expand article infoMaxim Yu. Proshchalykin, Vladimir E. Gokhman§
‡ Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, Russia
§ Moscow State University, Moscow, Russia
Open Access

On August 25, 2021, Arkady S. Lelej, an outstanding Russian entomologist and a very fine person, turns 75. In addition to their warmest regards to him, Arkady’s colleagues and friends prepared a Special Issue of the Journal of Hymenoptera Research on this special occasion. 53 authors from Belgium, China, Czech Republic, Estonia, Finland, Germany, Italy, Japan, the Netherlands, Trinidad and Tobago, Ukraine, United Kingdom, USA, and Russia (Feodosiya, Kaliningrad, Moscow, Nizhny Novgorod, Petrozavodsk, Saratov, St. Petersburg, and Vladivostok) took part in preparing this Issue. They have jointly presented 24 papers dedicated to various aspects of Hymenoptera studies, including descriptions of five insect genera, 28 species and one subspecies as new to science. Among them, 18 new taxa (three genera and 15 species) are named in Arkady’s honor.

Figure 1. 

9th International Congress of Hymenopterists. Ehime University, Matsuyama, Japan, 23-27 July 2018. Arkady S. Lelej presents his talk at the Congress. 26 July 2018.

Most papers in this Special Issue are dedicated to taxonomy and faunistics of various hymenopteran taxa. These include members of all traditional major subdivisions of the order, i.e., Symphyta, Parasitica and Aculeata. Among the extant taxa, there are members of the West Palaearctic Euura bergmanni and oligospila species groups (Tenthredinidae) (Prous et al. 2021), Ichneumonidae from the Eastern Palearctic and Oriental regions (Humala 2021; Kasparyan and Khalaim 2021), Afrotropical members of the braconid genus Sculptolobus Yang et al., 2008 (Samartsev and van Achterberg 2021), chalcid wasps of the genus Toxeuma Walker, 1833 (Pteromalidae) of the Russian fauna (Tselikh and Dale-Skey 2021), flat wasps of the genus Sierola Cameron, 1881 (Bethylidae) from China (Wang et al. 2021), and a new species of the genus Odontosphex Arnold, 1951 (Crabronidae) from Central Asia (Mokrousov and Proshchalykin 2021), a new species of the genus Anteon Jurine, 1807 (Dryinidae) from Turkey (Speranza et al. 2021) as well as cuckoo wasps (Chrysididae) from Pakistan and India (Rosa et al. 2021), two bee genera from the Caucasus, Mongolia, Middle East and North Africa (Astafurova and Proshchalykin 2021; Proshchalykin and Dathe 2021; Bogusch 2021) and new genera and species of spider wasps (Pompilidae) and velvet ants (Mutillidae) from the Oriental Region (Shimizu et al. 2021; Loktionov 2021; Okayasu 2021).

In addition, a few extinct groups were described, mostly from Baltic, Rovno and Sakhalinian ambers. These taxa include braconid parasitoids of ants from the tribe Neoneurini (Belokobylskij et al. 2021) as well as new genera and species of Encyrtidae (Simutnik et al. 2021) and Formicidae (Radchenko and Proshchalykin 2021). Moreover, a new genus and species of the extinct family Peleserphidae (Proctotrupoidea) are described by Rasnitsyn (2021).

In a single morphological study, the ultrastructure of unusual placoid sensilla of miniature Megaphragma Timberlake, 1924 wasps (Trichogrammatidae) was examined with various methods of electron microscopy (Diakova and Polilov 2021).

Several papers are dedicated to the bionomics of certain groups of Hymenoptera, including details of parasitism by two Eupelmus Dalman, 1820 species (Eupelmidae) on Aulacidea hieracii (Bouché, 1834) (Cynipidae) (Gokhman and Nikelshparg 2021), additional nesting data for Leptochilus regulus (de Saussure, 1855) (Vespidae) (Fateryga and Fateryga 2021) as well as defensive adaptations of Mutillidae (Schmidt et al. 2021) and caste and wing loading in Dolichovespula maculata (Linnaeus, 1763) (Vespidae) (Starr et al. 2021).

