Megamymar waorani Huber, here designated.

With the following combination of features: body huge (for a fairyfly), together with exserted section of the ovipositor well over 9 mm long; median ocellus abutting transverse trabecula; petiole distinctly shorter than gaster and, in dorsal view, completely hidden; metatarsus 1 longer than metatibia; gaster extending as horn anterodorsal to mesosoma.

Female. Head. Head slightly wider than mesosoma (21: 20), ~1.4× as wide as long, ~1.1× as wide as high and almost 1.25× as high as long, measured laterally; transverse trabecula entirely dark; supraorbital trabecula with well sclerotized (dark) anterior half short, extending posteriorly only to level of posterior margin of median ocellus, then continuing posteriorly as poorly sclerotized (light) section extending as far as anterior margin of lateral ocellus and further continuing obliquely towards but not reaching occipital foramen as a faint, fine line. Face ~0.5× as wide as high, in lateral view upper face receding to transverse trabecula and lower face flat and, in anterior view, with distinct depression medially dorsal to oral cavity and short, narrow vertical depression ventral to torulus; torulus ~0.5× its own height from transverse trabecula; preorbital sulci slightly converging medially just ventral to toruli then continuing straight ventrally to dorsolateral angle of oral cavity. Compound eye slightly shorter than malar space and apparently with very few short setae among the ommatidia. Vertex in lateral view slightly curved, posteriorly merging smoothly with occiput; median ocellus abutting transverse trabecula; POL 2.0× LOL and ~1.7× OOL. Gena ventrally in lateral view longer than eye length but dorsally much shorter. Back of head without sulci and oral cavity well separated from occipital foramen. Antenna. Scape with radicle barely differentiated; funicle 6-segmented; clava 1-segmented. Mouthparts. Mandibles apparently with 3 teeth, meeting when closed. Mesosoma. Mesosoma almost 2.6× as long as wide, almost 3.2× as long as high, and ~0.8× as wide as high. Pronotum entire, in lateral view almost horizontal, with flat dorsum, in dorsal view triangular and, including neck, almost as long as mesoscutum. Prosternum ~1.3× as long as wide, and entire. Mesoscutum just over 2.0× as long as scutellum + frenum, in lateral view flat; notauli incomplete, ~0.3 as long as mesoscutum as measured from their junction with anterior margin. Scutellum ~4.0× as long as poorly defined frenum; axilla barely advanced, each ~1.0× as wide as long. Metanotum slightly longer than frenum, without obvious dorsellum. Propodeum slightly longer than scutellum + frenum, with a shallow and narrow longitudinal median depression, in lateral view almost horizontal; spiracle in a shallow wide depression. Wings. Fore wing fairly wide, with apex rounded and slightly asymmetrical, and with almost straight margin behind venation; venation ~0.2× as long as wing length; parastigma with proximal but apparently without distal macrochaetae, with hypochaeta next to proximal macrochaeta. Hind wing narrow and almost straight. Legs. Legs long; tarsi 4-segmented, with tarsomere 1 of all legs longer than tibiae. Metasoma. Metasoma ~2.3× as long as mesosoma (Fig. 1). Petiole ~3.4× as long as wide, in dorsal view hidden by anterior extension of gaster. Gaster (measured to anterior apex of anteriorly truncate dorsal horn) ~6.4× as long as wide and ~1.2× as high as wide; gt₅ apparently the longest tergum (Fig. 1). Ovipositor with exserted portion extremely long.

Figure 1. 

Megamymar waorani Huber, holotype female, lateral habitus (ovipositor photoshopped to curve downward to fit page better).

Male. Unknown.

A euphonious combination from Greek: megas, meaning large, and Mymar, the name of the type genus of Mymaridae. The genus name is neuter. Mega refers to the large body of the only known species of the genus, which is over 1 mm longer than the next longest Neotropical species, Erdosiella mira (Annecke & Doutt), whose body length is 3.7 mm.

