Research Article |
Corresponding author: Marko Prous ( mprous@ut.ee ) Academic editor: Maksim Proshchalykin
© 2021 Marko Prous, Andrew Liston, Marko Mutanen.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Prous M, Liston A, Mutanen M (2021) Revision of the West Palaearctic Euura bergmanni and oligospila groups (Hymenoptera, Tenthredinidae). In: Proshchalykin MYu, Gokhman VE (Eds) Hymenoptera studies through space and time: A collection of papers dedicated to the 75th anniversary of Arkady S. Lelej. Journal of Hymenoptera Research 84: 187-269. https://doi.org/10.3897/jhr.84.68637
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Eight Western Palaearctic Euura species are here assigned to the bergmanni group (bergmanni, brevivalvis, dispar, glutinosae, leptocephalus, respondens, sylvestris, and viridis) and two species to the oligospila group (frenalis and oligospila). Euura pallens (Konow, 1903) (bergmanni group) is removed from the list of West Palaearctic taxa. Euura pyramidalis (Hellén, 1948) is treated as incertae sedis within the bergmanni group. Definitions of the bergmanni and oligospila groups are primarily based on genetic sequence data (mitochondrial COI and nuclear NaK and POL2). We report likely occurrence of heteroplasmy and amplification of NUMTs among some of the treated species, complicating the use of DNA barcoding in species discrimination. Based on morphological and genetic evidence, we establish that the correct name for the invasive willow sawfly in the southern hemisphere (South America, southern Africa, Australia, New Zealand), known there only in the female sex, is Euura respondens (Förster, 1854). The species is probably native to the Palaearctic (or even Holarctic) where males are common: possibly as common as females (examined from Europe and Central Asia). The name Euura oligospila (Förster, 1854) has been incorrectly used for the species in the southern hemisphere. The examination of type material and reliable association of males and females based on genetics revealed that females of E. oligospila are morphologically extremely similar to E. respondens (and to some other E. bergmanni group species), but male penis valves and genetics enable reliable separation of these species. Morphological separation of females of E. oligospila and E. respondens is possible, but challenging. Identification keys for males and females of the bergmanni and oligospila groups are provided. The following 15 new synonymies are proposed: Nematus validicornis Förster, 1854, syn. nov. with Euura bergmanni (Dahlbom, 1835); Pteronidea woollatti Lindqvist, 1971, syn. nov. and Nematus turgaiensis Safjanov, 1977, syn. nov. with Euura brevivalvis (Thomson, 1871); Pteronidea pseudodispar Lindqvist, 1969, syn. nov. with Euura dispar (Zaddach, 1876); Nematus (Pteronidea) fastosus var. ponojense Hellén, 1948, syn. nov. and N. (P.) fastosus var. punctiscuta Hellén, 1948, syn. nov. with Euura frenalis (Thomson, 1888); Nematus declaratus Muche, 1974, syn. nov. and N. desantisi D.R. Smith, 1983, syn. nov. with Euura respondens (Förster, 1854); Pteronidea straminea Lindqvist, 1958, syn. nov., P. angustiserra Lindqvist, 1969, syn. nov., and P. disparoides Lindqvist, 1969, syn. nov. with Euura sylvestris (Cameron, 1884); Pteronidea breviseta Lindqvist, 1946, syn. nov., P. breviseta Lindqvist, 1949, syn. nov., P. abscondita Lindqvist, 1949, syn. nov., and P. lauroi Lindqvist, 1960, syn. nov. with Euura viridis (Stephens, 1835). Lectotypes are designated for 18 nominal taxa: Amauronematus longicornis Konow, 1897; A. spurcus Konow, 1904; Nematus bergmanni Dahlbom, 1835; N. brevivalvis Thomson, 1871; N. curtispina Thomson, 1871; N. (Pteronidea) fastosus var. ponojense Hellén, 1948; N. (P.) fastosus var. punctiscuta Hellén, 1948; N. glutinosae Cameron, 1882; N. microcercus Thomson, 1871; N. polyspilus Förster, 1854; N. prasinus Hartig, 1837; N. respondens Förster, 1854; N. salicivorus Cameron, 1882; N. validicornis Förster, 1854; N. virescens Hartig, 1837; Pteronidea curtispina var. luctuosa Enslin, 1916; Pteronus fastosus Konow, 1904; and P. pallens Konow, 1903.
DNA barcode sharing, heteroplasmy, invasive species, lectotype, mitonuclear discordance, Nanopore sequencing, NUMTs, sawflies, synonyms
The genus Euura Newman, 1837, native to the Holarctic and Oriental regions, is the largest in the Nematinae (Tenthredinidae), containing about 650 species, or half of the subfamily (
Specimens examined or mentioned are deposited in the following collections:
CMH Collection of Mikk Heidemaa, Tartu, Estonia;
CVV Collection of Veli Vikberg, Turenki, Finland;
NIBIO Norwegian Institute of Bioeconomy Research, Svanhovd, Norway;
SMTP Swedish Malaise Trap Project, Station Linné, Öland, Sweden;
ZMHB Museum für Naturkunde der Humboldt-Universität, Berlin, Germany;
Other abbreviations:
BIN barcode index number;
HT holotype;
LT lectotype;
ST syntype or syntypes.
Specimens studied are listed in the Suppl. material
To photograph penis valves and lancets (valvula 1 or ventral part of the saw), genital capsules and ovipositors were separated from the specimen and macerated in KOH (10–15%) for 6–10 hours at room temperature, or treated with proteinase during DNA extraction. Temporary slide preparations of dissected lancets and penis valves in glycerine were made, and after photography, the parts were glued on a piece of cardboard, which was pinned with the corresponding specimen. In addition, relevant permanent slide preparations borrowed from institutional or personal (Veli Vikberg, Turenki) collections were photographed. Photos were taken with a digital camera attached to a microscope. Composite images with an extended depth of field were created from stacks of images using the software CombineZP (Alan Hadley; http://www.hadleyweb.pwp.blueyonder.co.uk/, although apparently no longer available) or Helicon Focus 7.6.4. Most of the lancets were photographed in two overlapping parts and a single image was created using the program Image Composite Editor (Microsoft) or with the plugin MosaicJ (
For species delimitation and to associate males and females, mitochondrial and nuclear DNA was sequenced from representatives of Euura belonging to the bergmanni and oligospila groups as well as other selected species. DNA was sequenced using Sanger (
The bergmanni and oligospila groups of Euura are here primarily defined based on phylogenetic analyses of DNA sequence data (Fig.
Based on three genes (COI, NaK, POL2), the branching order at the base of Euura phylogeny is poorly resolved, but several strongly supported groups can be identified, such as the bergmanni and oligospila groups (Fig.
Maximum likelihood tree of Euura based on three genes (mitochondrial COI and nuclear NaK and POL2). Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Support values for weakly supported branches (<90) are not shown. Letters “f” and “m” stand for “female” and “male” if known. Numbers at the end of the tip labels refer to the length of the sequence and the number of ambiguous positions (e.g., heterozygosities). The tree was arbitrarily rooted between Euura and other Nematini. The scale bar shows the number of estimated substitutions per nucleotide position.
Because of clear conflict between mitochondrial and nuclear data within the bergmanni and oligospila groups themselves, we analysed these gene sets separately for each group. Within the bergmanni group, the viridis subgroup is the only relationship above species level that is strongly supported both by COI and nuclear genes. The other relationships are poorly supported or even strongly contradictory (Fig.
Maximum likelihood trees of Euura bergmanni group (COI left, nuclear NaK and POL2 upper right) and viridis subgroup (NaK lower left, POL2 lower right). Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Support values for weakly supported branches (<90) are not shown. Letters “f” and “m” stand for “female” and “male” if known. Numbers at the end of the tip labels refer to the length of the sequence and the number of ambiguous positions (e.g., heterozygosities). Note the COI heteroplasmic variants for brevivalvis
Maximum likelihood trees of Euura oligospila group based on COI (left) and nuclear genes (NaK and POL2, right). Numbers at branches show SH-aLRT support (%) / ultrafast bootstrap support (%) values. Support values for weakly supported branches (<90) are not shown. Letters “f” and “m” stand for “female” and “male” if known. Numbers at the end of the tip labels refer to the length of the sequence and the number of ambiguous positions (e.g., heterozygosities). Note the COI NUMT for oligospila
Unfortunately, mitochondrial COI data is unreliable for species delimitation as it often conflicts with morphology and nuclear data, which is a common pattern in sawflies (
Due to the high degree of similarity of females in the bergmanni and oligospila groups, many of the names have confusing histories, and misinterpretations are also common in the recent literature. Here, only the more prominent or recent examples will be discussed. Particularly, there has been confusion about the identity of the invasive willow sawfly in the southern hemisphere (South America, southern Africa, Australia, New Zealand), which we here identify as Euura respondens (Förster, 1854). The name Nematus oligospilus [= E. oligospila (Förster, 1854)] has been incorrectly used for this sawfly in the southern hemisphere due to the high degree of similarity between females of E. oligospila and E. respondens. These two species prove to be genetically distant within Euura (Fig.
Remarkably, multiple mitochondrial COI variants are frequently observed within individuals of Euura (unpublished), including in the bergmanni and to a lesser extent in the oligospila groups. The viridis subgroup of the bergmanni group is the most problematic in this regard, where many individuals have secondary peaks in Sanger chromatograms, indicating the presence of multiple variants. Usually, the secondary peaks are rather weak, enabling the determination of the dominant variant, but sometimes different variants are represented at similar frequencies, making it necessary to code variable sequence positions with IUPAC ambiguity symbols. Some of the specimens referred to here were re-amplified and sequenced with Nanopore technology, enabling us to determine the exact variants and place them in the tree (Figs
Females
Abdomen posteriorly with modified segments 8–9 forming the ovipositor and its sheath (Fig.
Characters of Euura hypoxantha, bergmanni and oligospila groups, females A oligospila DEI-GISHym84738 B leptocephalus
1 | a Head and thorax mostly reddish (Fig. |
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b Abdomen dorsally mostly black (Fig. |
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c Frontal area between antennae protruding less abruptly below (Fig. |
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d Arctic species | 2 | |
– | aa Head and thorax mostly yellowish to mostly black (pale parts can be green in life) (e.g. Figs |
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bb Abdomen completely pale to dorsally mostly black (pale parts usually green in life) (e.g. Figs |
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cc Frontal area between antennae protruding more abruptly below (Fig. |
3 | |
2(1) | a Dorsal margin of lancet angulate at about second or third serrula from apex, apical sutures distinctly inclined apically (Fig. |
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b Valvula 3 in dorsal view narrow (Fig. |
E. leptocephalus (bergmanni group) | |
– | aa Dorsal margin of lancet not angulate at about second or third serrula from apex, apical sutures not distinctly inclined apically (Fig. |
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bb Valvula 3 in dorsal view usually broad at base (Fig. |
similar species mostly in flavescens group (E. reticulata, E. lienterica in part, E. pallidior…) | |
3(1) | a Tip of lancet rather broad and dorsally abruptly narrowing (Fig. |
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b Serrulae apically protruding and / or with large microdenticles (Fig. |
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c Malar space 1.5–2.2 times as long as diameter of front ocellus (Fig. |
4 | |
– | aa Tip of lancet usually narrower, acute, and dorsally narrowing less abruptly (Figs |
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bb Serrulae apically not protruding and with small microdenticles (Figs |
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cc Malar space 1.1–2.0 times as long as diameter of front ocellus (Fig. |
7 | |
4(3) | a Valvula 3 in lateral view shorter (Fig. |
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b Lancet shorter and broader (Fig. |
E. brevivalvis | |
– | aa Valvula 3 in lateral view longer (Fig. |
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bb Lancet longer and narrower (Fig. |
5 [the following three species not definitely morphologically separable] | |
5(4) | a Slightly larger denticles of serrulae (uncertain) (Fig. |
E. dispar |
– | aa Slightly smaller denticles of serrulae (uncertain) (Fig. |
6 |
6(5) | a On Betula (morphological differences unknown) | E. viridis |
– | aa On Alnus (morphological differences unknown) | E. glutinosae |
7(3) | a Ventral part of 2nd to 4th suture of lancet oblique and more or less straight or weakly curved apically (Fig. |
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b Gap between serrulae relatively large (Fig. |
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c Head dorsally more rounded (Fig. |
E. respondens | |
– | aa Ventral part of 2nd to 4th suture of lancet in most cases weakly or distinctly curved basally (Figs |
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bb Gap between serrulae large or small (Figs |
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cc Head dorsally rounded or rectangular (Fig. |
8 | |
8(7) | a Lancet usually relatively broad (Fig. |
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b Sutures of basal annulets usually weakly bent (Fig. |
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c Gap between serrulae small, relative to length of serrulae (Fig. |
E. bergmanni | |
– | aa Lancet narrower (Fig. |
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bb Sutures of basal annulets distinctly bent (Fig. |
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cc Gap between serrulae small or large, relative to length of serrulae (Fig. |
9 | |
9(8) | a Malar space more than 1.4 times as long as diameter of front ocellus (Fig. |
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b Clypeus usually less deeply emarginate (Fig. |
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c Head in dorsal view more rectangular (Fig. |
E. sylvestris | |
– | aa Malar space less than 1.5 times as long as diameter of front ocellus (Fig. |
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bb Clypeus usually more deeply emarginate (Fig. |
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cc Head in dorsal view more rounded (Fig. |
10 | |
10(9) | a Gap between serrulae smaller compared to length of serrulae (Fig. |
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b Terga 1–8 dorsally usually completely black (Fig. |
E. frenalis | |
– | aa Gap between serrulae larger compared to length of serrulae (Fig. |
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bb Terga 1–8 dorsally usually nearly completely pale or with narrow black median line (Fig. |
E. oligospila |
Males
Abdomen posteriorly with a genital capsule, ventrally covered by undivided sternum 9 (Fig.