We celebrated Arkady’s 70th anniversary just five years ago. On that occasion, a Special Issue of the Euroasian Entomological Journal was published (http://www.eco.nsc.ru/eej/supplement_2016_15_1.html) . In this Issue, Arkady’s biography, together with a full list of his published papers and taxa described by him, is given. Nevertheless, Arkady did not decrease his productivity since that time. He continues to work intensively, coupling research with organizational activities. Arkady continues to lead the Laboratory of Entomology of the Federal Scientific Center of the East Asia Terrestrial Biodiversity in Vladivostok. He is the Head of the Publishing Commission, and a member of both the Scientific Council and Dissertation Panel of the Center. During the last five years, Arkady led three projects funded by the Russian Foundation for Basic Research (RFBR), including IV Eurasian Symposium on Hymenoptera (September 9-15, 2019, Vladivostok). Starting from 2017, he published more than 50 papers dedicated to various hymenopteran taxa as well as to members of other insect orders, in addition to about 350 papers published during the previous years.

Figure 2. 

9th International Congress of Hymenopterists. Participants from Russia (left to right): Andrey I. Khalaim, Arkady S. Lelej, Ekaterina V. Tselikh, Vladimir E. Gokhman. 23 July 2018.

List of selected papers authored by Arkady Lelej (2017 – June 2021)


Brothers DJ, Lelej AS (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research 60: 1–97.

Zhou H-T, Lelej AS, Xu Z-F (2017) Discovery of the subfamily Rhopalomutillinae (Hymenoptera: Mutillidae) in China and description of Pherotilla brothersi sp. nov. Zootaxa 4273(4): 587–594.

Lelej AS, Williams KA, Loktionov VM, Pang H, Xu Z (2017) Review of the genus Zeugomutilla Chen, 1957 (Hymenoptera, Mutillidae, Mutillini), with description of two new species. Zootaxa 4247(1): 1–15.

Lelej AS, Proshchalykin MYu, Loktionov VM (Eds) (2017) Annotated catalog of the Hymenoptera of Russia. Volume I. Symphyta and Aculeata. Proceedings of the Zoological Institute RAS, Supplement 6: 5–475.

Loktionov VM, Lelej AS, Xu Z-F (2017) Review of the Priocnemis hyalinata species-group (Hymenoptera: Pompilidae), with description of a new species from China. Zootaxa 4254: 551–562.

Loktionov VM, Lelej AS, Xu Z-F (2017) Review of the genus Eopompilus Gussakovskij, 1932 (Hymenoptera: Pompilidae) with the description of new species from China. Zootaxa 4277: 413–426.

Loktionov VM, Lelej AS, Xu Z-F (2017) Discovery of the genus Nipponodipogon Ishikawa in the Oriental region, with description of two new species from China (Hymenoptera, Pompilidae). ZooKeys 692: 103–127.

Sakhvon VV, Lelej AS (2017) Review of the genus Habropogon Loew, 1847 (Diptera: Asilidae) from Russia, with description of new species from Siberia. Zootaxa 4311(3): 417–425.

Lelej AS, Mokrousov MV (2017) The types of Scoliidae (Hymenoptera), described by Eduard Eversmann, with some taxonomic notes and checklist of Russian Scoliidae. Far Eastern Entomologist 340: 1–17.

Rosa P, Lelej AS, Proshchalykin MYu, Loktionov VM, Mokrousov MV (2017) New records of Chrysididae (Hymenoptera) from Russia with description of five new species. Far Eastern Entomologist 345: 1–33.


Barthélémy C, Lelej AS, Loktionov VM (2018) A contribution to the Mutillidae (Hymenoptera) of Hong Kong. Zootaxa 4514(2): 230–242.

Zhou H-T, Lelej AS, Williams KA, Liu J-X (2018) Revision of the Oriental genus Zavatilla Tsuneki (Hymenoptera, Mutillidae, Trogaspidiini), with descriptions of two new species. Zootaxa 4418(2): 101–120.