Megamymar is best placed in Mymarini sensu Annecke and Doutt (1961) because of the combination of 6-segmented funicle, 4-segmented tarsi and distinctly petiolate gaster. Among the genera of Mymarini, the supraorbital trabecula extending posteriorly only to the level of the median ocellus, the fore wing shape, short fringe setae and venation with relatively long parastigma indicate that Megamymar is most closely related to Erdosiella Soyka and Tanyostethium Yoshimoto in the New World, and perhaps Narayanella Subba Rao in the Old World. Megamymar is clearly separated from these three genera by the petiole much shorter than the gaster and the anterodorsal extension of the base of the gaster over the apex of the propodeum; none of their species have this combination of features. Instead, in these genera the supraorbital trabecula extends to about the level of the lateral ocelli, the petiole is usually as long or longer than the gaster, and gt₁ does not project anteriorly over the propodeum though occasionally the base of gt₁ may slant slightly anteriorly before receding uniformly and smoothly posteriorly to the anterior margin of gt₂. It could perhaps be argued that the features of Megamymar are just one of degree rather than substance and therefore Megamymar should be treated as just an extreme representative of Erdosiella, but then many of the numerous genera of Mymarini should be placed in synonymy under one another as well. Whether such an approach would clarify generic relationships within Mymarini is debatable.

Megamymar superficially resembles several of the largest species of Australomymar Girault and the extralimital genera Borneomymar Huber, Neotriadomerus Huber and Paranaphoidea Girault. At least some species in all these genera, none of which are mophologically closely related to Megamymar, have a long ovipositor often greatly exserted posteriorly (Australomymar, Borneomymar, Polynemoidea Girault) or anteriorly (Neotriadomerus). In the Neotropical region, Australomymar appears superficially to be the most similar looking genus but the body of the largest species is at most ~3.0 mm long and the base of the female gaster does not extend anteriorly in a dorsal horn (the gastral sac). In other genera of Mymaridae the ovipositor may extend anteriorly but it is always ventral to the mesosoma.

Holotype female (USNM) on triangular card point labelled 1. “ECUADOR: NAPO Res. Ethnica Waorani, 1km S. Onkone Gare Camp, Trans.Ent. 9 Oct. 1994 220m 00°39'10"S, 76°26'W T.L. Erwin et al.”. 2. “Holotype Megamymar waorani Huber”. 2. “Insecticidal fogging of mostly bare green leaves, some with covering of lichenous or bryophytic plants in terra firme forest. At Trans 5m Sta. 4 Project MAXUS Lot 913” 3. “HOLOTYPE f# Megamymar waorani Huber”.

Megamymar waorani is the only described species in the genus. Its diagnosis is therefore the same as for the generic description, i.e., ovipositor extending anteriorly in a short, truncate gastral sac dorsal to propodeum (Fig. 1).

Female. Colour. Body orange-yellow, with metasoma except for apical tergum and sternum slightly lighter; transverse trabecula and mandibles dark reddish-brown; scape and pedicel dark yellow, fu₁–fu₄ brown except slightly lighter apex of fu₄, fu₅ cream coloured, fu₆ and clava dark brown; legs same colour as body except mesotibia slightly darker yellow, mesotarsomere brown, and metatibia and tarsus almost black. Fore wing clear except yellowish suffusion posterior to and slightly distal to venation; hind wing clear (Fig. 1). Body length. 4.8 mm, excluding exserted part of ovipositor, which extends 4.9 mm posterior to apex of gaster. Head. Head width ~535. Entire face with fine engraved but slightly granulate sculpture and with 2 sublateral setae, about 6 lateral setae and a dense patch of thicker setae sublaterally and laterally at mouth margin. Vertex with 2 setae between lateral ocelli, apparently 2 minute setae lateral to median ocellus, and engraved, slightly granulate reticulations at least anteriorly. Back of head with faint engraved reticulations, 1 short seta posterolateral to lateral ocellus, and a few minute setae lateral to foramen and sublaterally on gena. Antenna. Scape with faint, fine transverse sculpture on inner surface and apparently at least 1 very short seta, on dorsal margin at least. Fu₆ with at least one longitudinal row of 6 or 7 short mps, clava with numerous (11 on at least one surface) short scattered mps; funicle slender, with segments barely widening towards apex, each with apex squarely truncate and with numerous short setae. Length/width measurements (ratio, calculated from eyepiece micrometer measurements before converting to micrometres): scape 220/120 (1.83); pedicel 120/60 (2.00), fu₁ 240/30 (8.00); fu₂ 545/30 (18.33); fu₃ 910/40 (23.00); fu₄ 645/50 (13.00); fu₅ 495/60 (8.33); fu₆ 385/80 (4.87), clava 604/90 (6.78). Mesosoma. Pronotum with 2 setae anteriorly and 1 seta posterolaterally, and with faint raised reticulate sculpture. Mesoscutum with faint raised reticulate sculpture, 1 adnotaular seta at extreme anterior margin and 1 lateral seta posterolaterally on lateral lobe. Scutellum with faint engraved reticulate sculpture and without setae; axilla with 1 small seta; frenum apparently with faint sculpture. Propodeum with 1 propodeal setae well separated from spiracle and with faint reticulate sculpture. Wings. Fore wing with a single line of microtrichia separating bare space posterior to parastigma from bare space anterior to retinaculum; hypochaeta next to proximal macrochaeta, distal macrochaeta not visible (absent?); wing length 3900, width 845, length/width 4.62, longest marginal setae 100, venation length 820, spur 100. Hind wing with median row of microtrichia but only a few distally along anterior and posterior margins; wing length 2400, width 50, longest marginal setae 155, venation length 770. Legs. Metatibia and tarsomeres 1 and 2 with short, dense setae; tarsomere 1 slightly longer than metatibia. Metasoma. Petiole (mostly hidden by metacoxae and gt₁) ~310 long, width 90. Gaster with terga and sterna apparently transparent above yellow ground colour; middle terga apparently the longest segments (not accurately measurable). Ovipositor length (visible part only, the part concealed within gaster not measurable) 5500, the exserted length 4900 with hooked apex.