Characters of Euura bergmanni and oligospila groups, males. A respondens DEI-GISHym84697 B oligospila DEI-GISHym84695 C frenalis
1 | a Pronotum black (Fig. |
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b Paravalva basally with two-digit extension (Fig. |
E. leptocephalus | |
– | aa Pronotum extensively pale, often also rest of body (e. g. Figs |
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bb Paravalva basally simple (Figs |
2 | |
2(1) | a Pterostigma more or less uniformly pale (Figs |
3 |
– | aa Pterostigma dark brown to black, sometimes centrally somewhat paler (e.g. Figs |
11 |
3(2) | a Valviceps basally about as broad as apically and with weak or distinct constriction in middle (Fig. |
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b Invagination between valvispina and paravalva indistinct (Fig. |
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c Apical projection of tergum 8 about as broad as long or slightly longer than broad and about 0.5 times as long as tergum 8 (Fig. |
4 | |
– | aa Valviceps often basally narrower than apically and without distinct constriction in middle (Fig. |
|
bb Invagination between valvispina and paravalva distinct or indistinct (Fig. |
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cc Apical projection of tergum 8 about as broad as long or distinctly longer than broad and about 0.3–0.5 times as long as tergum 8 (Fig. |
5 | |
4(3) | a Valvispina possibly broader at base (Fig. |
|
b Mesepisternum (usually?) extensively pale (Fig. |
E. oligospila | |
– | aa Valvispina possibly narrower at base (Fig. |
|
bb Mesepisternum usually black (Fig. |
E. frenalis | |
5(3) | a Valvispina short (Fig. |
|
b Head in dorsal view rectangular (Fig. |
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c Antennae black or slightly pale ventrally (Fig. |
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d 6.0–8.0 mm | E. brevivalvis | |
– | aa Penis valve with different combination of characters (Fig. |
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bb Head in dorsal view rectangular to square or round (Fig. |
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cc Antennae usually distinctly pale at least ventrally (e.g. Figs |
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dd 4.5–8.0 mm | 6 | |
6(5) | a Pseudoceps apically with distinct filament-like constriction (Fig. |
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b Invagination between valvispina and paravalva distinct (Fig. |
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c Valvispina distinctly dorsally directed (Fig. |
E. respondens | |
– | aa Pseudoceps apically without filament-like constriction (Fig. |
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bb Invagination between valvispina and paravalva distinct or indistinct (Fig. |
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cc Valvispina in most species not dorsally directed (Fig. |
7 | |
7(6) | a Invagination between valvispina and paravalva indistinct (Fig. |
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b Valvispina relatively short (Fig. |
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c Paravalva apically distinctly broader than basally (Fig. |
E. sylvestris | |
– | aa Invagination between valvispina and paravalva distinct (Fig. |
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bb Valvispina relatively long (Fig. |
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cc Paravalva apically distinctly broader than basally or about as broad as basally (Fig. |
8 | |
8(7) | a Dorsal margins of valvura and pseudoceps distinctly angled (Fig. |
|
b Valvispina somewhat directed dorsally (Fig. |
E. bergmanni | |
– | aa Dorsal margins of valvura and pseudoceps not angled (Fig. |
|
bb Valvispina not directed dorsally (Fig. |
9 | |
9(8) | a Invagination between valvispina and paravalva deep (Fig. |
E. dispar |
– | aa Invagination between valvispina and paravalva shallower (Fig. |
10 |
10(9) | a Gap between valvispina and paravalva possibly smaller (Fig. |
|
b On Betula | E. viridis | |
– | aa Gap between valvispina and paravalva possibly larger (Fig. |
|
bb On Alnus | E. glutinosae | |
11(2) | a Valvispina long and somewhat directed dorsally (Fig. |
|
b Malar space 1.0–1.4 times as long as diameter of front ocellus (Fig. |
E. bergmanni | |
– | aa Valvispina long and not directed dorsally, or short and somewhat directed dorsally (Fig. |
|
bb Malar space 1.3–1.9 times as long as diameter of front ocellus (Fig. |
12 | |
12(11) | a Valvispina long (Fig. |
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b Invagination between valvispina and paravalva deep (Fig. |
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c Antennae usually distinctly pale at least ventrally (Fig. |
E. dispar | |
– | aa Valvispina short (Fig. |
|
bb Invagination between valvispina and paravalva shallow (Fig. |
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cc Antennae black or slightly pale ventrally (Fig. |
E. brevivalvis |
Penis valves of Euura bergmanni group. A bergmanni
The group is mostly defined based on phylogenetic analyses of sequence data (Fig.
Nematus bergmanni Dahlbom, 1835: 24–25. Type locality: Sweden, Lund area. LT designated below.
Nematus virescens Hartig, 1837: 217. Type locality: not stated, but presumably Germany according to the title of Hartig’s publication. LT designated below.
Nematus pallicarpus
Hartig, 1837: 215–216. Type locality: “in hiesiger Gegend” [Germany, Berlin area]. LT designated by
Nematus validicornis Förster, 1854a: 341–342. Type locality: Germany, Aachen area. LT designated below. Syn. nov.
Nematus curtispina Thomson, 1871: 152–153. Type locality: “Probably as widespread as the previous species” [translated from Swedish], which presumably refers to N. brevivalvis: “Probably occurs throughout Scandinavia” [translated from Swedish]. LT designated below.
Nematus varipictus
Holmgren, 1883: 147, Plate 2, Fig.
Nematus anthophilus
Zaddach, 1884 [in
Amauronematus longicornis Konow, 1897: 179. Type locality: Russia, Irkutsk. LT designated below.
Lygaeonematus pallens
Enslin, 1916: 500–501. Type locality: Germany, Dessau. LT designated by
Pteronidea curtispina var. luctuosa Enslin, 1916: 455. Type locality: Germany, Bavaria, Fürth. LT designated below.
Pteronidea vernalis
Lindqvist, 1937: 130–132. Type locality: southern Finland (HT). Not found in
Females are most similar to oligospila group, E. respondens, and E. sylvestris. Head more rectangular in dorsal view compared to oligospila group and E. respondens. Lancet usually broader and basal annular sutures usually less bent compared to E. sylvestris and E. respondens. Gap (cypsella) between serrulae small compared to oligospila group. Males distinguishable from other species by distinct penis valves (see key). Length of postocellar area in E. bergmanni is not a reliable character to distinguish it at least from E. sylvestris. Females of overwintering generation are dorsally largely black, the later generations largely or nearly completely pale (pterostigma is always pale). Males of overwintering generation are largely black (except legs and abdomen ventrally to various degrees), including pterostigma; the later generations are ventrally largely or nearly completely pale (including pterostigma) but dorsally mostly black. At least in females there tend to be distinct differences between the generations also in the length of malar space and perhaps postocellar area. In overwintering generations, the malar space tends to be distinctly shorter (Fig.
COI. Based on 13 specimens, maximum within species distance is 3.65% and the nearest neighbour, diverging by a minimum of 7.1%, is the viridis subgroup. Only one BIN: BOLD:AAG3539.
Nuclear. Based on 5 specimens, maximum within species distance is 0.19% (0.23% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 3.5%, is viridis.
Hosts: a wide variety of Salix species, including alba, fragilis (
Palaearctic (
Nematus bergmanni Dahlbom, 1835. Lectotype, here designated, ♀, MZLU2017334,
Nematus virescens Hartig, 1837. Lectotype, here designated, ♀, GBIF-GISHym3456,
Nematus pallicarpus Hartig, 1837. LT, “Cotype”, “Nematus pallicarpus Htg. Th. Hartig det.”, “Pteronidea curtispina Th. E. Clément det.”, DEI-GISHym84734,
Nematus validicornis Förster, 1854. Lectotype, here designated, ♂, GBIF-GISHym3451,
Nematus curtispina Thomson, 1871. Lectotype, here designated, ♀, MZLU2017334,
Amauronematus longicornis Konow, 1897. Lectotype, here designated, ♂, GBIF-GISHym3849,
Lygaeonematus pallens Enslin, 1916. LT, ♀, DEI-GISHym84733,
Pteronidea curtispina var. luctuosa Enslin, 1916. Lectotype, here designated, ♂, GBIF-GISHym3339,
Nematus brevivalvis Thomson, 1871: 151–152. Type locality: Sweden, Dalarna alpina. LT designated below.
Pteronus kriegeri
Konow, 1903a: 310 (key). Type locality: Germany, Saxony, Dorneichenbach..LT designated by
Amauronematus spurcus Konow, 1904: 261. Type locality: northern Russia. LT designated below.
Pteronidea absimilis Lindqvist, 1949: 79–80. Type locality: Finland, Pihtipudas (HT).
Pteronidea woollatti Lindqvist, 1971: 11–12. Type locality: Finland, Kilpisjärvi (HT). Syn. nov.
Nematus turgaiensis Safjanov, 1977: 98–103. Type locality: Russia, southern Kulunda steppe (ST). Syn. nov.
Most similar species are E. viridis, E. dispar, and E. glutinosae, from which it differs by having a shorter valvula 3 and lancet. Males distinguishable from other species by distinct penis valves.
COI. Based on 13 specimens, maximum within species distance is 3.19% and the nearest neighbour, diverging by a minimum of 0%, are dispar, viridis, and possibly glutinosae. BINs: BOLD:AEC8057, BOLD:ABZ5797, BOLD:ACF5540 (main cluster).
Nuclear. Based on 10 (only NaK) or 8 (NaK and POL2) specimens, maximum within species distance is 1.09% (only NaK) or 0.92% (NaK and POL2) and 0.12% based on haplotypes of individual females. The nearest neighbour, diverging by a minimum of 0% (only NaK) or 0.41% (NaK and POL2), is viridis. The 0% distance between viridis and brevivalvis for NaK is because one of the haplotypes of one female of viridis (
Hosts: Betula species, B. pendula (
Palaearctic (
Nematus brevivalvis Thomson, 1871. Lectotype, here designated, ♀, DEI-GISHym88904,
Pteronus kriegeri Konow, 1903a. LT, ♀, GBIF-GISHym3847,
Amauronematus spurcus Konow, 1904. Lectotype, here designated, ♀, GBIF-GISHym3848,
Pteronidea absimilis Lindqvist, 1949. HT, ♀, http://id.luomus.fi/GL.3465,
Pteronidea woollatti Lindqvist, 1971. HT, ♀, http://id.luomus.fi/GL.9217,
Nematus dispar
Zaddach, 1876: Plate II(5), 5. Type locality: Heubude (according to
Pteronidea pseudodispar Lindqvist, 1969: 242–245. Type locality: Finland, Helsinki, Munksnäs (HT). Syn. nov.
Most similar species are E. viridis and E. glutinosae, from which it possibly differs by having slightly larger denticles of serrulae. Males distinguishable from other species by their distinctive penis valves.
COI. Based on 11 specimens, maximum within species distance is 1.75% and the nearest neighbours, diverging by a minimum of 0%, are brevivalvis, glutinosae, and viridis. BINs: BOLD:AEC8057, BOLD:ABZ5797, BOLD:ACF5540 (main cluster).
Nuclear. Based on 6 specimens, maximum within species distance is 0.25% (0.15% based on haplotypes of individual females). The nearest neighbours, diverging by a minimum of 0.63%, are brevivalvis and glutinosae.
Hosts: Betula pendula (
Palaearctic (
Pteronidea pseudodispar Lindqvist, 1969. HT, ♀, http://id.luomus.fi/GL.3514,
Nematus glutinosae Cameron, 1882: 193–194. Type locality: Great Britain, England, Worcestershire. LT designated below.
Nematus viridissimus
Möller, 1882: 179. Type locality: Sweden, Skåne, Skrifvaremöllan vid Tvedörra lägerplats. LT designated by
Cameron’s article was published in February 1882, according to the issue wrappers and printed in the journal itself. Möller’s article was published no earlier than mid December 1882, based on an advertisement inside front issue wrapper dated December 1882, and the proceedings of the Entomologiska Föreningen meeting on 14 December 1882 published in the same issue (pp. 195–203). The seniority of the name glutinosae has previously been overlooked.
Most similar species are E. dispar and E. viridis. Compared to E. dispar it possibly has slightly smaller denticles of serrulae. Differences from E. viridis are not entirely clear. Male penis valves are most similar to E. viridis. There is possibly a larger gap between paravalva and valvispina compared to E. viridis.
COI. Based on 3 specimens, maximum within species distance is 0.15% and the nearest neighbour, diverging by a minimum of 0%, is dispar, but possibly also brevivalvis, and viridis. BINs: BOLD:ACF5540 (main cluster), but possibly also BOLD:AEC8057 and BOLD:ABZ5797.
Nuclear. Based on 2 specimens, maximum within species distance is 0% (0.02% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 0.63%, is dispar.
Hosts: Alnus species; A. glutinosa (
Nearly all published records are from the West Palaearctic (Europe, Georgia,
Nematus glutinosae Cameron, 1882. Lectotype, here designated, ♀, B.M.TYPE HYM.I.623,
Nematus viridissimus Möller, 1882. LT, ♀
Nematus leptocephalus Thomson, 1863: 632. Type locality: Scandinavia, “Lapponia intermedia” (HT).
Most similar species are E. reticulata and similar species in the flavescens group (character states in parentheses), from which it differs by having dorsal margin of lancet angulate at about second or third serrula from apex (not angulate) and apical sutures distinctly inclined apically (not inclined). Valvula 3 is also (usually) slightly narrower compared to flavescens group. Males distinguishable from other species by distinct penis valves.