Okayasu J, Williams KA, Lelej AS (2018) A remarkable new species of Sinotilla Lelej (Hymenoptera: Mutillidae: Smicromyrmini) from Taiwan and an overview of color diversity in East Asian mutillid females. Zootaxa 4446(3): 301–324.

Mokrousov MV, Lelej AS, Fadeev KI (2018) New data on the Palaearctic wasps of the genus Sierolomorpha Ashmead, 1903 (Hymenoptera, Sierolomorphidae) with description of S. trjapitzini sp. n. from Primorskii Territory, Russia. Entomological Review 98(6): 737–742.

Loktionov VM, Lelej AS (2018) Discovery of the spider wasp genus Sphictostethus Kohl, 1884 (Hymenoptera: Pompilidae: Pepsinae) in New Guinea, with description of two new species. Zootaxa 4429: 295–302.

Loktionov VM, Lelej AS (2018) To the knowledge of genus Nipponodipogon Ishikawa, 1965 (Hymenoptera: Pompilidae, Pepsinae) from Laos. Far Eastern Entomologist 362: 1–7.

Loktionov VM, Lelej AS, Liu J-X (2018) Contribution to the knowledge on the spider wasps genus Morochares Banks, 1934 (Hymenoptera: Pompilidae) from China. Zootaxa 4462: 497–510.

Mokrousov MV, Lelej AS, Gorbatovsky VV (2018) The types of described by Eduard Eversmann Tiphiidae and Sierolomorphidae (Hymenoptera) with taxonomic notes. Far Eastern Entomologist 349: 11–16.

Zhou H-T, Lelej AS, Liu J-X (2018) New records of velvet ants (Hymenoptera: Mutillidae) from China with taxonomic notes. Far Eastern Entomologist 354: 23–28.

Shimizu A, Lelej AS, Loktionov VM, Nishimoto Y, Endo T (2018) Revision of the subgenus Stigmatodipogon Ishikawa of the genus Dipogon Fox (Hymenoptera: Pompilidae: Pepsinae). Zootaxa 4514: 1–22.

Wahis R, Lelej AS, Loktionov VM (2018) Contribution to the knowledge of the genus Eopompilus Gussakovskij, 1932 (Hymenoptera, Pompilidae). Far Eastern Entomologist 361: 1–11.

Sakhvon VV, Lelej AS (2018) Review of the genus Heteropogon Loew, 1847 (Diptera: Asilidae) from Russia, and Central Asia, with description of two new species. Zootaxa 4486(4): 435–450.


Achterberg C, van, Sundukov YuN, Lelej AS, Belokobylskij SA (2019) To the knowledge of Gasteruptiidae (Hymenoptera) from Russian Far East. Far Eastern Entomologist 381: 1–8.

Brothers DJ, Lelej AS, Williams KA (2019) Genus-group names of Mutillidae (Hymenoptera): corrections and updates since 2008. Zootaxa 4651(3): 578–588.

Kochetkov DN, Olmi M, Lelej AS (2019) Contribution to the knowledge of dryinid wasps genus Anteon Jurine, 1807 (Hymenoptera: Dryinidae) in the Russian Far East. Far Eastern Entomologist 323: 11–17.

Williams KA, Lelej AS, Thaochan N (2019) New species of Myrmosinae (Hymenoptera: Mutillidae) from Southeastern Asia. Zootaxa 4656(3): 525–534.

Kumar GP, Lelej AS, Das D, Raveendran HKP, Loktionov VM (2019) Discovery of the genus Kudakrumia Krombein, 1979 (Hymenoptera: Mutillidae) in India and description of a new species. Zootaxa 4612(6): 260–266.

Loktionov VM, Lelej AS (2019) Contribution to the knowledge of the subgenus Stigmatodipogon Ishikawa, 1965 of the genus Dipogon Fox, 1897 (Hymenoptera: Pompilidae). Zootaxa 4613(2): 363–368.

Ilyasov RA, Youn HG, Lee M-l, Kim KW, Proshchalykin MYu, Lelej AS, Takahashi J-I, Kwon HW (2019) Phylogenetic relationships of Russian Far-East Apis cerana with other North Asian populations. Journal of Apicultural Science 63(2): 289–314.