The species is named after the indigenous Waorani people of Ecuador in whose reserve M. waorani was collected. Their way of life has been seriously affected by resource extraction and settlement by colonists. The species name is treated as a noun in apposition.

Unknown, but because of its size, likely a solitary parasitoid in large insect eggs. We suggest that the host is most likely a species of Orthoptera. First, despite M. waorani being the third longest fairyfly species known worldwide, after specimens of a species of Neotriadomerus from Australia and specimens of one species of Australomymar from New Zealand, it has apparently never been collected at ground level, despite considerable Malaise or pan trapping in equatorial rain forests of the Neotropical region. Second, the host egg must be at least 4.8 mm long and eggs of this size are mostly likely to be found among species of Orthoptera. Third, although hosts of any species of Mymaridae with body length over 3.0 mm are unknown, one relatively small (~1.3 mm) species of Australomymar has been reared from Tettigoniidae (Orthoptera) and one large (2.5 mm) species of Acmopolynema Ogloblin has been reared from Oecanthus spp. (Orthoptera: Gryllidae).

Neopolynemoidea chilensis Huber, here designated.

Female with the following combination of features: toruli almost touching transverse trabecula (Fig. 2a); scape over 10× as long as greatest width and 1.4× as long as head width, funicle 6-segmented and clava 3-segmented (Fig. 3); fore wing venation extending almost 0.7× wing length (Fig. 7).

Figures 2–4. 

Neopolynemoidea chilensis Huber, holotype female 2a head, anterior 2b head posterior (seen through head and flipped horizontally) 3 right antenna 4 left antenna.

