phrudus sp., Peucobius fulvus Townes, Stethantyx nearctica and Tersilochus sp.) did not cluster together in any analyses, except in a grouping that also included the two species of Sisyrostolinae (in both total-evidence analyses) (Table
In contrast, the Tersilochinae sensu stricto, which is equivalent to the Tersilochinae of
Our study generally supports a relatively close relationship of the Phrudus group and Tersilochinae sensu stricto; however, Erythrodolius and Brachyscleroma (Sisyrostolinae) clustered within this clade as follows: (Phrudus sp. + (Erythrodolius calamitosus + Peucobius fulvus)) + (Brachyscleroma sp. + Tersilochinae sensu stricto) (e.g., part 4 of Fig, 117, part 1 of Fig.
Tryphoninae
Tryphoninae (Eclytus sp., Idiogramma longicauda, Zagryphus nasutus (Cresson), Netelia sp., Phytodietus vulgaris, Cteniscus sp., Cycasis rubiginosa (Gravenhorst) and Polyblastus sp.) was not supported in any of our analyses (Table
Historically, Tryphoninae was one of the five major groups of Ichneumonidae (
In terms of phylogenetic analyses,
Based on the current study,
With respect to the findings of
Relationships within Tryphoninae based on the total-evidence parsimony analysis were as follows: (Neorhacodinae + Phytodietini) + (Idiogrammatini + (Polyblastus group of Tryphonini + (Eclytini + Oedemopsini) + Exenterus group of Tryphonini))) (part 4 of Fig.
Xoridinae
Xoridinae (Aplomerus sp., Odontocolon albotibiale (Bradley) and Xorides stigmapterus (Say)) was recovered as monophyletic in the parsimony analysis using only morphological characters, as well as both Bayesian analyses. In the morphological parsimony analysis (part 1 of Fig.
With respect to the monophyly of Xoridinae, on the basis of morphology alone, Xoridinae is well-suppported, with 23 synapomorphies, including one uniquely derived: character 113, state 1: larval mandible with spines at base of blade (part 1 of Fig.
In terms of the placement of Xoridinae within Ichneumonidae, the total-evidence parsimony analysis provides some support that Xoridinae (or a subset of xoridine taxa) is the sister group of all other Ichneumonidae.
Inclusion of biological characters in our data matrix allows an examination of the evolution of these characters within Ichneumonidae, at least with respect to the exemplar taxa used in this analysis. In terms of the validity of this kind of analysis, we agree that biological characters can be complex and our unweighted, unordered analysis may not take into account differences in the likelihood of particular character state changes evolving relative to others. Despite this, we believe that there is value in this kind of analysis. An unweighted, unordered analysis is objective. It does not place pre-conceived notions on the direction of evolution, nor does it make subjective decisions on the relative importance of characters. Whether the current analyses of the evolution of biological characters in Ichneumonidae is realistic or over-simplistic is a question that will hopefully foster discussion on the evolution of these interesting traits, producing hypotheses that can be tested by future analyses.
Three biological characters have been optimized on to the total-evidence parsimony strict consensus cladogram as follows: character 137: timing of larval maturation (Fig.
In the total-evidence parsimony analysis, character 137, timing of larval maturation, had a length of 8 steps (Fig.
The ancestral state for Ichneumonidae favours koinobiosis, although this requires some discussion. The state is unknown for Aplomerus sp. (Xoridinae) and is equivocal for the next node (the other two xoridines which are both idiobionts). There is strong morphological evidence that Aplomerus Provancher belongs to Xoridinae (
In terms of the direction of evolution of this character in the parsimony analysis, it has transitions in both directions, slightly favoured in the direction of koinobiosis to idiobiosis (five times) compared to vice versa (three times). The transition from a supposedly more specialized koinobiont to a less specialized strategy (idiobiosis) was not hypothesized by
Comparing the evolution of this character in the total-evidence Bayesian analysis, koinobiosis is favoured as plesiomorphic within Ichneumonidae (Fig.
Examination of the evolution of ectoparasitism versus endoparasitism (character 138) in the total-evidence parsimony analysis reveals that ectoparasitism is plesiomorphic in both Braconidae and Ichneumonidae (Fig.
In contrast, under DELTRAN optimization (not shown), transitions from ectoparasitism to endoparasitism occurred ten times: 1) Aleiodes terminalis (Braconidae); 2) Orthopelmatinae; 3) ancestor of Pimpliformes; 4) Pimpla annulipes; 5) Theronia bicincta; 6) Euceros sp. nov. 7) Microleptes sp.; 8) Ichneumoninae including Alomya debellator; 9) Neorhacodes enslini; 10) ancestor of Ophioniformes. Only one transition from endoparasitism to ectoparasitism occurred in the ancestor of higher Pimpliformes. Lycorininae had the same transition to state 2 as for ACCTRAN.
The order of host/ source of larval nutrition used by our exemplar taxa (character 139) is optimized on the total-evidence parsimony strict consensus tree in Fig.