Loktionov VM, Lelej AS, Liu J-X (2019) A new genus of spider wasps (Hymenoptera, Pompilidae) from China. Far Eastern Entomologist 376: 1–14.

Loktionov VM, Lelej AS (2019) The spider wasp genus Telostholus Haupt, 1929 (Hymenoptera, Pompilidae) in Indonesia, with the description of three new species. Zootaxa 4648(2): 260–272.


Engel MS, Lelej AS (2020) On the spelling of family-group names based on the genus Trigonalys Westwood (Hymenoptera: Trigonalidae versus Trigonalyidae). Entomologist’s Monthly Magazine 156(1): 58–60.

Gadallah NS, Lelej AS, Brothers DJ (2020) Biodiversity of the aculeate wasps (Hymenoptera: Aculeata) of the Arabian Peninsula: Vespoidea, Mutillidae. Zootaxa 4754(1): 141–152.

Lelej AS, Terine JB, Kumar GP, Das D, Sureshan PM (2020) Review of the genus Odontomutilla Ashmead, 1899 (Hymenoptera: Mutillidae) of the Indian subcontinent, with description of two new species. Zootaxa 4822(2): 191–208.

Lelej AS (2020) To the knowledge of velvet ants of the genera Artiotilla Invrea, Radoszkowskitilla Lelej and Taiwanomyrme Tsuneki (Hymenoptera: Mutillidae). Far Eastern Entomologist 410: 1–10.

Lelej AS (2020) Three new species of velvet ants from the tribe Smicromyrmini (Hymenoptera: Mutillidae) with unusual color syndrome. Far Eastern Entomologist 421: 1–9.

Pagliano G, Brothers DJ, Cambra R, Lelej AS, Lo Cascio P, Matteini Palmerini M, Scaramozzino PL, Williams KA, Romano M (2020) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali 36(1–2): 5–425.


Okayasu JA, Lelej AS, Williams KA (2021) Review of the genus Eotrogaspidia Lelej, 1996 (Hymenoptera: Mutillidae: Trogaspidiini). Zootaxa 4920(1): 56–90.

Sutanto DD, Williams KA, Nugroho H, Lelej AS (2021) To the knowledge of the velvet ant Genus Bischoffitilla Lelej (Hymenoptera: Mutillidae) in Indonesia. Treubia 48(1): 1–12.

In these works, 3 tribes, 1 subtribe, 1 genus, 41 species and 1 subspecies of Hymenoptera from 4 families are described in total as new to science. The whole list of the new taxa described by Arkady Lelej therefore includes 364 names.

List of the hymenopteran taxa described by Arkady Lelej (2017 – June 2021)