Female. Head. Head slightly wider than mesosoma (24: 21), ~1.8× as wide as long, ~1.6× as wide as high and almost 1.2× as high as long, measured laterally; transverse trabecula entire (Fig. 2a); supraorbital trabecula apparently entire. Face 1.2× as wide as high; torulus ~0.25× its own height from transverse trabecula; preorbital sulcus not bulging ventral to eye. Compound eye ~2.0× as long as malar space and with a few short setae among the ommatidia. Vertex in lateral view strongly curved, becoming vertical posteriorly so in same plane as occiput; median ocellus well separated from transverse trabecula; POL 2.0× LOL and ~2.7× OOL (holotype with left lateral ocellus missing entirely, with a seta where it should be). Back of head without sulci. Oral cavity posteriorly almost confluent with occipital foramen (Fig. 2b). Antenna. Scape with radicle barely differentiated; funicle 6-segmented; clava 3-segmented, with the sutures slightly oblique or transverse (Figs 3, 4). Mouthparts. Mandibles apparently with 3 teeth, barely meeting when closed (Fig. 2a). Mesosoma. Almost 2.0× as long as wide, almost 1.4× as long as high, and 0.75× as wide as high. Pronotum in lateral view short, vertical, in dorsal view apparently longitudinally divided medially and not visible except laterally. Prosternum ~1.9× as wide as long and longitudinally divided medially (Fig. 9b). Mesoscutum just over 0.9× as long as scutellum + frenum, in lateral view strongly curved anteriorly; notauli complete (Fig. 9a). Scutellum ~1.2× as long as frenum, with campaniform sensilla midway between anterior and posterior margins and fenestra an isosceles triangle with rounded corners; axilla slightly but distinctly advanced, ~1.8× as wide as long; second phragma extending to apex of propodeum, with rounded apex. Metanotum ~0.75× as long as frenum, with transversely oval/rhomboidal dorsellum. Propodeum medially about 0.6× as long as metanotum. Wings. Fore wing (Fig. 7) fairly wide, with slight lobe posterior to parastigma, and with apex symmetrical and narrowly rounded; venation ~0.6× as long as wing length; parastigma with proximal and slightly longer distal macrochaetae, with hypochaeta close to proximal macrochaeta (Fig. 7). Hind wing (Fig. 8) narrow and almost straight. Legs. Legs fairly long; tarsi 4-segmented, with tarsomere 1 of all legs slightly the longest segment or subequal to tarsomere 4 (Fig. 6). Metasoma. Metasoma ~0.8× as long as mesosoma (Fig. 6). Petiole ~3.2× as wide as long, barely visible in dorsal view. Gaster ~2.0× as long as wide, and ~1.3× as high as wide; gt₆ the longest tergum, gt₇ (syntergum) conical, gt₂–gt₆ subequal in length (Fig. 10a); cerci normal, but almost vertically positioned on sides of syntergum; apical sternum (outer ovipositor plate) much longer and narrower than preceeding sterna (Fig. 10b). Ovipositor arising at level of gt₂ (Fig. 10c) and exserted portion ~0.75× length of internal portion (Fig. 6).

Figures 5–8. 

Neopolynemoidea chilensis Huber, holotype female 5 holotype slide 6 habitus without head, dorsal 7 fore wing 8 hind wing.

Figure 9. 

Neopolynemoidea chilensis Huber, holotype female a mesosoma, dorsal b mesosoma, ventral (seen through mesosoma).

Figure 10. 

Neopolynemoidea chilensis Huber, holotype female a metasoma, dorsal b metasoma, ventral (seen through metasoma and flipped vertically), showing sterna c metasoma, ventral (seen through metasoma and flipped vertically), showing genitalia.

Male. Unknown.

From Greek: Neo- meaning new, referring to its occurrence in the New World and Polynemoidea, a monotypic genus known so far only from Tasmania. The name Neopolynemoidea is given to draw attention to the general similarity the two genera, one from the Old World and one from the New World.

Neopolynemoidea differs from Polynemoidea by: toruli almost in contact with transverse trabecula (separated from transverse trabecula by almost its own height in Polynemoidea), scape over 10× as long as wide (no more than 3× as long as wide in Polynemoidea); fore wing venation at least 0.6× as long as fore wing length (no more than 0.4 as long as fore wing length in Polynemoidea).

Worldwide, at least 14 genera or subgenera of Mymaridae have females with a 3-segmented clava: Allanagrus Noyes & Valentine, Allarescon Noyes & Valentine, Eustochomorpha Girault, Krokella Huber, Nesomymar Valentine, Nesopatasson Valentine, Neostethynium Ogloblin, Notomymar Doutt & Yoshimoto, Paracmotemnus Noyes & Valentine, Paranaphoidea (Idiocentrus) Gahan, Platystethynium (Platystethynium) Ogloblin, Polynemoidea, Pseudanaphes Noyes & Valentine, and Stethynium Enock. Except for Eustochomorpha, with an 8-segmented funicle in females, all have a 6-segmented funicle. The majority of these genera occur in the southern hemisphere, particularly in the southernmost areas, though several extend well into the northern hemisphere. Two other genera appear to have the clava with 3-segments in at least one species: one Eustochus (Eustochus) from China was described as having 3-segmented clava (the other described species have a 2-segmented clava) but this may be an artefact of partial antennal collapse, giving the appearance of a third segment; and Kompsomymar Huber, with a single described species from Australia, appears to have only partial divisions separating the claval segments. The hosts and biology of all but one (Stethynium) of the above genera are unknown. Among these, Krokella, Paracmotemnus and Polynemoidea have a fore wing venation longer than half the wing length, as in Neopolynemoidea, but none have the extremely long scape in females. Females of the only described species of Polynemoidea, however, have a long, exerted ovipositor. Therefore, based wing venation and ovipositor features a close relationship of Neopolynemoidea to Polynemoidea is proposed as being the most probable.