In terms of additional evidence supporting one of these three host orders as plesiomorphic for Ichneumonidae, all easily pre-date the origin of Ichneumonidae (at least 85 mya) (
A comparison of the relative frequencies of host use of the four major holometabolous orders in our parsimony analysis shows that Lepidoptera is the most prevalent host (for 35 % of our 131 exemplar ichneumonid species), compared to Hymenoptera (30 %), Coleoptera (12 %) and Diptera (5 %). These percentages are comparable to the known host use by ichneumonids at the subfamily level: Lepidoptera (18 subfamilies, 43 % of total); Hymenoptera (16 subfamilies, 38 %); Coleoptera (13 subfamilies, 31 %) and Diptera (8 subfamilies, 19%) (
Examination of the different transitions that occur within this character in the parsimony total-evidence tree reveals that of the 28 state changes, there were 13 different types of transitions, (e.g., Lepidoptera to Coleoptera, Hymenoptera to Coleoptera, etc). The order that was most often plesiomorphic in the state changes was Hymenoptera with the following apomorphic states and number of changes: changes to Lepidoptera (8); Coleoptera (5); Diptera (1); Neuroptera (1); Trichoptera (1) and facultatively herbivorous (1). The next most common order that was plesiomorphic in these transitions was Lepidoptera, as follows: changes to Coleoptera (4); Hymenoptera (2) and Diptera (1). Thirdly, there were two transitions from Diptera: one to Coleoptera and one to Hymenoptera. Lastly, there was one transition from Coleoptera to egg predation and one change from egg predation to parasitization of spiders. Therefore, ichneumonids that parasitize Hymenoptera appear much more likely to switch to different host orders compared to, for example, an ichneumonid that parasitizes Lepidoptera, Diptera or Coleoptera. In fact, over half of all of the total host transitions (17 of 28 state changes) in Fig.
Overall, the two total evidence analyses obtained the most resolution of relationships, followed by the molecular analyses and finally, the parsimony analysis with only morphological and biological characters. There was general congruence between the parsimony and Bayesian total-evidence analyses, except for at the base of Ichneumonidae (described below).
The relative support of different groupings within Ichneumonidae is shown in Table
In both total evidence analyses, Pimpliformes was sister group to Ichneumoniformes s.l., which agrees with
There were several other well-supported groupings of subfamilies including higher Pimpliformes, higher Ophioniformes, (Claseinae + Pedunculinae) and (Stilbops + Banchinae including Notostilbops). At the subfamily level, some subfamilies were well-supported across analyses: Banchinae (including Notostilbops), Campopleginae, Cremastinae, Diplazontinae, Ichneumoninae (including Alomya), Labeninae, Mesochorinae (excluding Chineater), Metopiinae (excluding Scolomus), Poemeniinae, Rhyssinae and Tersilochinae s.s. Moderate support (i.e., support in two to three analyses) was found for Anomaloninae, Cryptinae, Ophioninae, Phygadeuontinae and Xoridinae. Weak support (i.e., in only one analysis) was found for Acaenitinae, Nesomesochorinae and Orthocentrinae.
In contrast, the following subfamilies were never supported: Ctenopelmatinae, Pimplinae, Sisyrostolinae, Stilbopinae, Tersilochinae s.l. and Tryphoninae. Ctenopelmatinae was supported in the total-evidence parsimony analysis with the inclusion of Hybrizontinae, Lycorininae, Oxytorinae and Tatogastrinae. Tryphoninae was also supported in this analysis with the inclusion of Neorhacodinae. The most equivocally-placed subfamilies were Lycorininae, Neorhacodinae, Orthopelmatinae and Xoridinae.
Optimization of biological characters on the total evidence phylogenies hypothesized that for Ichneumonidae, ectoparasitism is plesiomorphic to endoparasitim. The ancestral state for timing of larval maturation is koinobiosis in both analyses; however, the lack of data for the sister taxon to all other ichneumonids in the parsimony analysis (Aplomerus) raises some doubt regarding this hypothesis. If Aplomerus is an idiobiont (as expected), then the parsimony analysis would support idiobiosis as plesiomorphic to koinobiosis. Finally, the ancestral host for Ichneumonidae is hypothesized to be Hymenoptera or Lepidoptera, although if the host of Aplomerus is determined to be Coleoptera (as expected), then Coleoptera would be the hypothesized ancestral host in one of the five analyses.
The authors would like to thank the curators of the institutions listed in the methods section for loan/ deposition of specimens and larval slides. In addition, Dr. G. Broad (Natural History Museum, London) provided host remains from which the larval exuvium of Phrudus defectus could be extracted. Dr. T. Shanower (US Department of Agriculture, Albany, California) provided larval specimens of Collyria catoptron. Cambridge University Press gave permission to copy the figures of the egg of Euceros frigidus (Figs
Taxonomic descriptions
Cephalic sclerites of mature larva of Phrudus defectus Stelfox.
