Deuteragenia Šustera, 1912.

The female possesses maxillary cardo with two tufts of curved bristles. The male with flagellomeres serrate beneath, the propodeum somewhat swollen and punctate, third submarginal cell of fore wing usually equal or slightly larger than second submarginal cell.

Currently the tribe includes the following six genera: Deuteragenia Šustera, 1912; Dipogon Fox, 1897; Myrmecodipogon Ishikawa, 1965; Nipponodipogon Ishikawa, 1965; Stigmatodipogon Ishikawa, 1965; and Winnemanella Krombein, 1962.

World-wide, except Australia (Lelej and Loktionov 2012).

Lelej and Loktionov (2012) analyzed 13 species from six generic groups Deuterageniini, with the genus Priocnemis Schiødte, 1837 as an outgroup. At that time the tribe comprised only the genus Dipogon with six subgenera. The analysis revealed the following relationships: Priocnemis + [Stigmatodipogon + {(Deuteragenia + Mesagenia) + [(Winnemanella + Nipponodipogon) + (Myrmecodipogon + Dipogon)]}]. As a result of this study, a new generic classification of the tribe was proposed, where Deuteragenia, Myrmecodipogon, Nipponodipogon, Stigmatodipogon, and Winnemanella were elevated to generic status. The females of Deuterageniini differ clearly from those of the tribes Ageniellini and Priocnemini, while male characters of Deuterageniini overlap with those of other tribes of the subfamily Pepsinae. For detailed information on the tribe see Lelej and Loktionov (2012).

Deuteragenia variegata (Linnaeus, 1758) [Sphex], by automatic designation.

The female and the male possess the following characters: fore wings with a basal or apical fascia; ratio of maximum width to maximum length of second radio-medial cell 2.0 times or less; the pterostigma normal-sized, its width similar to width of second submarginal cell (usually narrow); length of hind wing anal lobe more than 0.33 times width of submedial cell; the antenna elongated, F1 length more than 3.5 times (in female) and more than 3.0 times (in male) its width; mandible stout, with three teeth, including apical one.

56 species.

World-wide, except Australia (Lelej and Loktionov 2012). The genus has not been previously known from the Indonesian island of Sumatra.

For detailed information on the genus see Shimizu and Ishikawa (2002a), and Lelej and Loktionov (2012).

Holotype : female, “W Sumatra Padang Panjang XII.2003 St. Jakl leg.” [Indonesia, West Sumatra Prov., City of Padang Panjang] [OLL].

The female of this new species is unique among congeners of Deuteragenia in having bifid tarsal claws (Fig. 2E). Other characters of importance are: propodeum medial and posterior portion with coarse transverse rugae; clypeus anterior rim smooth and polished, not depressed, and not differentiated from dorsal portion (Fig. 2A); head and mesosoma mostly without setae; T1 petiolate basally (Fig. 2D); F2–F10 pale ventrally; legs partially brown (Fig. 1). The male is unknown.

Female, holotype (Figs 1, 2). Length: body 8.4 mm, fore wing 7.4 mm. Head width 1.21 times its height; MID 0.51 times head width in frontal view (Fig. 2A). Ocelli large, well raised; ocellar triangle slightly acute-angled; POD : OOD = 0.87 (Fig. 2C). Head in frontal view with vertex barely produced above dorsal eye margin (Fig. 2A). Posterior margin of vertex in dorsal view hardly concave (Fig. 2C). Head with frons almost flat in lateral view. Gena in profile well developed (Fig. 2B). Malar space very short. Clypeus moderately convex, its width 2.6 times its height, and 1.0 times LID; anterior margin straight, not depressed and not differentiated from dorsal portion; anterolateral corner rounded (Fig. 2A). Labrum not exposed. Bristles of maxillary cardo hard and long, reaching mandible ventral face. Antenna elongated; flagellomeres cylindrical; ratio of scape, pedicel and flagellomeres length (on dorsal side) 25 : 8 : 39 : 25 : 20 : 17 : 17 : 17 : 15 : 15 : 14 : 16; scape length 0.8 times UID; F1 length 5.0 times its maximum width (in dorsal view), and 1.27 times UID; apical flagellomere pointed apically.