Holotype female (CNC) in Canada balsam on slide (Fig. 5) labelled: 1. “Chile: Cautin, 1150m, Conguillio Nat. Park, Auraucaria Nothofagus Feb. 6 .1988 18PT L. Masner Chile Exp.” 2. “Neopolynemoidea chilensis f# Huber HOLOTYPE”. Paratypes: one female (CNC) on card, with same locality data as holotype.

Neopolynemoidea chilensis is the only described species in the genus so its diagnosis the same as for the generic description.

Female. Colour. Uniformly brown; scape, pedicel and legs except coxae and metafemur brownish yellow. Fore wing with brown suffusion except for a small clear area distal to stigma, a larger triangular area along posterior margin, a clear area anterior to retinaculum and a small one posterior to parastigma (Fig. 7). Body length. 1000 (paratype, card mount). Head. Head width ~226. Face with 6 setae ventral to level of ventral margin of torulus and faint engraved reticulations except for a band of more distinct striations extending between eyes ventral to toruli and a narrow dorsal band extending along preorbital groove (Fig. 2a). Vertex with 2 setae between lateral ocelli and slightly more distinct engraved reticulations. Back of head with faint transverse engraved reticulations dorsally, becoming vertical lateral to foramen, 2 short seta dorsal to foramen, and a few shorter setae laterally along eye (Fig. 2b). Antenna. Scape with faint, fine longitudinal sculpture on inner surface and several setae on ventral and dorsal margins (Fig. 3). Fu₄ with 1 mps, fu₅ and fu₆ each with 2 mps, clava with 2, 2, and 4 mps on segments 1–3, respectively; funicle with segments enlarging towards apex, each with apex squarely truncate and each with numerous fairly long setae (Figs 3, 4). Length/width (ratio) (holotype): scape ~320/30 (10.55); pedicel 76/36 (2.11), fu₁ 59/16 (3.08); fu₂ 114/19 (5.96); fu₃ 105/18 (5.70); fu₄ 93/21 (4.46); fu₅ 83/26 (3.19); fu₆ 72/28 (2.57), clava 185/47 (3.92). Mesosoma. Pronotum with 2 minute submedial and 2 longer sublateral setae. Mesoscutum with faint engraved reticulate sculpture, 1 adnotaular seta and 1 lateral seta on lateral lobe. Scutellum with faint engraved reticulate sculpture, campaniform sensilla midway between anterior and posterior margins and closer to each other than to scutellar seta, scutellar seta at margin of axillular carina, a minute seta/pit? anterior to apex of scutellar carina, and fenestra an isosceles triangle with rounded angles; axilla with 1 lateral seta and faint engraved reticulate sculpture; frenum with faint reticulate sculpture becoming longitudinal at lateral margin. Propodeum with 2 minute propodeal setae well separated from spiracle and with reticulate sculpture. Wings. Fore wing with a single line of microtrichia separating bare space posterior to parastigma from bare space anterior to retinaculum; hypochaeta close to proximal macrochaeta; wing length (holotype) 1210, width 222, length/width 5.46, longest marginal setae 312, venation length 739. Hind wing with a median row of about 15 microtrichia mostly in apical half beyond venation; wing length 1224, width 36, longest marginal setae 216, venation length 376. Legs. Metatibia with setae at least 2× as long as tibial width (Fig. 6). Metasoma. Petiole ~0.4× as long as wide, distinctly narrower than gt₁ (Figs 6, 9a). Gaster with segments subequal in length except gt₁ slightly shorter; gt₇ triangular, apically truncate. Gt₁ with 2 sublateral setae, gt₆ and gt₇ each with 1 submedian setae, remaining terga with 3 (sometimes 2) submedian/sublateral setae. Ovipositor length 833, ~1.8× as long as metatibia, and exserted length 400.

Unknown.

Porcepicus herison Huber, here designated.