Fig.
Cephalic sclerites mostly well-sclerotized. Epistomal suture (character 105): region distorted and not reconstructed in drawing, but possibly completely sclerotized and forming epistomal band (coded as “?”). Labral sclerite (character 108) & clypeolabral plates (character 130) unknown because labral region distorted (both scored as “?”). Stipital sclerite present, more or less horizontal and median end contacting labial sclerite (character 119. state 0) and without lateral plate-like extension (character 120, state 0). Pleurostoma only partially visible due to distortion; posterior struts of inferior mandibular processes not connected by band (character 114, state 0); inferior mandibular process dorsad to dorsal margin of labial sclerite (character 107, state 0); accessory pleurostomal area (character 106) not discernible (coded as “?”). Hypostoma well-sclerotized and long (character 115, state 0); lateral end simple, not divided or upcurved (character 116, state 0). Hypostomal spur present and long, about 2.0× as long as its basal width (character 117, state 0), meeting stipital sclerite near middle (character 118, state 0). Labial sclerite nearly circular (character 124, state 1), about as long as wide (character 125, state 0), not produced ventrally as a spine (character 126, state 0). Salivary orifice U-shaped (character 133, state 1). Prelabial sclerite absent (character 127, state 0). Sclerotized plate ventrad labial sclerite absent (character 129, state 0). Maxillary and labial palpi each bearing 2 sensilla (character 123, state 0). Mandible uniformly well-sclerotized (character 111, state 0), cone-shaped and apex with small, tooth-like projection (character 109, state 1) (Fig.
Material examined: Phrudus defectus Stelfox last larval instar exuvium slides: UNITED KINGDOM: Isle of Man, Laxey, Baldhoon Road, Crofton, SC4284; F.D. Bennett; from Epuraea melanocephala in sycamore flowers; exposed 1–2.vi.2008, adult emerged 4.v.2009 [DBW preparation 28.I.2012b] (EMUS); UNITED KINGDOM: Isle of Man, Laxey, Mooar Glen, SC43284; F.D. Bennett; from Epuraea melanocephala in sycamore flowers; collected 31.v.2008, adult emerged 11.v.2009 [DBW preparation 28.I.2012c] (EMUS).
Comments. Distortions of the two preparations (DBW preparations 28.I.2012b and 28.I.2012c) do not allow clear views of the dorsal portion of the cephalic capsule, and hence structures above the inferior mandibular processes are not shown. Figure
Cephalic sclerites of mature larva of Collyria spp.
Fig.
Cephalic sclerites mostly weakly sclerotized or absent. Epistomal suture unsclerotized (character 105, state 0). Labral sclerite absent (character 108, state 1). Clypeolabral plates absent (character 130, state 0). Stipital sclerite absent (character 119. state 2) and without lateral plate-like extension (character 120, state 0). Pleurostoma difficult to discern, but apparently not laterally expanded (character 106, state 0) and posterior struts of inferior mandibular processes short and not obviously connected by band (character 114, state 0). Hypostoma absent (character 115, state 2). Hypostomal spur absent (character 117, state 2). Labial sclerite (character 124) difficult to discern ventrally, therefore shape scored as “?”. Salivary orifice (character 133) not visible (scored as “?”). Prelabial sclerite absent (character 127, state 0). Sclerotized plate ventrad labial sclerite absent (character 129, state 0). Maxillary and labial palpi (character 123) not visible (scored as “?”). Mandible (scored for C. coxator from the description of
Material examined: Collyria coxator (Villers) last larval instar exuvium slide mount: Locality unspecified. Label 1: Slide no. 284. Don G. Salt. Label 2: Collyria calcitrator (Grav.) JRTS 1955 (NMNH); Collyria catoptron Wahl slit and macerated last instar whole larva slide mount: CHINA, Gansu Province, Yuzhong County, ix.1998, ex. Cephus fumipennis, T. Shanower et al. [DBW preparation 3.I.2011] (EMUS).
Comments: The nature of the cephalic sclerites of the mature larva of Collyria is not straightforward.
Between Salt’s detailed study and Short’s slide, Collyria appears to be quite unique amongst the Ichneumonidae for the drastic reduction of the cephalic sclerites. Reduction of the sclerites is associated with not spinning a cocoon (such as in the Anomaloninae, Ichneumoninae, Metopiinae, and Pimplini), but never to such an extent as in Collyria. Preserved larvae of a second species, Collyria catoptron Wahl, were available for comparison. A number of larvae were longitudinally slit and macerated in sodium hydroxide solution, with the resulting skins stained with acid fuschin and then slide mounted. Whole larvae were also stained with acid fuschin and then examined. No evidence of sclerotized structures could be found on the mounted skins. The stained whole specimens showed the general mouthpart regions as convexities and furrows but no sclerotized structures were present (Fig.