COI. Based on 4 specimens, maximum within species distance is 0.76% (4.56% when including representatives from BOLD:AAG3563 and BOLD:ACY4317) and the nearest neighbour, diverging by a minimum of 6.23%, is sylvestris. BINs: BOLD:ADS7391 (Holarctic), possibly also Nearctic BOLD:AAG3563 and BOLD:ACY4317.
Nuclear. Based on 3 specimens, maximum within species distance is 0.07% (0.05% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 2.96%, is viridis.
Hosts: Salix spec. (
West Palaearctic and Nearctic (
Nematus respondens Förster, 1854b: 427–428. Type locality: Austria. Lectotype designated below.
Nematus nitens
Thomson, 1888: 1212. Type locality: Sweden, Lund. Lectotype designated by
Pteronus balassagloi Jakowlew, 1891: 23–24. Type locality: Uzbekistan, Tashkent. Lectotype designated by Zhelohovtsev (1976). Mentioned as a synonym of Nematus respondens by Zhelohovtsev (1976).
Nematus declaratus Muche, 1974: 105–107. Type locality: Uzbekistan, Fergana (HT). Syn. nov.
Nematus desantisi D.R. Smith, 1983: 260–262. Type locality: Argentina, Chubut, Valle del Rio Chubut (HT). Syn. nov.
Females are most similar to oligospila group, E. bergmanni, and E. sylvestris. Head more rounded in dorsal view and lancet somewhat narrower compared to E. bergmanni. Ventral parts of 2nd to 4th suture of lancet are oblique and more or less straight or weakly curved apically in E. respondens, but weakly or distinctly curved basally in oligospila group and E. sylvestris. Males distinguishable from other species by distinct penis valves.
COI. Based on 10 specimens, maximum within species distance is 3.19% and the nearest neighbour, diverging by a minimum of 5.44%, is viridis subgroup. BIN: BOLD:ABU8945.
Nuclear. Based on 3 specimens, maximum within species distance is 0.18% (0.27% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 3.82%, is viridis.
Hosts: a large number of Salix species, as well as sometimes Populus species (
Palaearctic, Neotropic, Afrotropic, Australasian (
Nematus respondens Förster, 1854b. Lectotype, here designated, ♂, GBIF-GISHym3404,
Nematus nitens Thomson, 1888. LT, ♀, DEI-GISHym88903,
Pteronus balassagloi Jakowlew, 1891. Lectotype, ♂, DEI-GISHym30223,
Nematus declaratus Muche, 1974. HT, ♂, GBIF-GISHym2826, ZMHB.
Nematus sylvestris
Cameron, 1884: 266. Type locality: not stated in original description (ST); according to
Pteronidea straminea Lindqvist, 1958: 103. Type locality: Nurmes, Finland (HT). Syn. nov.
Pteronidea angustiserra Lindqvist, 1969: 241–242. Type locality: Mustasaari, Finland (HT). Syn. nov.
Pteronidea disparoides Lindqvist, 1969: 245. Type locality: Espoo, Finland (HT). Syn. nov.
Females are most similar to oligospila group, E. respondens, and E. bergmanni, from which it differs usually by having a longer malar space. Lancet is usually narrower compared to E. bergmanni. Clypeus is usually less deeply emarginate compared to oligospila group. Ventral part of 2nd to 4th suture of lancet is weakly or distinclty curved basally in E. sylvestris, but oblique and more or less straight or weakly curved apically in E. respondens. Males distinguishable from other species by their relatively distinct penis valves.
COI. Based on 22 specimens, maximum within species distance is 3.68% (5.68% when including also Nearctic-only BINs) and the nearest neighbour, diverging by a minimum of 5.45%, is viridis subgroup. BINs: BOLD:AAG3515 (Holarctic), BOLD:AEH2646 (
Nuclear. Based on 14 specimens, maximum within species distance is 0.97% (0.83% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 2.72%, is viridis.
Hosts: Salix spp. and at least occasionally Populus tremula.
Holarctic (
Pteronidea straminea Lindqvist, 1958. HT, ♀, http://id.luomus.fi/GL.3512 (saw slide PR240.AZ lost?),
Pteronidea angustiserra Lindqvist, 1969. HT, ♀, http://id.luomus.fi/GL.3443,
Pteronidea disparoides Lindqvist, 1969. HT, ♀, http://id.luomus.fi/GL.3493,
Nematus viridis Stephens, 1835: 30. Type locality: England, London area (HT).
Nematus prasinus Hartig, 1837: 216–217. Type locality: not stated, but presumably Germany according to the title of Hartig’s publication (ST). LT designated below.
Nematus polyspilus Förster, 1854a: 284, 286–288. Type locality: Germany, Aachen area (ST). LT designated below.
Pteronidea breviseta Lindqvist, 1946: 181. Type locality: Scandinavia, Lapland (ST). No identifiable syntypes were located. Syn. nov.
Pteronidea breviseta Lindqvist, 1949: 75–77. Type locality: Finland, Utsjoki, Outakoski (HT). Syn. nov.
Pteronidea abscondita
Lindqvist, 1949: 77. Type locality: Finland, Kuusamo (HT) [according to original description in
Pteronidea lauroi Lindqvist, 1960a: 35. Type locality: Finland, Ahlainen, Rankku (HT). Syn. nov.
Most similar species are E. dispar and E. glutinosae. Compared to E. dispar it possibly has slightly smaller denticles of serrulae. Differences from E. glutinosae are not entirely clear. Male penis valves are most similar to E. glutinosae. There is possibly a smaller gap between paravalva and valvispina compared to E. glutinosae. Since differences from E. glutinosae are not clear, the new synonymies proposed here might not be correct. Fortunately, even if the synonymies are more accurately treated under E. glutinosae, it would not disrupt the usage of currently valid names. The Pteronidea abscondita “allotype” (not a type specimen) (http://id.luomus.fi/GL.3442) is most likely E. flavescens (Stephens, 1835).
COI. Based on 4 specimens, maximum within species distance is 0.46% and the nearest neighbour, diverging by a minimum of 0%, are brevivalvis, dispar, and possibly glutinosae. BINs: BOLD:AEC8057, BOLD:ABZ5797, BOLD:ACF5540 (main cluster).
Nuclear. Based on 4 specimens, maximum within species distance is 0.09% (0.55% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 0% (only NaK) or 0.41% (NaK and POL2), is brevivalvis.
Hosts: most host records under the name viridis and all those under breviseta refer to Betula, e.g. B. pendula (
Palaearctic (
Nematus viridis Stephens, 1835. HT, ♀, B.M.TYPE HYM.I-697,
Nematus prasinus Hartig, 1837. Lectotype, here designated, ♀, GBIF-GISHym3388,
Nematus polyspilus Förster, 1854. Lectotype, here designated, ♀, GBIF-GISHym3386,
Pteronidea breviseta Lindqvist, 1946. No identifiable syntypes found: the HT of P. breviseta Lindqvist, 1949 was collected in 1948, and so cannot be a syntype of the taxon described in 1946.
Pteronidea breviseta Lindqvist, 1949. HT, ♀, http://id.luomus.fi/GL.3461,
Pteronidea abscondita Lindqvist, 1949. HT, ♀, http://id.luomus.fi/GL.3441,
Pteronidea lauroi Lindqvist, 1960. HT, ♀, http://id.luomus.fi/GL.3506,
Pteronus pallens Konow, 1903: 310 (key). Type locality: Irkutsk, Russia (LT).
Most similar species are sylvestris and respondens, from which it differs by having more prominent serrulae (cf. Figs
East Palaearctic. Removed from the list of West Palaearctic taxa. Specimens studied are from Russia (Irkutsk Oblast).
Pteronus pallens Konow, 1903. LT, here designated, ♀, GBIF-GISHym3863,
Nematus (Pteronidea) pyramidalis
Hellén, 1948: 114. Two ST ♀♀ not found in
Seems to belong to the bergmanni rather than the oligospila group, because of its long malar space according to the original description (
The group is mostly defined based on phylogenetic analyses of sequence data (Fig.
Nematus frenalis
Thomson, 1888: 1210–1212. Type locality: Sweden, Jämtland, Åreskutan. LT designated by
Pteronus fastosus Konow, 1904: 262–263. Type locality: northern Russia, Kanin Peninsula. LT designated below.
Nematus (Pteronidea) fastosus var. ponojense Hellén, 1948: 115. Type locality: Russia, Kola Peninsula, Ponoj (ST). Lectotype designated below. Syn. nov.
Nematus (Pteronidea) fastosus var. punctiscuta Hellén, 1948: 115. Type locality: Finland, Kilpisjärvi. LT designated below. Syn. nov.
See the key couplets 10 (females) and 4 (males).
COI. Based on 11 specimens, maximum within species distance is 6.38% and the nearest neighbour, diverging by a minimum of 0.3%, is oligospila. BINs: BOLD:AEA7654 and BOLD:ABZ2416, but possibly also Nearctic BOLD:ACA8095 and BOLD:AAV4677.
Nuclear. Based on 10 specimens, maximum within species distance is 0.67% (0.53% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 0.72%, is oligospila.
Hosts: according to
Two generations in southern Finland according to
Palaearctic (
Nematus frenalis Thomson, 1888. LT, ♀, DEI-GISHym88902,
Pteronus fastosus Konow, 1904. Lectotype, here designated, ♀, GBIF-GISHym3851,
Nematus (Pteronidea) fastosus var. ponojense Hellén, 1948. Lectotype, here designated, ♀, http://id.luomus.fi/GL.3546,
Nematus (Pteronidea) fastosus var. punctiscuta Hellén, 1948. Lectotype, here designated, ♀, http://id.luomus.fi/GL.3544,
Nematus oligospilus
Förster, 1854a: 284–286. Type locality: Germany, Aachen area. LT designated by
Nematus mendicus
Walsh, 1866: 261–262. Type locality: not stated (ST, probably destroyed by fire:
Nematus trivittatus
Norton, 1867: 218. Type locality: Canada, Mackenzie River and Great Slave Lake; USA, Illinois (ST). Synonymy with Pteronidea mendica by
Nematus microcercus Thomson, 1871: 152. Type locality: Sweden, Lund. Lectotype designated below.
Nematus dorsivittatus
Cresson, 1880: 10. Type locality: USA, Nevada (ST). Synonymised with Nematus oligospilus by
Nematus salicivorus Cameron, 1882: 194–195. Type locality: Great Britain, England, Worcestershire. Lectotype designated below.
Pteronus koebelei
Marlatt, 1896: 44–46 (key), 71. Type locality: USA, California and Oregon (ST). Synonymy with Nematus mendicus by
Pteronidea vanduzeei
Rohwer, 1913: 280–281. Type locality: USA, Virginia, Chain Bridge (HT). Synonymy with Nematus mendicus by
Pteronidea elelea
MacGillivray, 1923: 162. Type locality: Canada, Alberta, Edmonton (ST). Synonymy with Nematus mendicus by
See the key couplets 10 (females) and 4 (males). Can be small, about 4.5 mm (
COI. Based on 12 specimens, maximum within species distance is 5.93% and the nearest neighbour, diverging by a minimum of 0.3%, is frenalis. BINs: BOLD:AAV4676, BOLD:ABZ2416, BOLD:AEA6205, BOLD:ABY8224 (DEI-GISHym11390), BOLD:AEA3640 (Russian Far East), and possibly European BOLD:ADW3220 and Nearctic BOLD:ACA8095, BOLD:AAV4677. BOLD:ABW6676 is an artefact because of chimeric sequences composed of BOLD:AEA6205 and a possible NUMT cluster. Additionally, some specimens in BOLD possibly belong to a NUMT cluster because of a stop codon and in some cases also indels:
Nuclear. Based on 10 specimens, maximum within species distance is 0.68% (0.52% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 0.72%, is frenalis.
Hosts: Salix species (
West Palaearctic (see Material studied, below), East Palaearctic (
Nematus oligospilus Förster, 1854. LT, ♀, GBIF-GISHym3372,
Nematus microcercus Thomson, 1871. Lectotype, here designated, ♂, MZLU2017330,
Nematus salicivorus Cameron, 1882. Lectotype, here designated, ♀, B.M.TYPE HYM.I.625,
Euura glutinosae (A, B) and possible dispar (C–E) larvae A,B France 2007 Alnus glutinosa (photos: Henri Savina) C Pyrenees 2010 Betula (photo: Henri Savina) D Finland 2020 swept larva E DEI-GISHym12579 Germany 2020 Betula pendula. The larvae with paired red dots (C, D) have earlier been considered as representing pseudodispar that we consider synonymous to dispar, but whether this type of larvae truly represent dispar remains unconfirmed.
Euura sylvestris larvae. A, B Finland 2018 Salix caprea C, D Finland 2019 Populus tremula E Austria 2017 Salix hegetschweileri. Larvae of sylvestris found on Populus tremula in Finland in 2019 appear morphologically different from larvae on Salix, but based on genetics represent sylvestris.
Euura oligospila (A–C) and possible respondens (D–F) larvae from Salix fragilis (unconfirmed). A Finland 2018 Salix caprea B, C Sweden 2018 swept larva D Finland 2020 Salix fragilis E, F Finland 2020 Salix fragilis. Larva of respondens remain unconfirmed but adults of this species occurred at exactly the same spot as the larva D
The genus Euura has been taxonomically challenging and probably will remain so for years to come, but fortunately progress is also continuously being made. Here we revised the taxonomy of the bergmanni and oligospila groups, containing the bulk of the “green” Euura. These “green” species have been notoriously difficult to identify. In addition to very similar morphology, taxonomic oversplitting of species has further complicated identification of species in both groups. The most recent attempt to revise the “green” species was by
One issue that still needs attention is taxonomic revision of Nearctic taxa of the oligospila and bergmanni groups, which could affect the use of names for Palaearctic species.