Female with the following combination of features: back of head dorsal to foramen with median vertical occipital groove and transverse occipital groove/trabecula extending from eye to eye (Fig. 12a); mandible a small stub without teeth (Fig. 12a); gastral petiole apparently absent and gaster wider than long; body, legs and antenna with prominent stout spines (Figs 11–20).

Figure 11. 

Porcepicus herison Huber, holotype female on card, before slide mounting.

Female. Head. Head slightly narrower than mesosoma (14:17), ~2.0× as wide as long, ~1.7× as wide as high and ~1.15× as high as long, measured laterally; transverse and supraorbital trabeculae with short dark sections alternating with light sections. (Fig. 12a). Face slightly wider than high; torulus ~1.4× its own height from transverse trabecula; preorbital groove bulging laterally ventral to eye. Compound eye ~1.7× as long as malar space and with a few short blunt setae among ommatidia. Vertex posteriorly with sharp, slightly concave margin at junction with occiput (Fig. 12b); median ocellus well separated from transverse trabecula; ocelli in a low triangle, with lateral ocelli at posterolateral angle of vertex, and POL ~2× LOL and ~23× OOL. Back of head with vertical occipital groove, complete transverse occipital groove obtusely angled medially dorsal to occipital foramen, and oral cavity posteriorly almost confluent with occipital foramen (Fig. 12b). Antenna. Scape with radicle barely differentiated; funicle 6-segmented; clava 1-segmented (Figs 14a, b). Mouthparts. Mandibles without teeth, with rounded apex shorter than maxilla, presumably not meeting when closed (Fig. 12a). Mesosoma. Mesosoma ~0.8× as long as wide, ~1.2× as long as high, and ~1.4× as wide as high. Pronotum in lateral view almost vertical, apparently longitudinally divided medially, in dorsal view barely visible except laterally. Prosternum almost 2.0× as wide as long and longitudinally divided medially (Fig. 19). Mesoscutum as long as scutellum + frenum, in lateral view flat; notauli incomplete, barely indicated anteriorly (Fig. 18a). Scutellum ~0.25× as long as frenum, without campaniform sensilla or fenestra; axilla barely advanced, about as long as wide; second phragma extending to apex of propodeum, widely truncate apically. Metanotum linear and barely visible, without defined dorsellum. Propodeum medially ~5.0× as long as metanotum. Wings. Fore wing narrow and curved (Fig. 15) with distinct lobe posterior to parastigma; venation ~0.3× as long as wing length; parastigma with distal macrochaeta but without proximal macrochaeta or hypochaeta. Hind wing (Fig. 16) with distinct bend in basal third beyond venation, with anterior margin concave and posterior margin convex. Legs. legs short; tarsi 5-segmented, with protarsomere 1 the longest segment, and meso-and metararsomere 1 the shortest segments. Metasoma. Metasoma ~0.8× as long as mesosoma (Figs 18a, 20). Petiole vertical, not visible in dorsal view, ~10× as wide as long (high). Gaster ~1.2× as wide as long, and ~1.1× as wide as high; gt₂ and gt₃ the longest terga, gt₁ planoconvex, the almost straight anterior margin and curved posterior margin meeting laterally to form an acute angle (Fig. 18a); cercus apparently with only 2 cercal setae. Ovipositor arising in apical half of gaster, slightly less than 0.5× gaster length and slightly less than 0.5× metatibia length.

Figures 12–17. 

Porcepicus herison Huber, holotype female 12a head, anterior 12b head, posterior (seen through head and flipped horizontally) 13a right antenna (clava and fu₅ missing) 13b right antenna (opposite surface seen through antenna and flipped horizontally) 14a left antenna 14b left antanna (opposite surface seen through antenna) 15 fore wing 16 hind wing 17 holotype slide.

Figure 18. 

Porcepicus herison Huber, holotype female 18a habitus without head, dorsal 18b genitalia (seen through gaster).

Male. Unknown.

An arbitrary combination of letters based on the French word for porcupine, porc-épique, referring to the long and strong setae distributed on the antenna, body, and legs.

Porcepicus belongs to the Camptoptera group of genera. It appears to be most similar to Camptoptera by the back of the head having a vertical occipital groove, transverse occipital groove, and narrow and curved fore wing and lack of a hypochaeta. The 6-segmented funicle in females, slightly dorsoventrally flattened mesosoma, gaster wider than long, and apparent absence of a petiole distinguishes it from Camptoptera Foerster as well as the other genera in the genus group.