In summary, Collyria lacks all cephalic sclerites except for the pleurostoma, part of the labium and maxilla and the mandibles (at least in C. coxator). It might be noted that Short depicted the spiracle’s closing apparatus as extremely long and narrow. The spiracles in Short’s slide and the new specimens of catoptron are not nearly as long or as narrow (his depiction of the closing apparatus being separated from the atrium is accurate). Short apparently did not use a camera lucida or ocular grid, and his drawings are often strikingly distorted.
Cephalic sclerites of mature larva of Alomya semiflava Stephens
Fig.
The following description is based on a re-examination of two larval slides prepared by J.R.T. Short (
Cephalic sclerites with many prominent structures absent; remaining structures well-sclerotized. Epistomal suture completely sclerotized, uncertain if forming epistomal band because of distortion of larva, but coded as present (character 105, state 2). Labral sclerite absent (character 108, state 1); clypeolabral plates absent (character 130, state 0). Stipital sclerite absent (character 119, state 2). Pleurostoma well-sclerotized, not laterally expanded (character 106, state 0); posterior struts of inferior mandibular processes short and not connected by band (character 114, state 0). Hypostoma long and well-sclerotized (character 115, state 0), more or less straight, markedly angled ventrally towards cephalic midline. Hypostomal spur absent (character 117, state 2). Labial sclerite absent (character 124), originally coded as present, (circular to ovoid: state 1), but re-assessed when manuscript in press" (see comments, below). Region near salivary orifice distorted, but presumably U-shaped (character 133, state 1). Prelabial sclerite absent (character 127, state 0). Plate ventrad labial sclerite absent (character 129, state 0). Maxillary and labial palpi each bearing 5 sensilla (character 123, state 2). Mandible large, uniformly well-sclerotized (character 111, state 0) and triangular (character 109, state 0); blade without denticles (character 112, state 2). Antenna (character 131) not visible (coded as “?”). Spiracle with closing apparatus adjacent to atrium (character 132, state 1, but coded as separated from atrium, state 0). Skin smooth with scattered short setae.
Material examined: 1 last larval instar slide mount, larva reared under laboratory conditions from mummified final larval instar of Korscheltellus (= Hepialus) lupulinus (Linnaeus, 1758), 1979, R. Hinz (ANIC). 1 penultimate stage larval slide mount. Same data as final instar (ANIC).
Comments: Alomya semiflava, like other species in its genus, attack species of Hepialidae (Lepidoptera) (
1) There is no trace of the antennal disc.
2) The region of the epistomal suture is distorted, and it is not possible to determine if an epistomal band is present (it could be there but thin and not well sclerotized). The band is present in the penultimate larva slide, but ichneumonid larvae often lose structures upon maturity (
3) Short reported cuticular folds bearing setae on the clypeolabrum but these could not be seen. He apparently thought these to be analogous to the narrow clypeolabral plates found in Phaeogenini.
4) The maxillary apices are distorted and could not be reconstructed as depicted by Short. Both maxillary palpi are rotated so that only lateral views were possible (Fig.
5) The region of the salivary orifice is distorted and its shape cannot be determined (although it is presumably U-shaped). What Short depicts as the ‘silk press’ is the terminal end of the salivary duct.
6) Short shows the labial sclerite to be present, with the ventral section unsclerotized. The actual specimen has a crescentic structure on the right side in the vicinity of the labium (Fig.
7) The orientation of the epistoma+pleurostoma+hypostoma is difficult to determine, given the preparation’s distortion. The left-side hypostoma has broken off from the pleurostoma. The slide of the penultimate instar has the orientation of Short’s figure but the right side of the mature larval specimen belies that reconstruction. The actual arrangement is probably similar to that of Thyrateles procax (Cresson) or Trogus pennator (Fabricius) in
In summary, the cephalic morphology of the final-instar larva of A. semiflava is that of a standard ichneumonine, lacking only clypeolabral plates.