Although morphological differences between adults of different generations have already been observed in a few species of Tenthredinoidea, such as Pristiphora leucopus (Hellén, 1948) (
Funding by the Swedish Taxonomy Initiative (2012–2020) was the basis for this study. MP is supported by funding from the PUTTE programme by the Finnish Ministry of Environment. For the loan or gift of material, and access to collections, we thank Ewald Altenhofer (Etzen, Austria), Christer Hansson and Rune Bygebjerg (
List of specimens studied, available also as Suppl. material
C_ex GBIF.Nr catalogNumber scientificName scientificNameAuthorship lifeStage Sex individualCount country stateProvince locality decimalLatitude decimalLongitude minimumElevationInMeters maximumElevationInMeters eventDateBegin eventDateEnd recordedBy samplingProtocol identifiedBy identificationUncertain typeStatus originalCombination emergenceDateBegin emergenceDateEnd associatedTaxa institutionCode GenBankAccessionCOI GenBankAccessionNaK GenBankAccessionPOL2
179082 83979 DEI-GISHym83979 Euura abnormis (Holmgren, 1883) adult male 1 Norway Finnmark Grense Jakobselv 8 km S 69.7, 30.8 250 m 2019-06-24 A. Liston & M. Prous netting M. Prous
151358 84272 DEI-GISHym84272 Euura aegra (Konow, 1895) adult female 1 Finland Northern Ostrobothnia Juuma 2 km SE 66.261, 29.417 230 m 2018-06-10
152185 56635
158733 84060 DEI-GISHym84060 Euura annulata (Gimmerthal, 1834) adult male 1 Germany Thueringen Ilfeld: Netzkater: Brandesbachtal 51.6, 10.81 350 m 400 m 2019-05-30 23rd Symphyta Workshop netting M. Prous
166267 12212 DEI-GISHym12212 Euura anomaloptera (Foerster, 1854) adult female 1 Norway Finnmark Skoltebyen 3 km NW 69.697, 29.31 70 m 2019-06-26 A. Liston & M. Prous netting A. D. Liston
158186 83961 http://id.luomus.fi/GP.110174 Euura bergmanni (Dahlbom, 1835) adult male 1 Finland Uusimaa Helsinki 60.17555, 24.93416 1964-01-01 1964-12-31 E. O. Peltonen rearing M. Prous Salix pentandra
173497 108142
173503 108146
173583 110299
173601 110391
173692 110378
173838 108256
174482 106508
174499 106684
174546 106505
174547 106504
174604 106503
174605 106502
174606 106501
152924 57263
175197 109040
175211 109030
175382 111139
175568 108954
215668 12587 DEI-GISHym12587 Euura bergmanni (Dahlbom, 1835) adult female 1 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-24 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215669 12585 DEI-GISHym12585 Euura bergmanni (Dahlbom, 1835) adult female 1 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-24 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
176874 106693
158184 83942 http://id.luomus.fi/GP.110172 Euura bergmanni (Dahlbom, 1835) adult female 1 Finland Hame Paelkaene 61.33333, 24.26666 1959-06-01 J. Kangas rearing M. Prous Salix phylicifolia
158185 83960 http://id.luomus.fi/GP.110173 Euura bergmanni (Dahlbom, 1835) adult male 1 Finland Hame Paelkaene 61.33333, 24.26666 1983-06-19 J. Kangas rearing M. Prous Salix phylicifolia
152760 57099
152762 57101
152800 57139
152827 57166
215667 84689 DEI-GISHym84689 Euura bergmanni (Dahlbom, 1835) adult male 1 Finland Lapland Pousu 68.847 21.161 450 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
176993 107596
177115 107246
177118 107243
177688 107033
215709 84690 DEI-GISHym84690 Euura bergmanni (Dahlbom, 1835) adult female 1 Finland Lapland Saarikoski 1 km N 68.814, 21.238 440 m 2020-06-26 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
50633 3451 DEI-GISHym3451 Euura bergmanni (Dahlbom, 1835) adult male 1 Germany Nordrhein-Westfalen Aachen 50.76666, 6.1 M. Prous lectotype Nematus validicornis Foerster, 1854
80311 50479 BC
80312 50480 BC
217191 84733 DEI-GISHym84733 Euura bergmanni (Dahlbom, 1835) adult female 1 Germany Sachsen-Anhalt Dessau 51.85, 12.25 60 m E. Enslin lectotype Lygaeonematus pallens Enslin, 1916
217192 84734 DEI-GISHym84734 Euura bergmanni (Dahlbom, 1835) adult female 1 Germany Deutschland 51.5, 10.5 T. Hartig lectotype Nematus pallicarpus Hartig, 1837
51651 3339 DEI-GISHym3339 Euura bergmanni (Dahlbom, 1835) adult male 1 Germany Bayern Fuerth (Fuerth i. B. [= in Bayern] in coll.Enslin) 48.481, 10.969 lectotype Pteronidea curtispina var. luctuosa Enslin, 1916
49895 3456 DEI-GISHym3456 Euura bergmanni (Dahlbom, 1835) adult female 1 Germany Germany 51.5, 10.5 lectotype Nematus virescens Hartig, 1837
80266 50434 BC
50618 3849 DEI-GISHym3849 Euura bergmanni (Dahlbom, 1835) adult male 1 Russia Irkutsk 52.33333, 104.25 lectotype Amauronematus longicornis Konow, 1897
87284 31138 DEI-GISHym31138 Euura bergmanni (Dahlbom, 1835) adult female 1 Sweden Norrbottens Laen Abisko National Park, E10 68.353, 18.815 390 m 2012-06-16 A.D. Liston, A. Taeger & S.M. Blank netting M. Prous
69490 20294 DEI-GISHym20294 Euura bergmanni (Dahlbom, 1835) adult female 1 Sweden Gotlands Laen Gotlands kommun, Roleks, border between wood and open pasture 57.53677, 18.33787 2005-06-06 2005-07-05 SMTP - Swedish Malaise Trap Project Malaise trap A. Taeger SMTP KC974677 MW939725 MW939829
85614 20606 DEI-GISHym20606 Euura bergmanni (Dahlbom, 1835) adult female 1 Sweden Kopparbergs Laen Stenis 1 km N 60.947, 14.474 180 m 2013-06-15 A.D. Liston, M. Prous & A. Taeger netting A. D. Liston
155495 58933 MZLU2017334 Euura bergmanni (Dahlbom, 1835) adult female 1 Sweden Skane Vallby [Wallby], near Lund 55.65, 13.28333 0000-08-14 F. Koch lectotype Nematus curtispina Thomson, 1871
81971 52139 BC
216379 Euura bergmanni (Dahlbom, 1835) larva 1 United Kingdom Scotland Fife, Loch Gelly, South shore, VC85, NT205923 56.11562, -3.28381 2020-07-26 B. Little photography A. D. Liston Salix spec.
152815 57154
173199 107615
173287 108235
173693 110379
173824 110344
173907 110140
173979 109587
174100 109566
174109 107943
174153 110427
174278 109795
174391 108035
174507 106676
174582 106573
174726 110449
174807 106636
174810 106661
174847 110015
157367 59063
175110 106898
175142 106574
175182 111199
175300 111006
175302 110941
175306 110935
175307 110936
175308 110937
175314 110986
175315 110942
175342 110985
175478 108814
175481 108811
175485 108812
175486 108813
175585 108965
175686 111123
175691 111084
175692 111085
175701 111083
175718 111098
175722 111095
175724 111103
175726 111104
175735 111633
175802 108722
175804 108724
175948 106789
175949 106790
176049 106801
176050 106802
176051 106803
176545 111424
151655 56560 Prep.143.OR Euura bergmanni_group adult female 1 Finland Hame Paelkaene 61.33333, 24.26666 1959-06-17 M. Prous CVV
176878 106856
177842 107174
178135 111626
83913 54081 BC
83700 53868 BC
158810 12119 DEI-GISHym12119 Euura bergmanni_group adult male 1 Germany Thueringen Ilfeld: Netzkater: Brandesbachtal 51.6, 10.81 350 m 400 m 2019-05-31 23rd Symphyta Workshop netting M. Prous
80305 50473 BC
215430 84564 DEI-GISHym84564 Euura bergmanni_group adult male 1 Germany Brandenburg Muencheberg W, NSG Gumnitz 52.51183, 14.08223 2020-05-21 A. Liston & M. Prous netting M. Prous & A. Liston
83909 54077 BC
80945 51113 BC
80946 51114 BC
80947 51115 BC
87287 31141 DEI-GISHym31141 Euura bergmanni_group adult female 1 Sweden Norrbottens Laen Abisko National Park, E10 68.353, 18.815 390 m 2012-06-18 A.D. Liston, A. Taeger & S.M. Blank netting M. Prous
96685 31696 DEI-GISHym31696 Euura bigallae (Vikberg & Zinovjev, 2014) larva 1 Sweden Vaesterbotten, Norsjoe 5 km NNW 64.95492, 19.43269 2016-06-20 A.D. Liston photography A. D. Liston Salix caprea
94430 80206 DEI-GISHym80206 Euura bohemani (Thomson, 1871) adult male 1 Sweden Torne Lappmark Bjoerkliden 68.409, 18.639 500 m 600 m 2016-07-09 A. Liston & M. Prous netting M. Prous
151183 84190 DEI-GISHym84190 Euura brevivalvis (Thomson, 1871) adult female 1 Estonia Raplamaa Kastna 1 km N 58.863, 25.052 70 m 2018-05-22 M. Prous netting M. Prous
68090 11621 DEI-GISHym11621 Euura brevivalvis (Thomson, 1871) adult female 1 Finland Lapin Laeaeni Kevo 69.75, 27.01666 2001-06-09 2001-06-11 Kevo Subarctic Research Institute rearing A. D. Liston
68093 11624 DEI-GISHym11624 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapin Laeaeni Kevo 69.75, 27.01666 2001-06-09 2001-06-11 Kevo Subarctic Research Institute rearing A. D. Liston
75105 17593 DEI-GISHym17593 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapin Laeaeni Kevo 69.75, 27.01666 2001-06-09 2001-06-11 Kevo Subarctic Research Institute rearing A. Taeger
75106 17594 DEI-GISHym17594 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapin Laeaeni Kevo 69.75, 27.01666 2001-06-09 2001-06-11 Kevo Subarctic Research Institute rearing A. Taeger
68809 56573 http://id.luomus.fi/GL.9217 Euura brevivalvis (Thomson, 1871) adult female 1 Finland Lapin Laeaeni Kilpisjaervi 69.05, 20.8 1968-07-09 L. H. Woollatt holotype Pteronidea woollatti Lindqvist, 1971
215462 84593 DEI-GISHym84593 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapland Kilpisjaervi, Jehkas 69.086, 20.8 550 m 950 m 2020-06-28 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215672 84520 DEI-GISHym84520 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapland Kilpisjaervi, Jehkas 69.086, 20.8 550 m 950 m 2020-06-28 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215673 Euura brevivalvis (Thomson, 1871) adult 17 Finland Lapland Kilpisjaervi, Jehkas 69.086, 20.8 550 m 950 m 2020-06-28 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215555 84654 DEI-GISHym84654 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapland Kilpisjaervi, Malla 69.065, 20.722 500 m 850 m 2020-06-25 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215679 Euura brevivalvis (Thomson, 1871) adult 9 Finland Lapland Kilpisjaervi, Malla 69.065, 20.722 500 m 850 m 2020-06-25 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215674 Euura brevivalvis (Thomson, 1871) adult female 1 Finland Lapland Kilpisjaervi, Saana 69.037, 20.844 500 m 900 m 2020-06-29 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215493 84633 DEI-GISHym84633 Euura brevivalvis (Thomson, 1871) adult male 2 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215675 84494 DEI-GISHym84494 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215676 Euura brevivalvis (Thomson, 1871) adult 5 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215677 Euura brevivalvis (Thomson, 1871) adult 4 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-24 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
152532 56871
151400 84227 DEI-GISHym84227 Euura brevivalvis (Thomson, 1871) adult male 1 Finland Northern Ostrobothnia Oulanka station 5 km E 66.38, 29.426 230 m 2018-06-02 A. Liston & M. Prous netting M. Prous
64908 58962 http://id.luomus.fi/GL.3465 Euura brevivalvis (Thomson, 1871) adult female 1 Finland Pihtipudas 63.38333, 25.56666 1946-06-08 1946-06-09 E. Lindqvist M. Prous holotype Pteronidea absimilis Lindqvist, 1949
215678 Euura brevivalvis (Thomson, 1871) adult female 1 Finland Lapland Pousu 68.847, 21.161 450 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
151399 84214 DEI-GISHym84214 Euura brevivalvis (Thomson, 1871) adult female 1 Finland Northern Ostrobothnia Pudasjaervi 18 km E 65.409, 27.371 130 m 2018-06-01 A. Liston & M. Prous netting M. Prous
151295 56524 PR.586VV Euura brevivalvis (Thomson, 1871) adult female 1 Finland Lapin Laeaeni Utsjoki 69.88333 27 1960-06-08 V. Vikberg V. Vikberg CVV
151296 56525 PR.587VV Euura brevivalvis (Thomson, 1871) adult female 1 Finland Lapin Laeaeni Utsjoki 69.88333 27 1960-06-07 V. Vikberg V. Vikberg CVV
157334 59032
157335 59033
152507 56846
67818 11380 DEI-GISHym11380 Euura brevivalvis (Thomson, 1871) adult female 1 France Aulus-les-Bains 42.8, 1.33333 2011-05-05 H. Savina M. Prous
51848 3847 DEI-GISHym3847 Euura brevivalvis (Thomson, 1871) adult female 1 Germany Saxony-Anhalt Dornreichenbach 51.36666, 12.86666 1894-05-06 G. Enderlein lectotype Pteronus kriegeri Konow, 1903
216303 Euura brevivalvis (Thomson, 1871) adult 2 Norway Finnmark Grense Jakobselv 8 km S 69.7, 30.8 250 m 2019-06-24 A. Liston & M. Prous netting M. Prous
50628 3848 DEI-GISHym3848 Euura brevivalvis (Thomson, 1871) adult female 1 Russia Russia bor. lectotype Amauronematus spurcus Konow, 1904
87281 31135 DEI-GISHym31135 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Norrbottens Laen Abisko 9 km E (Stordalen) 68.35, 19.035 400 m 2012-06-17 A.D. Liston, A. Taeger & S.M. Blank netting M. Prous
146782 83696 DEI-GISHym83696 Euura brevivalvis (Thomson, 1871) larva 1 Sweden Norrbottens Laen Abisko 9 km E (Stordalen) 68.35, 19.035 400 m 2017-08-05 A. Liston & M. Prous M. Prous Betula nana
146785 83698 DEI-GISHym83698 Euura brevivalvis (Thomson, 1871) larva 1 Sweden Norrbottens Laen Abisko 9 km E (Stordalen) 68.35, 19.035 400 m 2017-08-05 A. Liston & M. Prous M. Prous Betula nana
96482 21375 DEI-GISHym21375 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Torne Lappmark Abisko: Mt. Njulla 68.362, 18.73 400 m 950 m 2016-06-30 A. Liston & M. Prous netting M. Prous
96486 21378 DEI-GISHym21378 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Torne Lappmark Abisko: Mt. Njulla 68.362, 18.73 400 m 950 m 2016-06-30 A. Liston & M. Prous netting M. Prous
155522 88904 DEI-GISHym88904 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Kopparbergs Laen Dalarna alpina C. H. Boheman M. Prous lectotype Nematus brevivalvis Thomson, 1871
87381 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Norrbottens Laen Kaunisvaara 2 km N 67.4, 23.345 170 m 2014-06-07 A. Liston & M. Prous netting A. D. Liston
87283 31137 DEI-GISHym31137 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Norrbottens Laen Kiruna nr. airport 67.84, 20.35 450 m 2012-06-21 A.D. Liston, A. Taeger & S.M. Blank netting M. Prous
87382 Euura brevivalvis (Thomson, 1871) adult female 2 Sweden Norrbottens Laen Muodoslompolo 10 km SW 67.894, 23.232 270 m 2014-06-13 A. Liston & M. Prous netting A. D. Liston
87378 Euura brevivalvis (Thomson, 1871) adult female 1 Sweden Norrbottens Laen Oeverkalix 22 km W 66.295, 22.356 140 m 2014-06-02 A. Liston & M. Prous netting A. D. Liston
87379 Euura brevivalvis (Thomson, 1871) adult female 2 Sweden Norrbottens Laen Oeverkalix 22 km W 66.295, 22.356 140 m 2014-06-04 A. Liston & M. Prous netting A. D. Liston
87380 Euura brevivalvis (Thomson, 1871) adult female 4 Sweden Norrbottens Laen Pajala 67.204, 23.409 150 m 2014-06-05 A. Liston & M. Prous netting A. D. Liston
150906 84184 DEI-GISHym84184 Euura clibrichella (Cameron, 1878) adult male 1 Sweden Torne Lappmark Bjoerkliden 68.409, 18.639 500 m 850 m 2016-07-01 A. Liston & M. Prous netting M. Prous
63033 Euura dispar (Zaddach, 1876) larva 1 Europe Pyrenaeen 42.5 2 2010-07-24 H. Savina photography H. Savina Betula spec.