Holotype female (CNC) in Canada balsam on slide (Fig. 17) labelled: 1. “Peru: Loreto 220m Teniente Lopez 2°36'S, 76°07'W, 22.VII.1993, R. Leschen FIT” 2. “Porcepicus herison f# HOLOTYPE”.

Paratypes : Four females. PERU. Same locality data as holotype (2 females on cards, CNC); Junín River, NW of San Ramón, Río Oxabamba, San Fernan Farm, 925 m, 11°5'36"S, 75°23'43"W, 30.vi.2010, M. Hoddle, MT [Malaise trap] (1 female on slide, UCRC, UCRC ENT 285052); San Martín, 19 km NE Tarupoto, 950 m, 6–8.vii.2004, B. V. Brown, MT (1 female on point, UCRC, UCRC ENT 457917).

Porcepicus herison is the only described species in the genus so its diagnosis is the same as for the generic description.

Female. Colour. Body brown (mesoscutum light brown in one paratype) except anterior surface of gt₁ white; legs light brown, pedicel and tarsi almost white; thick setae and their sockets on body, antenna and legs dark brown (Figs 11, 20). Body length. 310–330 μm (paratypes). Head. Head width 145. Face almost smooth, with 4 fairly long strong setae at or ventral to level of ventral margin of torulus. Vertex with shallow transversely reticulate sculpture and 2 long strong erect setae on each side of lateral ocellus. Gena with 1 strong seta lateral to oral cavity (Fig. 12a, b). Back of head (Fig. 12b) with faint transverse to oblique reticulate sculpture and 2 short setae dorsal to occipital groove. Antenna. Scape with faint trace of longitudinal sculpture on inner surface and about 4 long setae on ventral and dorsal margins; pedicel 0.8× as long as scape, with ventral margin strongly indented basally; funicle with segments enlarging towards apex, each with apex obliquely truncate (Figs 13, 14) and with 1 or 2 long, fairly strong setae on ventral margin; fu₂–fu₅ each with 1 (possible 2) mps; clava with 1 (possibly 2) longitudinal mps (Fig. 14). Length/width (ratio) (holotype): scape ~55/23 (~2.30), pedicel ~35/25 (1.40), fu₁ 15/9 (1.67), fu₂ 11/11 (1.00), fu₃ 11/12 (0.92), fu₄ 15/13 (1.15), fu₅ 16/13 (1.23), fu₆ 17/16 (1.06), clava 50/24 (2.08). Mesosoma. Pronotum with 1 long strong lateral seta; mesoscutum with transverse reticulate sculpture and 1 long strong seta at lateral angle; scutellum without campaniform sensilla and apparently smooth; axilla smooth, with 1 long strong seta; frenum with faint transverse reticulations medially, becoming longitudinal at lateral margin. Propodeum with long strong seta, its base almost in contact with small circular spiracle. Wings. Fore wing with a row of about 9 microtrichia between usual anterior and posterior rows; wing length (holotype) 368, width 32, length/width 11.5, longest marginal setae 157, venation length 114. Hind wing medially with anterior row of about 15 microtrichia and posterior row of 3 microtrichia; wing length 379, width 12, longest marginal setae 114, venation length 103. Legs. Meso- and metatibia each with 3 long strong setae on dorsal margin; coxae and femora with shorter, moderately strong setae (Fig. 18a). Metasoma. Gaster apparently smooth (Fig. 18a); gt₂–gt₅ each with 1 or 2 long strong sublateral and lateral setae; apical tergum with 1 median seta (Fig. 18b). Ovipositor length 45, its apex distinctly anterior to apex of gaster (Figs 18b, 19).

Figures 19, 20. 

Porcepicus herison Huber 19 holotype, habitus without head, ventral (seen through body and flipped vertically) 20 paratype, habitus, dorsal, fore wings missing.

The name is an arbitrary combination of letters similar to the French word for hedgehog, hérisson. The name is treated as a noun in apposition.

Unknown. As with most species in the Camptoptera group of genera the hosts are unknown. We suggest the hosts are Coleoptera based on at least one record from that order for Camptoptera and one for Litus.