Taxa sequenced, countries of collection, specimen voucher numbers and Genbank accession numbers for molecular vouchers. All sequences were original to this study except as noted by superscripts indicating literature reference: 1
Taxa | Country of collection | Voucher depository | Voucher number | Genbank accession numbers | ||
---|---|---|---|---|---|---|
COI | 28s D2 | EF1a | ||||
Braconidae | ||||||
Doryctes erythromelas (Brullé) | ? | UKY | ? | GQ374627 1 | GQ374709 1 | GQ410706 1 |
Rhysipolis sp. | ? | UKY | ? | GQ374626 1 | GQ374708 1 | GQ410705 1 |
Aleiodes terminalis Cresson | ? | UKY | ? | – | GQ374710 1 | GQ410707 1 |
Aleiodes pictus (Herrich-Schäffer) | England | ? | ? | EF115464 2 | – | – |
Ichneumonidae | ||||||
Acaenitinae | ||||||
Spilopteron occiputale (Cresson) | United States | CNC | CNC 422320 | MK959483 | MK851161 | MK851398 |
Coleocentrus rufus Provancher | United States | CNC | CNC 422321 | MK959401 | MK851078 | MK851315 |
Adelognathinae | ||||||
Adelognathus sp. | United States | CNC | CNC 422322 | MK959374 | MK851051 | MK851288 |
Agriotypinae | ||||||
Agriotypus armatus Curtis | Czech Republic | CNC | CNC 422323 | MK959376 | MK851053 | MK851290 |
Alomyinae | ||||||
Alomya debellator (Fabricius) | Switzerland | CNC | CNC 422374 | MK959378 | MK851055 | MK851292 |
Anomaloninae | ||||||
Anomalonini | ||||||
Anomalon picticorne (Viereck) | United States | CNC | CNC 422324 | MK959379 | MK851056 | MK851293 |
Gravenhorstiini | ||||||
Therion texanum (Ashmead) | United States | CNC | CNC 422325 | MK959490 | MK851168 | MK851405 |
Ateleutinae | ||||||
Ateleute sp. nov. | United States | CNC | CNC 422344 | MK959384 | MK851061 | MK851298 |
Banchinae | ||||||
Atrophini | ||||||
Lissonota scutellaris (Cresson) | United States | CNC | CNC 422326: (COI, 28S D2); CNC 422489: (EF1a) | MK959436 | MK851113 | MK851350 |
Exetastes bioculatus Cresson | United States | CNC | CNC 422327 | MK959424 | MK851101 | MK851338 |
Glyptini | ||||||
Apophua simplicipes (Cresson) | Canada | CNC | CNC 422328 | MK959382 | MK851059 | MK851296 |
Sphelodon phoxopteridis (Weed) | United States | CNC | CNC 422329 | MK959482 | MK851160 | MK851397 |
Brachycyrtinae | ||||||
Brachycyrtus wardae Bennett | Fiji | CNC | CNC 422490: (COI, 28S D2); CNC 422330: (EF1a) | MK959389 | MK851066 | MK851303 |
Campopleginae | ||||||
Bathyplectes infernalis (Gravenhorst) | United States | CNC | CNC 422331 | MK959388 | MK851065 | MK851302 |
Campoletis sonorensis (Cameron) | United States | CNC | CNC 422332 | MK959391 | MK851068 | MK851305 |
Campoplex sp. | United States | CNC | CNC 422333 | MK959392 | MK851069 | MK851306 |
Casinaria grandis Walley | United States | CNC | CNC 422334 | MK959393 | MK851070 | MK851307 |
Dusona egregia (Viereck) | United States | CNC | CNC 422335 | MK959415 | MK851092 | MK851329 |
Hyposoter sp. | United States | CNC | CNC 422336 | MK959429 | MK851106 | MK851343 |
Olesicampe sp. | United States | CNC | CNC 422491: (COI, 28S D2); CNC 422337: (EF1a) | MK959452 | MK851129 | MK851366 |
Rhimphoctona macrocephala (Provancher) | Canada | CNC | CNC 422338 | MK959474 | MK851151 | MK851389 |
Claseinae | ||||||
Clasis sp. nov. | Chile | CNC | CNC 422339 | MK959398 | MK851075 | MK851312 |
Collyriinae | ||||||
Collyria catoptron Wahl | China | CNC | CNC 422340 | MK959402 | MK851079 | MK851316 |
Cremastinae | ||||||
Eiphosoma pyralidis Ashmead | United States | CNC | CNC 422341 | MK959418 | MK851095 | MK851332 |
Xiphosomella setoni Johnson | United States | CNC | CNC 422342 | MK959496 | MK851174 | MK851411 |
Cryptinae | ||||||
Aptesini | ||||||
Polytribax contiguus (Cresson) | Canada | CNC | CNC 422349 | MK959471 | MK851148 | MK851386 |
Rhytura pendens Townes | United States | CNC | CNC 422350 | MK959477 | MK851155 | MK851393 |
Cryptini | ||||||
Agonocryptus chichimecus (Cresson) | United States | CNC | CNC 422343 | MK959375 | MK851052 | MK851289 |
Baryceros texanus (Ashmead) | United States | CNC | CNC 422345 | MK959386 | MK851063 | MK851300 |