Figure 1. 

Habitus of Deuteragenia leleji sp. nov., holotype, female: A lateral view B dorsal view. Scale bars: 1.0 mm.

Pronotum length 0.42 times its maximum width in dorsal view; anterior face not differentiated from dorsum; posterior margin moderately rounded (Fig. 1B). Dorsum of mesoscutum hardly convex. Dorsum of mesoscutellum and metanotum slightly convex like dorsum of propodeum in lateral view. Metapostnotum noticeably depressed, barely arcuately emarginated postero-medially, its length 0.15 times metanotum length medially. Propodeum somewhat elongated, its length 1.05 times its maximum width in dorsal view; dorsum and posterior face not differentiated from each other and evenly convex (Fig. 2D).

Figure 2. 

Deuteragenia leleji sp. nov., holotype, female: A head in frontal view B head in lateral view C head in dorsal view D mesoscutellum, metanotum, metapostnotum, propodeum and T1 in dorsal view E metaclaws F fore wing G hind wing. Scale bars: 1.0 mm (F, G); 0.5 mm (D); 0.2 mm (A–C); 0.1 mm (E).

All femora without spines. Protibia ventral face apically with few short spines. Meso- and metatibia dorsal and outer-lateral face with scattered short spines. Tarsomeres 1–3 of all legs with very short spines ventrally; tarsomeres 4 and 5 of all legs without spines ventrally. Tarsal claws of all legs symmetrical and bifid; inner tooth as long as apical one, very broad at base, and narrowing apically, with pointed apex; outer margins of inner and apical teeth parallel (Fig. 2E).

Fore wing (Fig. 2F) translucent, with subapical and subbasal brown fasciae. Pterostigma brown, its length 4.75 times its height (on inner distance), and 5.2 times Rs₂. Second submarginal cell narrowed on vein Rs by 0.77 times its own length on vein M, receiving crossvein 1m-cu at basal 0.55. Third submarginal cell narrowed on vein Rs by 0.43 times its own length on vein M, receiving crossvein 2m-cu at basal 0.3. Crossvein 2rs-m slightly bent near M. Crossvein 3rs-m weakly arcuate. Crossvein cu-a straight, originating beyond separation of vein M+CuA. Vein M touching wing margin. Hind wing (Fig. 2G) translucent, with slightly brownish apical portion.

Metasoma with T1 distinctly petiolate (Fig. 2D).

Head , meso- and metasoma matt and punctate. Clypeus anterior rim smooth and polished, with other part densely punctate (Fig. 2A). Mandible apical portion polished. Frons finely and densely punctate, median line distinct. Metapostnotum polished, with indistinct transverse striae. Propodeum (Fig. 2D) anterior portion with dense and coarse merging punctures, its medial and posterior portions with coarse transverse rugae which somewhat arcuate posteriorly. Antenna and legs matt.

Body black (Figs 1, 2). Antenna black, with F2–F10 pale ventrally. Mandible partially brown apically. Bristles of maxillary cardo brown. Legs black, with following brown: all coxae and trochanters, fore- and mesofemora, foretibia partially, metafemure except apical portion, all claws.

Body without setae except following: upper frons along inner orbits with one long setae; gena with short scattered setae; all coxae anterior face and pronotum with scattered short setae; propodeum postero-laterally with scattered gray short setae; T6 and S6 with long and dense pale setae. Body with sparse gray pubescence most intensive on propodeum postero-laterally.

Male is unknown.

Indonesia: West Sumatra.

The specific epithet is a pathronym honoring Prof. Arkady Lelej (FSC Biodiversity FEB RAS, Vladivostok, Russia), my scientific advisor and inspirer, on the occasion of his 75^th birthday.