152770 57109
152516 56855
152517 56856
166360 84146 DEI-GISHym84146 Euura dispar (Zaddach, 1876) adult female 1 Finland Kitee, Papinniemi [6883:3656] 62.025, 29.99 2019-06-12 A.D. Liston netting A. D. Liston
178870 111659 http://id.luomus.fi/GL.3514 Euura dispar (Zaddach, 1876) adult female 1 Finland Uusimaa Munksnaes 60.19944, 24.87527 1967-08-11 E. Lindqvist E. Lindqvist holotype Pteronidea pseudodispar Lindqvist, 1969
152508 56847
158337 83956 DEI-GISHym83956 Euura dispar (Zaddach, 1876) adult female 1 Germany Sachsen-Anhalt Blankenburg/Harz: Steinkoepfe 51.739, 10.967 420 m 2019-06-01 23rd Symphyta Workshop netting M. Prous
158338 83953 DEI-GISHym83953 Euura dispar (Zaddach, 1876) adult female 1 Germany Sachsen-Anhalt Blankenburg/Harz: Steinkoepfe 51.739, 10.967 420 m 2019-06-01 23rd Symphyta Workshop netting M. Prous
158339 Euura dispar (Zaddach, 1876) adult female 1 Germany Sachsen-Anhalt Blankenburg/Harz: Steinkoepfe 51.739, 10.967 420 m 2019-06-01 23rd Symphyta Workshop netting M. Prous
148324 88519 DEI-GISHym88519 Euura dispar (Zaddach, 1876) adult male 1 Germany Brandenburg Eberswalde, Waldstrasse 19: Garten mit Waldrand 52.8264, 13.84136 2017-05-18 2017-05-25 A. Taeger Malaise trap
215418 12579 DEI-GISHym12579 Euura dispar (Zaddach, 1876) larva 1 Germany Brandenburg Hoppegarten 52.518, 13.674 2020-06-01 A. Liston & M. Prous netting M. Prous ? Betula pendula
80306 50474 BC
75108 17596 DEI-GISHym17596 Euura dispar (Zaddach, 1876) adult male 1 Germany Brandenburg Noerdlich Golzow, 5.M 52.96667, 13.804 1994-06-01 DEI-Projekt S. M. Blank
80274 50442 BC
80307 50475 BC
80308 50476 BC
67743 11328 DEI-GISHym11328 Euura dispar (Zaddach, 1876) adult female 1 Germany Mecklenburg-Vorpommern Ranzin, Oldenb. Holz 53.95114, 13.57204 2008-08-16 H.-J. Jacobs netting M. Prous ?
63245 11633 DEI-GISHym11633 Euura dispar (Zaddach, 1876) adult male 1 Italy Sicilia Etna Massif North, ca. 10 km SW Linguaglossa, Piano Provenzana 37.8, 15.03 1600 m 2010-05-16 A.D. Liston netting A. D. Liston
76419 19632 DEI-GISHym19632 Euura dispar (Zaddach, 1876) adult male 1 Portugal Viana do Castelo Mon‡ao 5 km SSE 42.04128, -8.44533 300 m 2012-05-13 Blank, Jacobs, Liston & Taeger netting M. Prous
76420 19631 DEI-GISHym19631 Euura dispar (Zaddach, 1876) adult female 1 Portugal Viana do Castelo Mon‡ao 5 km SSE 42.04128, -8.44533 300 m 2012-05-13 Blank, Jacobs, Liston & Taeger netting M. Prous
149748 21499 DEI-GISHym21499 Euura dispar (Zaddach, 1876) adult male 1 Sweden Torne Lappmark Bjoerkliden 68.409, 18.639 500 m 600 m 2016-07-09 A. Liston & M. Prous netting
131807 21388 DEI-GISHym21388 Euura dispar (Zaddach, 1876) adult male 1 Sweden Torne Lappmark Bjoerkliden 2 km SE 68.395, 18.705 350 m 2016-07-01 A. Liston & M. Prous netting M. Prous
63275 Euura dispar (Zaddach, 1876) larva 1 United Kingdom Scotland Dundreggan 57.2, -4.76666 2010-08-14 2010-08-21 G. T. Knight photography A. D. Liston Betula spec.
151389 84279 DEI-GISHym84279 Euura excisa (Thomson, 1863) adult female 1 Finland Northern Ostrobothnia Oulanka station 13 km E 66.361, 29.596 280 m 2018-06-12 A. Liston & M. Prous netting M. Prous
133677 83588 DEI-GISHym83588 Euura fahraei (Thomson, 1863) larva 1 Germany Brandenburg Muencheberg, Dahmsdorf 52.53333, 14.1 2017-05-28 A.D. Liston A. D. Liston Populus tremula
151185 84191 DEI-GISHym84191 Euura flavescens (Stephens, 1835) adult male 1 Estonia Raplamaa Kastna 1 km N 58.863, 25.052 70 m 2018-05-22 M. Prous netting
151640 56553 http://id.luomus.fi/GL.3442 Euura flavescens (Stephens, 1835) adult male 1 Finland Lapponia inarensis Inari 69 28 1967-06-28 M. Prous ?
68826 111737 http://id.luomus.fi/GL.3544 Euura frenalis (Thomson, 1888) adult female 1 Finland Lapin Laeaeni Kilpisjaervi 69.05, 20.8 1935-07-11 lectotype Nematus (Pteronidea) fastosus var. punctiscuta Hellen, 1948
215698 Euura frenalis (Thomson, 1888) adult female 1 Finland Lapland Kilpisjaervi, Malla 69.065, 20.722 500 m 850 m 2020-06-25 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston ?
175171 111195
152534 56873
151404 84232 DEI-GISHym84232 Euura frenalis (Thomson, 1888) adult male 1 Finland Northern Ostrobothnia Oulanka station 8 km E 66.367, 29.503 270 m 2018-06-02 A. Liston & M. Prous netting M. Prous
151656 56561 Prep.108.OR Euura frenalis (Thomson, 1888) adult female 1 Finland Hame Paelkaene 61.33333, 24.26666 1959-06-16 CVV
215696 12616 DEI-GISHym12616 Euura frenalis (Thomson, 1888) adult male 1 Finland Lapland Pousu 68.847, 21.161 450 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston ?
215697 12604 DEI-GISHym12604 Euura frenalis (Thomson, 1888) adult male 1 Finland Lapland Pousu 68.847, 21.161 450 m 2020-06-26 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
151403 84220 DEI-GISHym84220 Euura frenalis (Thomson, 1888) adult male 1 Finland Northern Ostrobothnia Taivalkoski 22 km NE 65.722, 28.569 250 m 2018-06-01 A. Liston & M. Prous netting M. Prous
151405 84222 DEI-GISHym84222 Euura frenalis (Thomson, 1888) adult male 1 Finland Northern Ostrobothnia Taivalkoski 22 km NE 65.722, 28.569 250 m 2018-06-01 A. Liston & M. Prous netting M. Prous
151406 84219 DEI-GISHym84219 Euura frenalis (Thomson, 1888) adult male 1 Finland Northern Ostrobothnia Taivalkoski 22 km NE 65.722, 28.569 250 m 2018-06-01 A. Liston & M. Prous netting M. Prous
158898 12057 DEI-GISHym12057 Euura frenalis (Thomson, 1888) adult male 1 Norway Finnmark Batsfjord 33 km SW 70.47, 28.97 280 m 2019-06-27 A. Liston & M. Prous netting M. Prous
50638 3851 DEI-GISHym3851 Euura frenalis (Thomson, 1888) adult female 1 Russia Kanin, Halbinsel 68 45 B. Poppius lectotype Pteronus fastosus Konow, 1904
68827 111964 http://id.luomus.fi/GL.3546 Euura frenalis (Thomson, 1888) adult female 1 Russia Murmanskaya Oblast Kola-Halbinsel, Ponoj 67.06666, 41.11666 1913-07-14 M. Prous lectotype Nematus (Pteronidea) fastosus var. ponojense Hellen, 1948
150441 88724 DEI-GISHym88724 Euura frenalis (Thomson, 1888) adult male 1 Sweden Torne Lappmark Abisko: Mt. Njulla 68.362, 18.73 400 m 950 m 2016-06-30 A. Liston & M. Prous netting M. Prous
87275 31129 DEI-GISHym31129 Euura frenalis (Thomson, 1888) adult female 1 Sweden Norrbottens Laen Abisko: Mt. Njulla above treeline 68.362, 18.73 700 m 900 m 2012-06-29 A.D. Liston & A. Taeger netting M. Prous ?
87278 31132 DEI-GISHym31132 Euura frenalis (Thomson, 1888) adult female 1 Sweden Norrbottens Laen Abisko: Mt. Njulla above treeline 68.362, 18.723 900 m 2012-07-05 A.D. Liston & A. Taeger netting A. D. Liston
87279 31133 DEI-GISHym31133 Euura frenalis (Thomson, 1888) adult female 1 Sweden Norrbottens Laen Abisko: Mt. Njulla above treeline 68.362, 18.723 900 m 2012-07-05 A.D. Liston & A. Taeger netting A. D. Liston
155524 88902 DEI-GISHym88902 Euura frenalis (Thomson, 1888) adult female 1 Sweden Jamtlands Lan Areskutan 63.43333, 13.1 C. G. Thomson F. Koch lectotype Nematus frenalis Thomson, 1888
158214 12026 DEI-GISHym12026 Euura frenalis (Thomson, 1888) adult male 1 Sweden Norrbottens Laen Karesuando 27 km SE 68.249, 22.879 310 m 2014-06-13 A. Liston & M. Prous netting M. Prous
69847 20422 DEI-GISHym20422 Euura frenalis (Thomson, 1888) adult female 1 Sweden Norrbottens Laen Kiruna kommun, Abisko nationalpark, Nuolja Kalfjaell 68.35958, 18.71927 900 m 2006-06-26 2006-07-15 SMTP - Swedish Malaise Trap Project Malaise trap M. Prous SMTP
213598 12556 DEI-GISHym12556 Euura frenalis (Thomson, 1888) adult female 1 Sweden Torne Lappmark Nikkaluokta 7 km NE 67.89, 19.138 500 m 1050 m 2016-07-08 A. Liston & M. Prous netting M. Prous ?