Diapetimorpha brunnea Townes | United States | CNC | CNC 422346 | MK959411 | MK851088 | MK851325 |
Echthrus reluctator (Linnaeus) | Germany | CNC | CNC 422348 | MK959416 | MK851093 | MK851330 |
Lymeon orbus (Say) | United States | CNC | CNC 422347 | MK959438 | MK851115 | MK851352 |
Ctenopelmatinae | ||||||
Ctenopelmatini | ||||||
Ctenopelma sanguineum (Provancher) | United States | CNC | CNC 422354 | MK959405 | MK851082 | MK851319 |
Xenoschesis limata (Cresson) | United States | CNC | CNC 422355 | MK959495 | MK851173 | MK851410 |
Euryproctini | ||||||
Euryproctus sentinis Davis | United States | CNC | CNC 422356 | MK959423 | MK851100 | MK851337 |
Mesoleptidea decens (Cresson) | United States | CNC | CNC 422357 | MK959443 | MK851120 | MK851357 |
Mesoleiini | ||||||
Barytarbes honestus (Cresson) | United States | CNC | CNC 422358 | MK959387 | MK851064 | MK851301 |
Himerta luteofacia Leblanc | United States | CNC | CNC 422359 | MK959427 | MK851104 | MK851341 |
Perilissini | ||||||
Lathrolestes asperatus Barron | United States | CNC | CNC 422360 | MK959433 | MK851110 | MK851347 |
Perilissus concolor (Cresson) | United States | CNC | CNC 422361 | MK959461 | MK851138 | MK851375 |
Pionini | ||||||
Rhorus bartelti Luhman | Canada | CNC | CNC 422362 | – | MK851152 | MK851390 |
Sympherta fucata (Cresson) | United States | CNC | CNC 281120 | MK959487 | MK851165 | MK851402 |
Scolobatini | ||||||
Onarion sp. | Bolivia | CNC | CNC 422365 | MK959453 | MK851130 | MK851367 |
Seleucini | ||||||
Seleucus cuneiformis Holmgren | Japan | CNC | CNC 422364 | MK959479 | MK851157 | MK851395 |
Westwoodiini | ||||||
Westwoodia sp. | Australia | CNC | CNC 422366 | MK959493 | MK851171 | MK851408 |
Tribe indet. | ||||||
Ctenopelmatinae Genus NZ | New Zealand | CNC | CNC 422367 | MK959406 | MK851083 | MK851320 |
Cyllocerinae | ||||||
Cylloceria melancholica (Gravenhorst) | United States | CNC | CNC 422368 | MK959409 | MK851086 | MK851323 |
Diacritinae | ||||||
Diacritus incompletus Momoi | Japan | CNC | CNC 422369 | MK959410 | MK851087 | MK851324 |
Diplazontinae | ||||||
Diplazon laetatorius (Fabricius) | Canada | CNC | CNC 422370 | MK959413 | MK851090 | MK851327 |
Woldstedtius flavolineatus (Gravenhorst) | United States | CNC | CNC 422371 | MK959494 | MK851172 | MK851409 |
Eucerotinae | ||||||
Euceros sp. nov. | United States | CNC | CNC 422372 | MK959422 | MK851099 | MK851336 |
Hybrizontinae | ||||||
Hybrizon rileyi (Ashmead) | United States | CNC | CNC 422373: (COI, 28S D2); CNC 422392: (EF1a) | MK959428 | MK851105 | MK851342 |
Ichneumoninae | ||||||
Heresiarchini | ||||||
Coelichneumon eximius (Stephens) | United States | CNC | CNC 422378 | MK959400 | MK851077 | MK851314 |
Protichneumon grandis (Brullé) | United States | CNC | CNC 422379 | MK959473 | MK851150 | MK851388 |
Ichneumonini | ||||||
Barichneumon neosorex Heinrich | United States | CNC | CNC 422380 | MK959385 | MK851062 | MK851299 |
Cratichneumon w-album (Cresson) | United States | CNC | CNC 422381 | MK959403 | MK851080 | MK851317 |
Orgichneumon calcatorius (Thunberg) | United States | CNC | CNC 422382 | MK959455 | MK851132 | MK851369 |
Patrocloides montanus (Cresson) | United States | CNC | CNC 422383 | MK959459 | MK851136 | MK851373 |
Joppocryptini | ||||||
Plagiotrypes concinnus (Say) | United States | CNC | CNC 422384 | MK959468 | MK851145 | MK851382 |
Listrodromini | ||||||
Dilopharius otomitus (Cresson) | United States | CNC | CNC 422385 | MK959412 | MK851089 | MK851326 |
Phaeogenini | ||||||
Centeterus euryptychiae (Ashmead) | Canada | CNC | CNC 422375 | MK959394 | MK851071 | MK851308 |
Phaeogenes hebrus (Cresson) | United States | CNC | CNC 422376 | MK959464 | MK851141 | MK851378 |
Stenodontus sp. nov. | United States | CNC | CNC 422377 | MK959484 | MK851162 | MK851399 |
Platylabini | ||||||
Cyclolabus impressus (Provancher) | United States | CNC | CNC 422386 | MK959408 | MK851085 | MK851322 |