Family Mutillidae

Apteromutillini Brothers & Lelej, 2017

Ctenotillini Brothers & Lelej, 2017

Dasymutillini Brothers & Lelej, 2017

Euspinoliina Brothers & Lelej, 2017

Eotrogaspidia adhabar Okayasu, Lelej & Williams, 2021

Eotrogaspidia buddha Okayasu, Lelej & Williams, 2021

Eotrogaspidia hauseri Okayasu, Lelej & Williams, 2021

Eotrogaspidia melanopleura Okayasu, Lelej & Williams, 2021

Ephucilla hejunhuai Lelej, 2020

Krombeinella brothersi Williams, Lelej & Thaochan, 2019

Kudakrumia loaibonho Williams, Lelej & Thaochan, 2019

Kudakrumia malaenglek Williams, Lelej & Thaochan, 2019

Kudakrumia rangnekari Girish Kumar & Lelej, 2019

Odontomutilla fletcheri Lelej, Terine & Girish Kumar, 2020

Odontomutilla sairandhriensis Terine, Girish Kumar & Lelej, 2020

Pherotilla brothersi Zhou, Lelej & Xu, 2017

Promecidia abnormis Lelej, 2016

Promecidia chui Lelej et Xu, 2016

Promecilla ovtchinnikovi Lelej, 2020

Sinotilla nigrothoracica Okayasu, Williams & Lelej, 2018

Smicromyrme burgeri Lelej, 2020

Taiwanomyrme cheni Lelej, 2020

Zavatilla nepalensis Zhou et Lelej, 2018

Zavatilla xuzaifui Zhou, Lelej et Williams, 2018

Zeugomutilla pangi Lelej, 2017

Family Pompilidae

Eopriocnemis Loktionov & Lelej, 2019

Dipogon lao Loktionov & Lelej, 2018

Dipogon wasbaueri Loktionov & Lelej, 2019

Dipogon siam Loktionov & Lelej, 2019

Eopompilus pseudominor Loktionov, Lelej & Xu, 2017

Eopriocnemis albiflagellata Loktionov & Lelej, 2019

Morochares sinica Loktionov & Lelej, 2018

Morochares wahisi Loktionov & Lelej, 2018

Morochares wolfi Loktionov & Lelej, 2018

Morochares xuzaifui Loktionov, Lelej & Liu, 2018

Nipponodipogon gusenleitnerorum Loktionov & Lelej, 2018

Nipponodipogon orientalis Loktionov, Lelej & Xu, 2017

Nipponodipogon shimizui Loktionov, Lelej & Xu, 2017

Priocnemis yunnanensis Loktionov, Lelej & Xu, 2017

Sphictostethus arfak Loktionov & Lelej, 2018

Sphictostethus papua Loktionov & Lelej, 2018

Stigmaporus volgadon Loktionov & Lelej, 2016

Telostholus celebes Loktionov & Lelej, 2019

Telostholus rinjani Loktionov & Lelej, 2019

Telostholus sulawesi Loktionov & Lelej, 2019

Family Apidae

Apis cerana ussuriensis Ilyasov, Takahashi, Proshchalykin, Lelej & Kwon, 2019

Family Sierolomorphidae

Sierolomorpha trjapitzini Mokrousov & Lelej, 2018

In addition to his talent as a scientist, Arkady possesses a number of other remarkable features. First, he is kind and respectful to all surrounding people, including colleagues at work and people on the street. His wonderful politeness and ability to avoid conflicts, i.e., features which are definitely rare and needed nowadays, allow him to quickly establish connections with unfamiliar people who are always positively impressed by their first meeting with him. We know that Arkady’s personality and proficiency in insect collecting, are highly valued by his colleagues (in total, 92 entomologists from 20 countries, including authors of this Special Issue), who named 103 (!) insect taxa in Arkady’s honor (COLEOPTERA: Salpingidae (1); DIPTERA: Asilidae (5), Cecidomyiidae (1), Chironomidae (1), Dolichopodidae (1), Hybotidae (2), Phoridae (1), Rhagionidae (2), Syrphidae (4); HEMIPTERA: Aphilidae (1); HYMENOPTERA: Andrenidae (2), Apidae (1), Aulacidae (1), Bethylidae (3), Braconidae (14), Bradynobaenidae (1), Cephidae (1), Chrysididae (3), Colletidae (2), Crabronidae (6), Cynipidae (1), Diapriidae (2), Drynidae (2), Encyrtidae (2), Eulophidae (1), Eurytomidae (2), Formicidae (3), Halictidae (1), Ichneumonidae (8), Mutillidae (5), Mymaridae (1), Pamphiliidae (1), Peleserphidae (2), Pompilidae (4), Proctotrupidae (1), Pteromalidae (3), Sphecidae (1), Tenthredinidae (1), Torymidae (1), Vespidae (3); LEPIDOPTERA: Sesiidae (1), Uraniidae (1); ORTHOPTERA: Acrididae (1), Gryllidae (2). We believe we can hardly find similar examples among modern entomologists, and we feel privileged to know him as one of the great modern entomologists, a friend, and a delightful human being.

The Editors and the Editorial Office of the Journal of Hymenoptera Research, numerous colleagues acting as reviewers of the papers, as well as the Administration of the Federal Scientific Center of the East Asia Terrestrial Biodiversity, were of great help in preparing this Special Issue.

Happy 75th birthday, Arkady Stepanovich! We wish you a long and happy life and many new amazing discoveries!

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