131840 21332 DEI-GISHym21332 Euura frenalis (Thomson, 1888) adult male 1 Sweden Torne Lappmark: Rensjoen, near Kiruna 68.068, 19.022 2016-06-22 A.D. Liston netting M. Prous
158846 83976 DEI-GISHym83976 Euura freyja (Liston, Taeger & Blank, 2009) adult female 1 Norway Finnmark Grense Jakobselv 8 km S 69.7, 30.8 250 m 2019-06-24 A. Liston & M. Prous netting A. D. Liston
146879 80498 DEI-GISHym80498 Euura glutinosae Cameron, 1882 adult female 1 Austria Lower Austria Etzen 48.11666, 15.23333 2016-09-06 E. Altenhofer rearing M. Prous 2017-08-06 Alnus glutinosa
67843 11403 DEI-GISHym11403 Euura glutinosae Cameron, 1882 adult female 1 France Aulus-les-Bains 42.8, 1.33333 2010-07-01 H. Savina rearing A. D. Liston Alnus spec.
62541 Euura glutinosae Cameron, 1882 larva 1 France Bonac-sur-Lez 42.88333, 0.96666 2008-09-01 2008-09-30 H. Savina A. D. Liston Alnus glutinosa FOTO
151566 84469 DEI-GISHym84469 Euura glutinosae Cameron, 1882 larva 1 Germany Rhineland Palatinate Schoenenberg-Kuebelberg, 1 km S 49.3988, 7.3865 250 m 2018-07-14 M. Prous M. Prous ? Alnus glutinosa
158771 84096 DEI-GISHym84096 Euura glutinosae Cameron, 1882 adult male 1 Germany Sachsen-Anhalt Stolberg: Grube Luise, Bachtal 51.556, 10.976 350 m 2019-05-28 23rd Symphyta Workshop netting M. Prous
216953 Euura glutinosae Cameron, 1882 adult female 1 Russia Moskowskaja Oblast Sergiev-Posad district, Zolotilovo village 56.275, 37.987 2019-08-24 A. Boldyrev A. Boldyrev 2020-03-20 Alnus spec. FOTO
216955 Euura glutinosae Cameron, 1882 adult female 1 Russia Moskowskaja Oblast Sergiev-Posad district, Zolotilovo village 56.275, 37.987 2019-09-01 A. Boldyrev A. Boldyrev 2020-03-14 Alnus spec. FOTO
151604 111986
216659 111909 B.M.TYPE HYM1.623 Euura glutinosae Cameron, 1882 adult female 1 United Kingdom England Worcestershire [original label], Worcester [original description] 1875-09-21 J. E. Fletcher rearing lectotype Nematus glutinosae Cameron, 1882 Alnus glutinosa
133501 80261 DEI-GISHym80261 Euura humeralis (Serville, 1823) adult male 1 Germany Brandenburg Strausberg 3 km E 52.566, 13.934 80 m 2017-04-09 A. Liston & M. Prous netting M. Prous
166674 12505 DEI-GISHym12505 Euura hypoxantha (Foerster, 1854) adult female 1 Austria Lower Austria Rappottenstein 48.51666, 15.06666 2002-08-07 E. Altenhofer rearing A. Taeger 2003-05-13 Salix purpurea
166818 Euura hypoxantha (Foerster, 1854) adult male 1 Estonia Dorpat 58.36611, 26.73611 1883-06-01 M. Prous
166819 Euura hypoxantha (Foerster, 1854) adult male 1 Estonia Dorpat 58.36611, 26.73611 1883-05-08 M. Prous
173750 110182
155486 58924
152804 57143
176717 106507
176718 106506
49889 3301 DEI-GISHym3301 Euura hypoxantha (Foerster, 1854) adult male 1 Germany Nordrhein-Westfalen Aachen 50.76666, 6.1 M. Prous syntype Nematus hypoxanthus Foerster, 1854
80289 50457 BC
166479 83946 DEI-GISHym83946 Euura hypoxantha (Foerster, 1854) adult male 1 Germany Thueringen Ilfeld: Netzkater: Brandesbachtal 51.6, 10.81 350 m 400 m 2019-05-31 23rd Symphyta Workshop netting M. Prous
80288 50456 BC
166264 12209 DEI-GISHym12209 Euura hypoxantha (Foerster, 1854) adult male 1 Norway Finnmark Skoltebyen 3 km NW 69.697, 29.31 70 m 2019-06-26 A. Liston & M. Prous netting M. Prous
213605 12054 DEI-GISHym12054 Euura hypoxantha (Foerster, 1854) adult male 1 Norway Finnmark Skoltebyen 3 km NW 69.697, 29.31 70 m 2019-06-26 A. Liston & M. Prous netting M. Prous
216301 Euura hypoxantha (Foerster, 1854) adult male 1 Norway Finnmark Skoltebyen 3 km NW 69.697, 29.31 70 m 2019-06-26 A. Liston & M. Prous netting M. Prous
152120 86211 DEI-GISHym86211 Euura hypoxantha (Foerster, 1854) adult male 1 Russia Primorskiy Kray Gornotajozhnoe, 1 km E 43.694, 132.168 150 m 2016-05-22 K. Kramp, M. Prous & A. Taeger netting M. Prous
158245 80649 DEI-GISHym80649 Euura hypoxantha (Foerster, 1854) adult female 1 Russia Primorsky Krai Steklyanukha 43.349, 132.458 40 m 2019-05-07 M. Prous netting M. Prous ?
152128 86752 DEI-GISHym86752 Euura hypoxantha (Foerster, 1854) adult female 1 Russia Primorsky Krai Zolotaya Dolina E 42.943, 133.161 40 m 2016-06-10 K. Kramp, M. Prous & A. Taeger netting
155485 58923
66030 11928 DEI-GISHym11928 Euura infirma (Foerster, 1854) adult male 1 Germany Thueringen Brandesbachtal bei Netzkater N Ilfeld, Wiese, mittlere Hanglage 51.6, 10.81 350 m 2010-05-23 A. Taeger A. Taeger
158222 80655 DEI-GISHym80655 Euura krausi (Taeger & Blank, 1998) adult female 1 Russia Primorskiy Kray Anisimovka: Gribanovka 1 km N 43.126, 132.797 450 m 2019-05-08 M. Prous netting M. Prous
175205 108977
215726 84503 DEI-GISHym84503 Euura leptocephalus (Thomson, 1863) adult female 1 Finland Lapland Leutsuvaara 68.917, 20.933 550 m 700 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
215725 12610 DEI-GISHym12610 Euura leptocephalus (Thomson, 1863) adult female 1 Finland Lapland Pousu 68.847, 21.161 450 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
157411 Euura leptocephalus (Thomson, 1863) adult female 1 Finland Utsjoki, Outakoski 69.6, 25.96666 1948-06-22 E. Lindqvist M. Prous
213701 12065 DEI-GISHym12065 Euura leptocephalus (Thomson, 1863) adult male 1 Norway Finnmark Batsfjord 9 km SW 70.568, 29.554 230 m 2019-06-27 A. Liston & M. Prous netting M. Prous
131804 21386 DEI-GISHym21386 Euura leptocephalus (Thomson, 1863) adult female 1 Sweden Torne Lappmark Abisko: Mt. Njulla 68.362, 18.73 400 m 950 m 2016-06-30 A. Liston & M. Prous netting M. Prous
150447 88730 DEI-GISHym88730 Euura leptocephalus (Thomson, 1863) adult male 1 Sweden Torne Lappmark Abisko: Mt. Njulla 68.362, 18.73 400 m 950 m 2016-06-30 A. Liston & M. Prous netting M. Prous
69796 59083
178904 12535 DEI-GISHym12535 Euura leptocephalus (Thomson, 1863) adult female 1 Sweden Torne Lappmark Nikkaluokta 7 km NE 67.89, 19.138 500 m 1050 m 2016-07-08 A. Liston & M. Prous netting M. Prous
151423 84256 DEI-GISHym84256 Euura lichtwardti (Konow, 1903) adult male 1 Finland Northern Ostrobothnia Oulanka station 18 km SE 66.283, 29.653 150 m 2018-06-08
179080 12072 DEI-GISHym12072 Euura longicauda (Hellen, 1948) adult male 1 Norway Finnmark Batsfjord 2 km SW 70.624, 29.672 70 m 2019-06-27 A. Liston & M. Prous netting M. Prous
151424 56546
213610 12074 DEI-GISHym12074 Euura malaisei (Hellén, 1970) adult male 1 Norway Finnmark Batsfjord 2 km SW 70.624, 29.672 70 m 2019-06-27 A. Liston & M. Prous netting M. Prous
157574 12015 DEI-GISHym12015 Euura miltonota (Zaddach, 1883) adult female 1 Germany Berlin, Wuhletal 52.51, 13.572 2019-04-19 K. Kramp, A. Liston & M. Prous netting M. Prous
151385 84264 DEI-GISHym84264 Euura montana (Zaddach, 1883) adult male 1 Finland Lapland Tolva 7 km W 66.223, 28.552 330 m 450 m 2018-06-09
89412 31247 DEI-GISHym31247 Euura myosotidis (Fabricius, 1804) adult male 1 Estonia Laanemaa Virtsu 2 km E 58.574, 23.543 5 m 2015-06-06 A.D. Liston, M. Prous & A. Taeger netting A. D. Liston
150903 84182 DEI-GISHym84182 Euura nigricornis (Serville, 1823) adult female 1 Germany Brandenburg Trebnitz 52.53333, 14.21666 2017-05-21 A.D. Liston netting A. D. Liston
148321 88516 DEI-GISHym88516 Euura obducta (Hartig, 1837) adult female 1 Germany Brandenburg Eberswalde, Waldstrasse 19: Garten mit Waldrand 52.8264, 13.84136 2017-05-18 2017-05-25 A. Taeger Malaise trap
67481 17065 DEI-GISHym17065 Euura oligospila (Foerster, 1854) adult female 1 Austria Salzburg Alpendorf Dachstein-West 47.514, 13.451 950 m 2010-07-25 A.D. Liston & A. Taeger A. Taeger
89414 31226 DEI-GISHym31226 Euura oligospila (Foerster, 1854) adult female 1 Estonia Ida-Virumaa Ontika, 1.5 km NE 59.443, 27.315 50 m 2015-06-03 A.D. Liston, M. Prous & A. Taeger netting A. D. Liston
149738 31239 DEI-GISHym31239 Euura oligospila (Foerster, 1854) adult male 1 Estonia Parnumaa Riisa 1 km N 58.489, 24.982 25 m 2015-06-05 A.D. Liston, M. Prous & A. Taeger netting M. Prous
152784 57123
151657 56562 Prep.122.OR Euura oligospila (Foerster, 1854) adult female 1 Finland Uusimaa Helsinki 60.17555, 24.93416 1962-06-12 CVV
151671 56577 Prep.104.OR Euura oligospila (Foerster, 1854) adult female 1 Finland Uusimaa Helsinki, Villinki 60.15861, 25.11361 1961-08-05 CVV
152767 57106
155601 58983
175081 106948
175384 111137
175437 111191
175714 111090
175756 111164
175760 111160
67828 11390 DEI-GISHym11390 Euura oligospila (Foerster, 1854) adult female 1 France Bethmale 42.86666, 1.08333 2010-07-04 H. Savina A. D. Liston
68227 17894 DEI-GISHym17894 Euura oligospila (Foerster, 1854) adult male 1 France Bethmale 42.86666, 1.08333 2009-09-01 2009-09-30 H. Savina A. D. Liston 2010-05-20 Salix spec.