Linycus exhortator (Fabricius) | United States | CNC | CNC 422387 | MK959435 | MK851112 | MK851349 |
Labeninae | ||||||
Orthognathelini | ||||||
Grotea anguina Cresson | United States | CNC | CNC 422388 | MK959426 | MK851103 | MK851340 |
Labium sp. | Australia | CNC | CNC 422389 | MK959432 | MK851109 | MK851346 |
Labenini | ||||||
Apechoneura sp. | Bolivia | CNC | CNC 422390 | MK959380 | MK851057 | MK851294 |
Labena grallator (Say) | United States | CNC | CNC 422391 | MK959431 | MK851108 | MK851345 |
Poecilocryptini | ||||||
Poecilocryptus nigromaculatus Cameron | Australia | ? (28s); CNC (EF1a) | ? (28s); CNC 422392 (EF1a) | – | AJ302921 3 | MK851383 |
Lycorininae | ||||||
Lycorina glaucomata (Cushman) | United States | CNC | CNC 422393 | MK959437 | MK851114 | MK851351 |
Mesochorinae | ||||||
Astiphromma sp. nov. | United States | CNC | CNC 422394 | MK959383 | MK851060 | MK851297 |
Chineater masneri Wahl | Chile | CNC | CNC 422395 | MK959395 | MK851072 | MK851309 |
Cidaphus paniscoides (Ashmead) | United States | CNC | CNC 422396 | MK959397 | MK851074 | MK851311 |
Lepidura collaris Townes | Chile | CNC | CNC 422397 | MK959434 | MK851111 | MK851348 |
Mesochorus sp. | United States | CNC | CNC 422398 | MK959442 | MK851119 | MK851356 |
Metopiinae | ||||||
Exochus semirufus Cresson | United States | CNC | CNC 422399 | MK959425 | MK851102 | MK851339 |
Metopius pollinctorius (Say) | United States | CNC | CNC 422400 | MK959444 | MK851121 | MK851358 |
Scolomus sp. | Chile | CNC | CNC 422401 | MK959478 | MK851156 | MK851394 |
Seticornuta terminalis (Ashmead) | United States | CNC | CNC 422402 | MK959480 | MK851158 | MK851396 |
Microleptinae | ||||||
Microleptes sp. | South Korea | CNC | CNC 422403 | MK959445 | MK851122 | MK851359 |
Neorhacodinae | ||||||
Neorhacodes enslini (Ruschka) | Spain | CNC | CNC 422434 | MK959446 | MK851123 | MK851360 |
Nesomesochorinae | ||||||
Chriodes sp. | Madagascar | CNC | CNC 422404 | MK959396 | MK851073 | MK851310 |
Nonnus sp. | Argentina | CNC | CNC 422405 | MK959449 | MK851126 | MK851363 |
Ophioninae | ||||||
Enicospilus flavostigma Hooker | United States | CNC | CNC 422406 | MK959420 | MK851097 | MK851334 |
Hellwigia obscura Gravenhorst | France | ? | ? | – | AJ302858 4 | – |
Ophion sp. | United States | CNC | CNC 422407 | MK959454 | MK851131 | MK851368 |
Skiapus sp. | Mozambique | CNC | CNC 422408 | MK959481 | MK851159 | – |
Thyreodon sp. | Guyana | CNC | CNC 422409 | MK959492 | MK851170 | MK851407 |
Orthocentrinae | ||||||
Megastylus sp. nov. | United States | CNC | CNC 422410 | MK959441 | MK851118 | MK851355 |
Orthocentrus sp. | United States | CNC | CNC 422411 | MK959456 | MK851133 | MK851370 |
Proclitus speciosus Dasch | United States | CNC | CNC 422412 | MK959472 | MK851149 | MK851387 |
Orthopelmatinae | ||||||
Orthopelma mediator (Thunberg) | Sweden | CNC | CNC 422413 | MK959457 | MK851134 | MK851371 |
Oxytorinae | ||||||
Oxytorus albopleuralis (Provancher) | United States | CNC | CNC 422414 | MK959458 | MK851135 | MK851372 |
Pedunculinae | ||||||
Pedunculus sp. nov. | Chile | CNC | CNC 422415 | MK959460 | MK851137 | MK851374 |
Phygadeuontinae | ||||||
Acrolyta sp. | United States | CNC | CNC 422351 | MK959372 | MK851049 | MK851286 |
Endasys patulus (Viereck) | United States | CNC | CNC 422352 | MK959419 | MK851096 | MK851333 |
Mastrus sp. | United States | CNC | CNC 422353 | MK959439 | MK851116 | MK851353 |
Pimplinae | ||||||
Delomeristini | ||||||
Perithous divinator (Rossi) | Canada | CNC | CNC 422416 | MK959462 | MK851139 | MK851376 |
Ephialtini | ||||||
Acrotaphus wiltii (Cresson) | United States | CNC | CNC 422417 | MK959373 | MK851050 | MK851287 |
Clistopyga recurva (Say) | United States | CNC | CNC 422418 | MK959399 | MK851076 | MK851313 |
Dolichomitus irritator (Fabricius) | United States | CNC | CNC 422419 | MK959414 | MK851091 | MK851328 |
Zaglyptus pictilis Townes | United States | CNC | CNC 422420 | MK959498 | MK851176 | MK851413 |
Pimplini | ||||||