49891 3372 DEI-GISHym3372 Euura oligospila (Foerster, 1854) adult female 1 Germany Nordrhein-Westfalen Aachen Umgebung 50.76666, 6.1 lectotype Nematus oligospilus Foerster, 1854
215748 Euura oligospila (Foerster, 1854) adult 7 Germany Brandenburg Muencheberg, Seelower Str. 52.50167, 14.15334 2013-05-17 A.D. Liston M. Prous
215749 84695 DEI-GISHym84695 Euura oligospila (Foerster, 1854) adult male 1 Germany Brandenburg Muencheberg, Seelower Str. 52.50167, 14.15334 2013-05-17 A.D. Liston M. Prous
217266 84738 DEI-GISHym84738 Euura oligospila (Foerster, 1854) adult female 1 Germany Brandenburg Muencheberg, Seelower Str. 52.50167, 14.15334 2013-05-17 A.D. Liston M. Prous
67908 11482 DEI-GISHym11482 Euura oligospila (Foerster, 1854) adult female 1 Germany Brandenburg Praedikow 52.63333, 14.01666 2011-05-01 A.D. Liston netting A. D. Liston
152122 86305 DEI-GISHym86305 Euura oligospila (Foerster, 1854) adult female 1 Russia Primorsky Krai Arsenyev, Ski-Base Bodrost 44.122, 133.27 200 m 2016-05-26 K. Kramp, M. Prous & A. Taeger netting M. Prous
166478 80721 DEI-GISHym80721 Euura oligospila (Foerster, 1854) adult female 1 Russia Primorsky Krai Gornotajozhnoe, 2 km W 43.694, 132.135 80 m 2019-05-14 M. Prous netting M. Prous
166745 80754 DEI-GISHym80754 Euura oligospila (Foerster, 1854) adult female 1 Russia Primorsky Krai Gornotajozhnoe, 2 km W 43.694, 132.135 2019-05-16 M. Prous & S. Tuerk netting M. Prous
67907 11481 DEI-GISHym11481 Euura oligospila (Foerster, 1854) adult female 1 Slovakia Liptovsky Hradok ESE 10 km, Svarinska valley, Svarin S 49.00916, 19.853 695 m 730 m 2005-06-19 A.D. Liston netting A. D. Liston
149740 21410 DEI-GISHym21410 Euura oligospila (Foerster, 1854) adult female 1 Sweden Torne Lappmark Abisko 6 km E 68.348, 18.969 400 m 2016-07-04 A. Liston & M. Prous netting M. Prous
149739 21395 DEI-GISHym21395 Euura oligospila (Foerster, 1854) adult male 1 Sweden Torne Lappmark Abisko 6 km W 68.342, 18.691 650 m 900 m 2016-07-02 A. Liston & M. Prous netting M. Prous
152057 83843 DEI-GISHym83843 Euura oligospila (Foerster, 1854) larva 1 Sweden Torne Lappmark Abisko National Park, Abisko Oestra 68.35088, 18.83086 400 m 2018-08-29 A.D. Liston netting M. Prous
155494 88899 DEI-GISHym88899 Euura oligospila (Foerster, 1854) adult female 1 Sweden Skane Igelosa 55.763, 13.274 1835-08-16 rearing M. Prous
217184 111959 MZLU2017330 Euura oligospila (Foerster, 1854) adult male 1 Sweden Malmoehus Laen Lund 55.7, 13.18333 lectotype Nematus microcercus Thomson, 1871
96476 21367 DEI-GISHym21367 Euura oligospila (Foerster, 1854) adult female 1 Sweden Torne Lappmark: Abisko: lowest 3 km of Abisko River 68.356, 18.769 350 m 2016-06-28 A.D. Liston netting A. D. Liston
216660 111910 B.M.TYPE HYM.1.625 Euura oligospila (Foerster, 1854) adult female 1 United Kingdom England Worcestershire [original label], Worcester [original description] 1877-10-01 1877-10-31 R. B. Benson Nematus salicivorus Cameron, 1882 Salix viminalis
50662 3863 DEI-GISHym3863 Euura pallens (Konow, 1903) adult female 1 Russia Irkutsk 52.33333, 104.25 lectotype Pteronus pallens Konow, 1903
50663 3864 DEI-GISHym3864 Euura pallens (Konow, 1903) adult female 1 Russia Irkutsk 52.33333, 104.25 paralectotype Pteronus pallens Konow, 1903
94847 84966 DEI-GISHym84966 Euura papillosa (Retzius, 1783) adult male 1 Japan Nagano Yamanouchi, Road 471/502 36.744, 138.524 1600 m 2016-07-17 A. Taeger netting M. Prous
93919 80108 DEI-GISHym80108 Euura parvilabris (Thomson, 1863) adult male 1 Sweden Torne Lappmark Bjoerkliden 68.409, 18.639 500 m 850 m 2016-07-01 A. Liston & M. Prous netting M. Prous
89415 31242 DEI-GISHym31242 Euura poecilonota (Zaddach, 1876) adult female 1 Estonia Viljandimaa Sandra 58.453, 25.037 25 m 2015-06-05 A.D. Liston, M. Prous & A. Taeger netting M. Prous
90029 31243 DEI-GISHym31243 Euura poecilonota (Zaddach, 1876) adult female 1 Estonia Viljandimaa Sandra 58.453, 25.037 25 m 2015-06-05 A.D. Liston, M. Prous & A. Taeger netting M. Prous
90031 Euura poecilonota (Zaddach, 1876) adult female 1 Estonia Viljandimaa Sandra 58.453, 25.037 25 m 2015-06-05 A.D. Liston, M. Prous & A. Taeger netting M. Prous
173133 106442
75131 17620 DEI-GISHym17620 Euura poecilonota (Zaddach, 1876) adult female 1 Finland Lapin Laeaeni Kevo 69.75, 27.01666 2001-06-03 2001-06-07 Kevo Subarctic Research Institute rearing A. Taeger Betula pubescens
178869 12527 DEI-GISHym12527 Euura poecilonota (Zaddach, 1876) adult male 1 Finland Lapin Laeaeni Kevo Subarctic Research Institute 69.75, 27.01659 240 m 2001-06-09 2001-06-11 Kevo Subarctic Research Institute M. Prous ? 2001-06-16 Betula pubescens
151401 84462 DEI-GISHym84462 Euura poecilonota (Zaddach, 1876) adult female 1 Finland Northern Ostrobothnia Oulanka station 18 km SE 66.283, 29.653 150 m 2018-06-13 A. Liston & M. Prous netting M. Prous
215755 Euura poecilonota (Zaddach, 1876) adult female 1 Finland Lapland Pousu 68.847, 21.161 450 m 2020-06-27 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
75154 17640 DEI-GISHym17640 Euura poecilonota (Zaddach, 1876) adult female 1 Germany Bayern Dirnberg bei Boebrach Muehle 49.05, 13.05 S. M. Blank M. Prous
80279 50447 BC
80280 50448 BC
1166 17621 DEI-GISHym17621 Euura poecilonota (Zaddach, 1876) adult female 1 Germany Brandenburg Noerdlich Britz, 6.M 52.91667, 13.804 1993-05-11 1993-05-13 DEI-Projekt Malaise trap A. Taeger
152113 86351 DEI-GISHym86351 Euura poecilonota (Zaddach, 1876) adult female 1 Russia Primorsky Krai Arsenyev, Ski-Base Bodrost 44.122, 133.27 200 m 2016-05-30 K. Kramp, M. Prous & A. Taeger netting M. Prous ?
152119 86176 DEI-GISHym86176 Euura poecilonota (Zaddach, 1876) adult female 1 Russia Primorskiy Kray Gornotajozhnoe, 1 km E 43.694, 132.168 150 m 2016-05-22 K. Kramp, M. Prous & A. Taeger netting M. Prous ?
152114 86727 DEI-GISHym86727 Euura poecilonota (Zaddach, 1876) adult female 1 Russia Primorsky Krai Nakhodka 10 km ESE: Beregovoj 42.788, 133.03 30 m 2016-06-07 K. Kramp, M. Prous & A. Taeger netting M. Prous ?
152115 86753 DEI-GISHym86753 Euura poecilonota (Zaddach, 1876) adult female 1 Russia Primorsky Krai Nikolaevka 5 km NNE 43.137, 133.249 140 m 2016-06-11 K. Kramp, M. Prous & A. Taeger netting M. Prous ?
149750 21400 DEI-GISHym21400 Euura poecilonota (Zaddach, 1876) adult female 1 Sweden Norrbottens Laen Abisko 9 km E (Stordalen) 68.35, 19.035 400 m 2016-07-04 A. Liston & M. Prous netting M. Prous
87286 31140 DEI-GISHym31140 Euura poecilonota (Zaddach, 1876) adult female 1 Sweden Norrbottens Laen Abisko National Park, E10 68.353, 18.815 390 m 2012-06-19 A.D. Liston netting M. Prous
87291 31145 DEI-GISHym31145 Euura poecilonota (Zaddach, 1876) adult female 1 Sweden Norrbottens Laen Abisko National Park, E10 68.353, 18.815 390 m 2012-06-23 A.D. Liston & A. Taeger netting M. Prous
210680 Euura poecilonota (Zaddach, 1876) adult female 1 Sweden Torne Lappmark Bjoerkliden 68.409, 18.639 500 m 850 m 2016-07-01 A. Liston & M. Prous netting M. Prous
210678 Euura poecilonota (Zaddach, 1876) adult female 1 Sweden Torne Lappmark: Abisko, Mt Njulla 68.365, 18.72 600 m 900 m 2016-06-23 A.D. Liston netting M. Prous
210679 Euura poecilonota (Zaddach, 1876) adult female 1 Sweden Torne Lappmark: Bjoerkliden 2 km SE 68.395, 18.705 500 m 800 m 2016-06-27 A.D. Liston netting M. Prous
155571 58973
151402 84261 DEI-GISHym84261 Euura punctifrons (Malaise, 1921) adult male 1 Finland Northern Ostrobothnia Oulanka station 8 km E 66.367, 29.503 270 m 2018-06-08
89193 31253 DEI-GISHym31253 Euura punicea (Christ, 1791) adult female 1 Estonia Laanemaa Karuse 1 km S 58.608, 23.695 25 m 2015-06-06 A.D. Liston, M. Prous & A. Taeger netting A. D. Liston
63749 Euura respondens (Foerster, 1854) adult female 1 Argentina Argentinien (Staat) 2003-11-15 S. M. Blank & C. Kutzscher S. M. Blank
68083 11613 DEI-GISHym11613 Euura respondens (Foerster, 1854) adult female 1 Argentina Rio Negro El Condor near Viedma -40.8, -63 2001-12-05 2001-12-14 C. Kutzscher netting S. M. Blank
53155 Euura respondens (Foerster, 1854) adult female 1 Argentina Jujuy Tilcara N 1 km -23.5605, -65.39166 2003-10-29 S. M. Blank S. M. Blank Salix babylonica
68070 11597 DEI-GISHym11597 Euura respondens (Foerster, 1854) adult female 1 Argentina Jujuy Tilcara N 1 km -23.5605, -65.39166 2003-10-29 S. M. Blank S. M. Blank Salix babylonica
157315 19889 DEI-GISHym19889 Euura respondens (Foerster, 1854) adult female 1 Argentina Jujuy Tilcara N 1 km -23.5605, -65.39166 2530 m 2003-10-29 S. M. Blank & C. Kutzscher A. D. Liston
68077 11605 DEI-GISHym11605 Euura respondens (Foerster, 1854) adult female 1 Australia Australian Capital Territory (ACT) Campbell -35.28809, 149.15415 2005-01-11 J. LaSalle S. M. Blank
68097 11629 DEI-GISHym11629 Euura respondens (Foerster, 1854) adult female 1 Australia Australian Capital Territory (ACT) Campbell -35.28809, 149.15415 2005-01-11 J. LaSalle S. M. Blank
79967 50135 BC
49893 3404 DEI-GISHym3404 Euura respondens (Foerster, 1854) adult male 1 Austria Austria 47.33333, 13.33333 syntype Nematus respondens Foerster, 1854
150436 84169 DEI-GISHym84169 Euura respondens (Foerster, 1854) adult female 1 Bulgaria Varna Goren Chiflik 43.014, 27.626 30 m 2018-04-13 A. Liston & M. Prous netting M. Prous
216266 84701 DEI-GISHym84701 Euura respondens (Foerster, 1854) adult female 1 Czech Republic Kralovehradecky kraj Bedrichovka 50.3, 16.43333 2009-06-09 J. Macek rearing M. Prous Salix fragilis
216268 Euura respondens (Foerster, 1854) adult male 1 Czech Republic Bil‚ Karpaty CHKO Machova res. (potok) 49.25, 17.55 2007-05-18 J. Macek M. Prous
216265 84700 DEI-GISHym84700 Euura respondens (Foerster, 1854) adult male 1 Czech Republic Stredocesky Kraj Lazec 49.68333, 13.95 2008-08-17 J. Macek M. Prous
215699 84697 DEI-GISHym84697 Euura respondens (Foerster, 1854) adult male 1 Finland Hame Janakkala 60.9, 24.58333 1987-01-01 1987-08-26 J. Kangas rearing M. Prous 1987-08-26 Salix fragilis
166483 59331 tO2 Euura respondens (Foerster, 1854) adult female 1 Finland Joensuu 2008-08-24 T. Nyman rearing M. Prous 2008-08-30 Salix fragilis NIBIO KF936599 KF935897
173521 108268
173524 108270
173525 108269
216175 Euura respondens (Foerster, 1854) adult male 1 Germany Brandenburg Fuerstenwalde 0000-06-12 rearing M. Prous ZMHB
152076 12514 DEI-GISHym12514 Euura respondens (Foerster, 1854) adult male 1 Germany Mecklenburg-Vorpommern Neubrandenburg 53.56666, 13.26666 30 m 1895-04-30 M. Prous
133779 80370 DEI-GISHym80370 Euura respondens (Foerster, 1854) adult female 1 Greece Elis Kryovrysi 37.9191, 21.8041 1000 m 2017-04-26
216267 Euura respondens (Foerster, 1854) adult female 1 Slovakia Moravsky Sv. Mikulas 2008-06-12 J. Macek M. Prous ? 2008-07-03 Salix alba
155523 88903 DEI-GISHym88903 Euura respondens (Foerster, 1854) adult female 1 Sweden Malmoehus Laen Lund 55.7, 13.18333 F. Koch lectotype Nematus nitens Thomson, 1888
216174 Euura respondens (Foerster, 1854) adult female 1 Uzbekistan Bakhmal 39.71666, 68.01666 1600 m 1979-05-21 1979-05-23 W. H. Muche M. Prous ZMHB
52078 2826 http://coll.mfn-berlin.de/u/dbcac9 Euura respondens (Foerster, 1854) adult male 1 Uzbekistan Fergana Oblast Fergana 40.38333, 71.76666 1973-05-21 1973-05-23 W. H. Muche holotype Nematus declaratus Muche, 1974 ZMHB