Pimpla annulipes Brullé | Canada | CNC | CNC 422421 | MK959467 | MK851144 | MK851381 |
Theronia bicincta (Cresson) | United States | CNC | CNC 422422 | MK959491 | MK851169 | MK851406 |
Poemeniinae | ||||||
Neoxorides caryae (Harrington) | United States | CNC | CNC 422423 | MK959447 | MK851124 | MK851361 |
Poemenia albipes (Cresson) | United States | CNC | CNC 422424 | MK959469 | MK851146 | MK851384 |
Rhyssinae | ||||||
Megarhyssa greenei Viereck | Canada | CNC | CNC 422425 | MK959440 | MK851117 | MK851354 |
Rhyssa crevieri (Provancher) | Canada | CNC | CNC 422426 | MK959475 | MK851153 | MK851391 |
Rhyssella nitida (Cresson) | Canada | CNC | CNC 422427 | MK959476 | MK851154 | MK851392 |
Sisyrostolinae | ||||||
Brachyscleroma sp. | Kenya | CNC | CNC 422428 | MK959390 | MK851067 | MK851304 |
Erythrodolius calamitosus Seyrig | Madagascar | CNC | CNC 422429 | MK959421 | MK851098 | MK851335 |
Stilbopinae | ||||||
Notostilbops sp. nov. | Chile | CNC | CNC 422430 | MK959450 | MK851127 | MK851364 |
Stilbops vetulus (Gravenhorst) | Hungary | CNC | CNC 422431 | MK959486 | MK851164 | MK851401 |
Tatogastrinae | ||||||
Tatogaster nigra Townes | Chile | CNC | CNC 422432 | MK959488 | MK851166 | MK851403 |
Tersilochinae | ||||||
Allophrys divaricata Horstmann | United States | CNC | CNC 422493: (COI, 28S D2); CNC 422433: (EF1a) | MK959377 | MK851054 | MK851291 |
Peucobius fulvus Townes | United States | CNC | CNC 422435 | MK959463 | MK851140 | MK851377 |
Phrudus sp. | South Korea | CNC | CNC 422436 | MK959465 | MK851142 | MK851379 |
Stethantyx nearctica Townes | United States | CNC | CNC 422437 | MK959485 | MK851163 | MK851400 |
Tersilochus sp. | United States | CNC | CNC 422438: (COI, EF1a); CNC 422494: (28s D2) | MK959489 | MK851167 | MK851404 |
Tryphoninae | ||||||
Eclytini | ||||||
Eclytus sp. | United States | CNC | CNC 422495: (COI, 28S D2); CNC 422439: (EF1a) | MK959417 | MK851094 | MK851331 |
Idiogrammatini | ||||||
Idiogramma longicauda (Cushman) | United States | CNC | CNC 422440 | MK959430 | MK851107 | MK851344 |
Oedemopsini | ||||||
Zagryphus nasutus (Cresson) | United States | CNC | CNC 422441: (COI, EF1a); CNC 422496: (28s D2) | MK959499 | MK851177 | MK851414 |
Phytodietini | ||||||
Netelia sp. | United States | CNC | CNC 422442 | MK959448 | MK851125 | MK851362 |
Phytodietus vulgaris Cresson | United States | CNC | CNC 422443 | MK959466 | MK851143 | MK851380 |
Tryphonini | ||||||
Cteniscus sp. | United States | CNC | CNC 422444 | MK959404 | MK851081 | MK851318 |
Cycasis rubiginosa (Gravenhorst) | Switzerland | CNC | CNC 422445 | MK959407 | MK851084 | MK851321 |
Polyblastus sp. | United States | CNC | CNC 422446 | MK959470 | MK851147 | MK851385 |
Xordinae | ||||||
Aplomerus sp. | United States | UKY | CNC 681999 (COI, 28s D2), CNC | MK959381 | MK851058 | MK851295 |
CNC | 422447: (EF1a) | |||||
Odontocolon albotibiale (Bradley) | United States | CNC | CNC 422497: (COI, 28s D2); CNC 422448: (EF1a) | MK959451 | MK851128 | MK851365 |
Xorides stigmapterus (Say) | Canada | CNC | CNC 422449 | MK959497 | MK851175 | MK851412 |
Ichneumonidae complete phylogenetic matrix, molecular characters coded as 0123
Data type: phylogenetic data (nexus format)
Ichneumonidae complete phylogenetic matrix, molecular characters coded as 0123
Data type: phylogenetic data (Hennig86 format)
Ichneumonidae phylogenetic matrix, molecular characters only, coded as ACGT
Data type: phylogenetic data (nexus format)
Ichneumonidae phylogenetic matrix, molecular characters only, coded as ACGT
Data type: phylogenetic data (Hennig86 format)
Total evidence parsimony trees
Data type: Phylogenetic trees (Winclada format)
Morphology-only parsimony trees
Data type: Phylogenetic trees (Winclada format)
Molecular-only parsimony trees
Data type: Phylogenetic trees (Winclada format)
Bayesian total evidence trees
Data type: Phylogenetic trees (Winclada format)
Bayesian molecular-only trees
Data type: Phylogenetic trees (Winclada format)