215629 84686 DEI-GISHym84686 Euura respondens (Foerster, 1854) adult female 1 Uzbekistan Fergana 10 km S 40.28, 71.8 1975-05-20 W. H. Muche M. Prous
216172 84694 DEI-GISHym84694 Euura respondens (Foerster, 1854) adult male 1 Uzbekistan Fergana 10 km S 40.28, 71.8 1975-05-20 W. H. Muche M. Prous
216173 Euura respondens (Foerster, 1854) adult 7 Uzbekistan Fergana 10 km S 40.28, 71.8 1975-05-20 W. H. Muche M. Prous ZMHB
216176 111833 http://coll.mfn-berlin.de/u/111027 Euura respondens (Foerster, 1854) adult female 1 Uzbekistan Fergana 10 km S 40.28, 71.8 1975-05-20 W. H. Muche M. Prous ZMHB
150058 30223 DEI-GISHym30223 Euura respondens (Foerster, 1854) adult male 1 Uzbekistan Taschkent 41.33333, 69.3 Balassoglo M. Prous lectotype Pteronus balassagloi Jakowlew, 1891
151299 56528 PR.588VV Euura reticulata (Holmgren, 1883) adult female 1 Finland Lapin Laeaeni Pallastunturi 68.1 24 1966-06-15 V. Vikberg V. Vikberg CVV
148466 88661 DEI-GISHym88661 Euura ribesii (Scopoli, 1763) adult male 1 Germany Brandenburg Eberswalde, Waldstrasse 19: Garten mit Waldrand 52.8264, 13.84136 2017-07-15 2017-07-28 A. Taeger Malaise trap
151354 84243 DEI-GISHym84243 Euura schlueteri (Enslin, 1915) adult male 1 Finland Northern Ostrobothnia Oulanka station 18 km SE 66.283, 29.653 150 m 2018-06-02 A. Liston & M. Prous netting M. Prous
93331 80058 DEI-GISHym80058 Euura scutellata (Hartig, 1837) adult male 1 Germany Brandenburg Muencheberg,
148341 88536 DEI-GISHym88536 Euura spiraeae (Zaddach, 1883) adult female 1 Germany Brandenburg Eberswalde, Waldstrasse 19: Garten mit Waldrand 52.8264, 13.84136 2017-05-25 2017-06-05 A. Taeger Malaise trap
146883 83744 DEI-GISHym83744 Euura sylvestris (Cameron, 1884) larva 1 Austria Nordtirol Hochgurgl, 500 m N, side of Timmelsjoch road 46.90496, 11.05144 2017-08-17 C. Michell & A. Liston M. Prous Salix hegetschweileri
158316 59158 http://id.luomus.fi/GL.3493 Euura sylvestris (Cameron, 1884) adult female 1 Finland Uusimaa Espoo 60.21666, 24.66666 1964-06-02 J. Perkioemaeki E. Lindqvist holotype Pteronidea disparoides Lindqvist, 1969
151659 56564 Prep.310.OR Euura sylvestris (Cameron, 1884) adult female 1 Finland Uusimaa Helsinki 60.17555, 24.93416 1963-01-01 1963-12-31 CVV
173821 110342
151667 56572 http://id.luomus.fi/GL.9216 Euura sylvestris (Cameron, 1884) adult male 1 Finland Lapin Laeaeni Kilpisjaervi 69.05, 20.8 1968-07-09 L. H. Woollatt M. Prous paratype Pteronidea woollatti Lindqvist, 1971
155573 58975
166247 59290 GP.97871 Euura sylvestris (Cameron, 1884) adult female 1 Finland Kumpula, Botanical garden 60.2016, 24.9607 2016-08-15 2016-08-21 J. Paukkunen Malaise trap M. Prous
151292 56521 PR.574VV Euura sylvestris (Cameron, 1884) adult female 1 Finland Northern Ostrobothnia Kuusamo [736:60] 66.359, 29.342 1979-07-01 V. Vikberg V. Vikberg CVV
175156 111639
175360 111044
175703 111102
175708 111111
152174 56629 http://id.luomus.fi/GL.3443 Euura sylvestris (Cameron, 1884) adult female 1 Finland Ostrobottnia australis Mustasaari 63.16666, 21.66666 1963-07-24 J. Perkioemaeki E. Lindqvist holotype Pteronidea angustiserra Lindqvist, 1969
64947 111965 http://id.luomus.fi/GL.3512 Euura sylvestris (Cameron, 1884) adult female 1 Finland Nurmes 63.55, 29.11666 1939-06-18 A. Saarinen M. Prous ? holotype Pteronidea straminea Lindqvist, 1958
176710 111252
176711 111253
176712 111254
217164 111940
157558 59131
217165 111941
217166 111942
158211 59151
136171 20993 DEI-GISHym20993 Euura sylvestris (Cameron, 1884) adult female 1 Lithuania Panevezys Pajiesmeniai 2 km S 56.094, 24.479 35 m 2015-06-08 A.D. Liston & A. Taeger netting M. Prous
157617 12018 DEI-GISHym12018 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark Abisko 68.349, 18.839 370 m 2016-07-04 A. Liston & M. Prous netting M. Prous
96411 21407 DEI-GISHym21407 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark Abisko 6 km E 68.348, 18.969 400 m 2016-07-04 A. Liston & M. Prous netting M. Prous
149746 21405 DEI-GISHym21405 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark Abisko 6 km E 68.348, 18.969 400 m 2016-07-04 A. Liston & M. Prous netting M. Prous
96483 21376 DEI-GISHym21376 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark Abisko: Mt. Njulla 68.362, 18.73 400 m 950 m 2016-06-30 A. Liston & M. Prous netting M. Prous
157483 12008 DEI-GISHym12008 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark Nikkaluokta 7 km NE 67.89, 19.138 500 m 1050 m 2016-07-08 A. Liston & M. Prous netting M. Prous
133762 83594 DEI-GISHym83594 Euura sylvestris (Cameron, 1884) adult male 1 Sweden Gotland Roma 57.5, 18.447 2017-06-08 A.D. Liston netting M. Prous
96487 21351 DEI-GISHym21351 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark: Abisko Oe 1-5 km SW 68.346, 18.781 350 m 2016-06-25 A.D. Liston netting M. Prous
131796 21339 DEI-GISHym21339 Euura sylvestris (Cameron, 1884) adult female 1 Sweden Torne Lappmark: Kiruna, near airport 67.84, 20.35 450 m 2016-06-22 A.D. Liston netting M. Prous
148499 88694 DEI-GISHym88694 Euura tibialis (Newman, 1837) adult female 1 Germany Brandenburg Eberswalde, Waldstrasse 19: Garten mit Waldrand 52.8264, 13.84136 2017-08-27 2017-09-29 A. Taeger Malaise trap
78329 15293 DEI-GISHym15293 Euura vaga (Fabricius, 1781) adult male 1 Canada Quebec Gatineau Park 1.8 km N Eardley, Juniperus virginiana stand 45.56667, -76.09139 60 m 80 m 2012-05-31 2012-06-07 CNC Hymenoptera Team Malaise trap A. D. Liston
150911 84189 DEI-GISHym84189 Euura variator (Ruthe, 1859) adult male 1 Sweden Torne Lappmark Bjoerkliden 68.409, 18.639 500 m 850 m 2016-07-01 A. Liston & M. Prous netting
148289 88484 DEI-GISHym88484 Euura vicina (Serville, 1823) adult male 1 Germany Brandenburg Eberswalde, Waldstrasse 19: Garten mit Waldrand 52.8264, 13.84136 2017-05-18 2017-05-25 A. Taeger Malaise trap
157550 12011 DEI-GISHym12011 Euura viduata (Zetterstedt, 1838) adult male 1 Germany Brandenburg Muencheberg, Gr. Schlagenthinsee 52.51979, 14.10212 2019-04-04 A. Liston & M. Prous M. Prous
158696 12064 DEI-GISHym12064 Euura villosa (Thomson, 1863) adult female 1 Norway Finnmark Batsfjord 9 km SW 70.568, 29.554 230 m 2019-06-27 A. Liston & M. Prous netting M. Prous
64984 59355 http://id.luomus.fi/GL.3506 Euura viridis (Stephens, 1835) adult female 1 Finland Ahlainen, Rankku 61.61666, 21.56666 1958-07-06 V. Lauro holotype Pteronidea lauroi Lindqvist, 1960
215759 84547 DEI-GISHym84547 Euura viridis (Stephens, 1835) adult male 1 Finland Lapland Kilpisjaervi, Saana 69.037, 20.844 500 m 900 m 2020-06-29 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
155487 58925
155488 58926
46497 56552 http://id.luomus.fi/GL.3441 Euura viridis (Stephens, 1835) adult female 1 Finland Kuusamo 65.96666, 29.18333 J. Sahlberg M. Prous holotype Pteronidea abscondita Lindqvist, 1949
215760 12603 DEI-GISHym12603 Euura viridis (Stephens, 1835) adult male 1 Finland Lapland Lammasoaivi 68.778, 21.338 600 m 2020-06-26 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
152418 56757
152420 56759
152528 56867
152510 56849
152511 56850
152512 56851
215666 111829 http://id.luomus.fi/GL.3461 Euura viridis (Stephens, 1835) adult female 1 Finland Utsjoki, Outakoski 69.6, 25.96666 1948-06-24 E. Lindqvist E. Lindqvist holotype Pteronidea breviseta Lindqvist, 1949
51190 3386 DEI-GISHym3386 Euura viridis (Stephens, 1835) adult female 1 Germany Nordrhein-Westfalen Aachen Umgebung 50.76666, 6.1 lectotype Nematus polyspilus Foerster, 1854
49892 3388 DEI-GISHym3388 Euura viridis (Stephens, 1835) adult female 1 Germany Germany 51.5, 10.5 lectotype Nematus prasinus Hartig, 1837
158841 12150 DEI-GISHym12150 Euura viridis (Stephens, 1835) adult male 1 Germany Thueringen Ilfeld: Netzkater: Brandesbachtal 51.6, 10.81 350 m 400 m 2019-05-28 23rd Symphyta Workshop netting M. Prous ?
80275 50443 BC
149747 21406 DEI-GISHym21406 Euura viridis (Stephens, 1835) adult female 1 Sweden Torne Lappmark Abisko 6 km E 68.348, 18.969 400 m 2016-07-04 A. Liston & M. Prous netting M. Prous
87277 31131 DEI-GISHym31131 Euura viridis (Stephens, 1835) adult female 1 Sweden Norrbottens Laen Abisko National Park 68.342, 18.755 410 m 2012-06-27 A.D. Liston & A. Taeger netting M. Prous
87288 31142 DEI-GISHym31142 Euura viridis (Stephens, 1835) adult female 1 Sweden Norrbottens Laen Abisko National Park, E10 68.353, 18.815 390 m 2012-06-25 A.D. Liston & A. Taeger netting M. Prous ?
96499 21364 DEI-GISHym21364 Euura viridis (Stephens, 1835) adult female 1 Sweden Torne Lappmark: Bjoerkliden 2 km SE 68.395, 18.705 500 m 800 m 2016-06-27 A.D. Liston netting M. Prous
216658 111908 B.M.TYPE HYM1.697 Euura viridis (Stephens, 1835) adult female 1 United Kingdom England London, in the vicinity of Stephens, J. F. holotype Nematus viridis Stephens, 1835
150434 84172 DEI-GISHym84172 Euura vittata (Serville, 1823) adult male 1 Bulgaria Sliven Ichera 3 km SW 42.749, 26.421 730 m 2018-04-14 A. Liston & M. Prous netting M. Prous
131794 21330 DEI-GISHym21330 Euura weiffenbachiella Liston & Vikberg, 2017 larva 1 Sweden Skane Sandhammeren [N55.387 E14.191] 55.387, 14.191 2016-06-14 A.D. Liston photography A. D. Liston Salix repens
149240 83827 DEI-GISHym83827 Megadineura grandis (Andre, 1882) adult male 1 Japan Nagano Suzaka 10 km SSW 36.538, 138.295 1350 m 2017-05-30 A. Taeger netting M. Prous
88820 80015 DEI-GISHym80015 Mesoneura opaca (Fabricius, 1775) adult female 1 Morocco Meknes-Tafilalet Region Ifrane 7 km NW 33.552, -5.175 1590 m 2015-04-20 A. Liston & M. Prous netting M. Prous
95621 86259 DEI-GISHym86259 Nematus adult male 1 Russia Primorsky Krai Arsenyev, Ski-Base Bodrost 44.122, 133.27 200 m 2016-05-25 K. Kramp, M. Prous & A. Taeger netting M. Prous
155310 58866 TUZ109010 Nematus alniastri (Scharfenberg, 1805) adult female 1 Estonia Saaremaa Abruka [58.159N 22.501E] 58.159, 22.501 2017-07-25 V. Soon A. D. Liston
133692 80332 DEI-GISHym80332 Nematus lucidus (Panzer, 1801) adult male 1 Greece Achaia Achaiko Chorio S 38.1371, 22.061 1150 m 2017-04-25
89023 31229 DEI-GISHym31229 Nematus princeps Zaddach, 1876 adult male 1 Estonia Ida-Virumaa Vasavere 1.5 km E 59.296, 27.543 50 m 2015-06-04 H. Vardal netting M. Prous
215573 84672 DEI-GISHym84672 Nematus tulunensis Vikberg, 1972 larva 1 Finland Northern Ostrobothnia Oulu, kasvipuutarha [7219:3427] 65.06365, 25.45948 2020-06-30 M. Mutanen, M. Prous, A. Liston M. Prous & A. Liston Lonicera tatarica
87921 31044 DEI-GISHym31044 Nematus umbratus Thomson, 1871 adult female 1 Sweden Norrbottens Laen Pajala 67.204, 23.409 150 m 2014-06-05 A. Liston & M. Prous netting A. D. Liston
216201 84540 DEI-GISHym84540 Pristiphora astragali Vikberg, 1978 adult female 1 Finland Lapland Kilpisjaervi, Jehkas 69.086, 20.8 550 m 950 m 2020-06-28 M. Mutanen, M. Prous, A. Liston netting M. Prous & A. Liston
94230 80158 DEI-GISHym80158 Pristiphora cincta Newman, 1837 adult male 1 Sweden Torne Lappmark Abisko 6 km W 68.342, 18.691 650 m 900 m 2016-07-02 A. Liston & M. Prous netting M. Prous
133670 80321 DEI-GISHym80321 Pristiphora cretica W. Schedl, 1981 adult male 1 Greece Achaia Achaiko Chorio S 38.1371, 22.061 1150 m 2017-04-25